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<mods:title id="81566E2BDBA023F67CB0077072BC075D">A new Moenkhausia (Characiformes: Characidae) from rio Braço Norte, rio Tapajós basin, with comments on the fish endemism of Serra do Cachimbo plateau</mods:title>
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<mods:namePart id="0DC116AB7A804086651DD2609B4FD53A">Vita, George</mods:namePart>
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<mods:affiliation id="02F31EB2108A0C808180571E9E9772A3">Museu de Zoologia da Universidade de São Paulo, Av. Nazaré 481, 04263000, São Paulo, SP, Brazil &amp; george. vitao @ gmail. com; https: // orcid. org / 0000 - 0003 - 2527 - 1950</mods:affiliation>
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<mods:affiliation id="4623D2182FF1E3A67561E106059A75A4">Universidade do Estado de Minas Gerais, Unidade Acadêmica Passos, Avenida Juca Stockler, 1130, 37900106, Passos, Minas Gerais, Brazil &amp; guilhermedutr @ yahoo. com. br; https: // orcid. org / 0000 - 0002 - 1010 - 3738</mods:affiliation>
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(
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SL, 
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and 
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from 
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, 
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drainage, 
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basin
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: 
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(32, 2 C&amp;S, 
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SL)
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<specimenCode id="561F9EA1EA502676FE5FD3AE572CF876" box="[454,594,1950,1977]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="587">INPA 59895</specimenCode>
(5, 
<quantity id="C1419B3FEA502676FD1BD3AF5659F876" box="[642,807,1950,1977]" metricMagnitude="-2" metricUnit="m" metricValue="6.305" metricValueMax="7.75" metricValueMin="4.86" pageId="1" pageNumber="587" unit="mm" value="63.05" valueMax="77.5" valueMin="48.6">48.6–77.5 mm</quantity>
SL), collected with holotype
</materialsCitation>
. 
<materialsCitation id="B6D13C87EA502675FB1AD3AE5481FF7E" collectingDate="2015-08-03" collectionCode="MZUSP" collectorName="O. Oyakawa &amp; W. Ohara &amp; Pastana" country="Brazil" lastPageId="2" lastPageNumber="588" latitude="-9.366639" location="Igarape Olho" longLatPrecision="1" longitude="-54.91539" municipality="rio Braco Norte" pageId="1" pageNumber="587" specimenCode="MZUSP 119401" specimenCount="1" typeStatus="paratype">
<specimenCode id="561F9EA1EA502676FB1AD3AE5045F876" box="[1155,1339,1951,1977]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="587">MZUSP 119401</specimenCode>
(1, 
<quantity id="C1419B3FEA502676FAF2D3AE55BDF812" metricMagnitude="-2" metricUnit="m" metricValue="6.76" pageId="1" pageNumber="587" unit="mm" value="67.6">67.6 mm</quantity>
SL), 
<location id="03666001EA502676FE9AD3F254E8F812" LSID="urn:lsid:plazi:treatment:8E1087CCEA502671FF0ED24F546AF9C7:03666001EA502676FE9AD3F254E8F812" box="[259,406,1987,2013]" country="Brazil" latitude="-9.366639" longLatPrecision="1" longitude="-54.91539" municipality="rio Braco Norte" name="Igarape Olho" pageId="1" pageNumber="587">Igarapé Olho</location>
d´água, a tributary of 
<collectingMunicipality id="E662ACA0EA502676FD0DD3F25636F812" box="[660,840,1987,2013]" pageId="1" pageNumber="587">rio Braço Norte</collectingMunicipality>
, 
<geoCoordinate id="638D501DEA502676FCCAD3F356A1F812" box="[851,991,1986,2013]" degrees="9" direction="south" minutes="21" orientation="latitude" pageId="1" pageNumber="587" precision="1" seconds="59.9" value="-9.366639">9°21’59.9”S</geoCoordinate>
<geoCoordinate id="638D501DEA502676FC7FD3F351F5F812" box="[998,1163,1986,2013]" degrees="54" direction="west" minutes="54" orientation="longitude" pageId="1" pageNumber="587" precision="1" seconds="55.4" value="-54.91539">54°54’55.4”W</geoCoordinate>
, 
<collectorName id="AB4C530CEA502676FB0ED3F25059F812" box="[1175,1319,1987,2013]" pageId="1" pageNumber="587">O. Oyakawa</collectorName>
, 
<collectorName id="AB4C530CEA502676FAA8D3F250E2F812" box="[1329,1436,1987,2013]" pageId="1" pageNumber="587">W. Ohara</collectorName>
and M. 
<collectorName id="AB4C530CEA532675FF69D4A65436FF7E" box="[240,328,151,177]" pageId="2" pageNumber="588">Pastana</collectorName>
, 
<date id="7207101AEA532675FECCD4A65481FF7E" box="[341,511,150,177]" pageId="2" pageNumber="588" value="2015-08-03">
<collectingDate id="6243E9F2EA532675FECCD4A65481FF7E" box="[341,511,150,177]" pageId="2" pageNumber="588" value="2015-08-03">3 August 2015</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="B6D13C87EA532675FD93D4A65674FF1A" collectionCode="MZUSP" collectorName="F. Dagosta &amp; M. M. F. Marinho &amp; Camelier" country="Brazil" latitude="-9.304444" location="rio Braco Norte" longLatPrecision="21" longitude="-54.94528" pageId="2" pageNumber="588" specimenCode="MZUSP 119869" specimenCount="1" typeStatus="paratype">
<specimenCode id="561F9EA1EA532675FD93D4A657BCFF7E" box="[522,706,151,177]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="588">MZUSP 119869</specimenCode>
(1, 
<quantity id="C1419B3FEA532675FD68D4A65628FF7E" box="[753,854,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="7.709999999999999" pageId="2" pageNumber="588" unit="mm" value="77.1">77.1 mm</quantity>
SL), small tributary of 
<location id="03666001EA532675FBFDD4A65064FF7E" LSID="urn:lsid:plazi:treatment:8E1087CCEA502671FF0ED24F546AF9C7:03666001EA532675FBFDD4A65064FF7E" box="[1124,1306,151,177]" country="Brazil" latitude="-9.304444" longLatPrecision="21" longitude="-54.94528" name="rio Braco Norte" pageId="2" pageNumber="588">rio Braço Norte</location>
, 
<geoCoordinate id="638D501DEA532675FABFD4A650E2FF7E" box="[1318,1436,151,177]" degrees="9" direction="south" minutes="18" orientation="latitude" pageId="2" pageNumber="588" precision="15" seconds="16" value="-9.304444">9°18’16”S</geoCoordinate>
<geoCoordinate id="638D501DEA532675FF0ED48B5459FF1A" box="[151,295,186,213]" degrees="54" direction="west" minutes="56" orientation="longitude" pageId="2" pageNumber="588" precision="15" seconds="43" value="-54.94528">54°56’43”W</geoCoordinate>
, 
<collectorName id="AB4C530CEA532675FEA8D48A54D5FF1A" box="[305,427,187,213]" pageId="2" pageNumber="588">F. Dagosta</collectorName>
, 
<collectorName id="AB4C530CEA532675FE2CD48A5701FF1A" box="[437,639,187,213]" pageId="2" pageNumber="588">M. M. F. Marinho</collectorName>
, P. 
<collectorName id="AB4C530CEA532675FD3AD48A5674FF1A" box="[675,778,187,213]" pageId="2" pageNumber="588">Camelier</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="B6D13C87EA532675FCB5D48A510DFF1A" box="[812,1139,186,213]" collectingDate="2015-08-03" collectionCode="V" country="Brazil" location="Giovanetti" pageId="2" pageNumber="588" specimenCount="1" typeStatus="paratype">
<collectionCode id="60A8AE1FEA532675FCB5D48A563EFF1A" box="[812,832,187,213]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="588" type="Museum">V</collectionCode>
. 
<location id="03666001EA532675FCD3D48A56BDFF1A" LSID="urn:lsid:plazi:treatment:8E1087CCEA502671FF0ED24F546AF9C7:03666001EA532675FCD3D48A56BDFF1A" box="[842,963,187,213]" country="Brazil" name="Giovanetti" pageId="2" pageNumber="588">Giovanetti</location>
, 
<date id="7207101AEA532675FC54D48A510DFF1A" box="[973,1139,186,213]" pageId="2" pageNumber="588" value="2015-08-03">
<collectingDate id="6243E9F2EA532675FC54D48A510DFF1A" box="[973,1139,186,213]" pageId="2" pageNumber="588" value="2015-08-03">3 August 2015</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="B6D13C87EA532675FBE4D48A50E3FF36" collectionCode="MZUSP" collectorName="F. Dagosta &amp; M. M. F. Marinho &amp; Camelier" country="Brazil" latitude="-9.275472" location="rio Braco Norte" longLatPrecision="1" longitude="-54.82039" pageId="2" pageNumber="588" specimenCode="MZUSP 119922" specimenCount="1" typeStatus="paratype">
<specimenCode id="561F9EA1EA532675FBE4D48A504CFF1A" box="[1149,1330,187,213]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="588">MZUSP 119922</specimenCode>
(2, 
<quantity id="C1419B3FEA532675FAC4D48A5585FF36" metricMagnitude="-2" metricUnit="m" metricValue="7.584999999999999" metricValueMax="7.890000000000001" metricValueMin="7.279999999999999" pageId="2" pageNumber="588" unit="mm" value="75.85" valueMax="78.9" valueMin="72.8">72.8– 78.9 mm</quantity>
SL), tributary of 
<location id="03666001EA532675FE26D4EE570CFF36" LSID="urn:lsid:plazi:treatment:8E1087CCEA502671FF0ED24F546AF9C7:03666001EA532675FE26D4EE570CFF36" box="[447,626,223,249]" country="Brazil" latitude="-9.275472" longLatPrecision="1" longitude="-54.82039" name="rio Braco Norte" pageId="2" pageNumber="588">rio Braço Norte</location>
, 
<geoCoordinate id="638D501DEA532675FDE4D4EE5676FF36" box="[637,776,223,249]" degrees="9" direction="south" minutes="16" orientation="latitude" pageId="2" pageNumber="588" precision="1" seconds="31.7" value="-9.275472">9°16’31.7”S</geoCoordinate>
<geoCoordinate id="638D501DEA532675FC96D4EF56CDFF36" box="[783,947,222,249]" degrees="54" direction="west" minutes="49" orientation="longitude" pageId="2" pageNumber="588" precision="1" seconds="13.4" value="-54.82039">54°49’13.4”W</geoCoordinate>
, 
<collectorName id="AB4C530CEA532675FC26D4EE5147FF36" box="[959,1081,223,249]" pageId="2" pageNumber="588">F. Dagosta</collectorName>
, 
<collectorName id="AB4C530CEA532675FBDDD4EE506EFF36" box="[1092,1296,223,249]" pageId="2" pageNumber="588">M. M. F. Marinho</collectorName>
, P. 
<collectorName id="AB4C530CEA532675FAAFD4EE50E3FF36" box="[1334,1437,223,249]" pageId="2" pageNumber="588">Camelier</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="B6D13C87EA532675FF2AD5325480FED2" box="[179,510,258,285]" collectingDate="2015-08-04" collectionCode="V" country="Brazil" location="Giovanetti" pageId="2" pageNumber="588" specimenCount="1" typeStatus="paratype">
<collectionCode id="60A8AE1FEA532675FF2AD53255B9FED2" box="[179,199,259,285]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="588" type="Museum">V</collectionCode>
. 
<location id="03666001EA532675FF4BD5325435FED2" LSID="urn:lsid:plazi:treatment:8E1087CCEA502671FF0ED24F546AF9C7:03666001EA532675FF4BD5325435FED2" box="[210,331,259,285]" country="Brazil" name="Giovanetti" pageId="2" pageNumber="588">Giovanetti</location>
, 
<date id="7207101AEA532675FECFD5325480FED2" box="[342,510,258,285]" pageId="2" pageNumber="588" value="2015-08-04">
<collectingDate id="6243E9F2EA532675FECFD5325480FED2" box="[342,510,258,285]" pageId="2" pageNumber="588" value="2015-08-04">4 August 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="52C66652EA532675FF0ED69D5435FD27" ID-DOI="http://doi.org/10.5281/zenodo.8255380" ID-Zenodo-Dep="8255380" httpUri="https://zenodo.org/record/8255380/files/figure.png" pageId="2" pageNumber="588" startId="2.[151,250,684,708]" targetBox="[378,1209,309,660]" targetPageId="2" targetType="figure">
<paragraph id="060636DAEA532675FF0ED69D5435FD27" blockId="2.[151,1436,684,744]" pageId="2" pageNumber="588">
<emphasis id="34CDEAC8EA532675FF0ED69D546DFD0B" bold="true" box="[151,275,684,708]" pageId="2" pageNumber="588">FIGURE 1.</emphasis>
Holotype of 
<taxonomicName id="C1B94D59EA532675FE01D69D57A3FD0B" authority=", MZUSP" authorityName="MZUSP" box="[408,733,684,708]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA532675FE01D69D5703FD0B" box="[408,637,684,708]" italics="true" pageId="2" pageNumber="588">Moenkhausia guaruba</emphasis>
, MZUSP
</taxonomicName>
128235, 96.9 mm SL, Brazil, Pará, Novo Progresso, rio Braço Norte, rio Tapajós basin.
</paragraph>
</caption>
<caption id="52C66652EA532675FF0ED10B55BEFA55" ID-DOI="http://doi.org/10.5281/zenodo.8255382" ID-Zenodo-Dep="8255382" httpUri="https://zenodo.org/record/8255382/files/figure.png" pageId="2" pageNumber="588" startId="2.[151,250,1338,1362]" targetBox="[388,1200,774,1306]" targetPageId="2" targetType="figure">
<paragraph id="060636DAEA532675FF0ED10B55BEFA55" blockId="2.[151,1436,1338,1434]" pageId="2" pageNumber="588">
<emphasis id="34CDEAC8EA532675FF0ED10B546BFA9D" bold="true" box="[151,277,1338,1362]" pageId="2" pageNumber="588">FIGURE 2.</emphasis>
Number of branched anal-fin rays in 
<emphasis id="34CDEAC8EA532675FD3CD10B56F2FA9D" box="[677,908,1338,1362]" italics="true" pageId="2" pageNumber="588">
Moenkahusia 
<taxonomicName id="C1B94D59EA532675FCAFD10B56F2FA9D" box="[822,908,1338,1362]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="guaruba">guaruba</taxonomicName>
</emphasis>
(blue) and 
<taxonomicName id="C1B94D59EA532675FB91D10B519FFA9D" authorityName="Gery, Planquette &amp; Le Bail" authorityYear="1995" box="[1032,1249,1338,1362]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="moisae">
<emphasis id="34CDEAC8EA532675FB91D10B519FFA9D" box="[1032,1249,1338,1362]" italics="true" pageId="2" pageNumber="588">Moenkhausia moisae</emphasis>
</taxonomicName>
(red). Data of 
<taxonomicName id="C1B94D59EA532675FA18D10A5593FAB9" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA532675FA18D10A50E8FA9D" box="[1409,1430,1339,1362]" italics="true" pageId="2" pageNumber="588">M</emphasis>
. 
<emphasis id="34CDEAC8EA532675FF0ED16F5593FAB9" box="[151,237,1374,1398]" italics="true" pageId="2" pageNumber="588">guaruba</emphasis>
</taxonomicName>
from the type series (MZUSP 128235 and MZUSP 119389; n = 30), and of 
<taxonomicName id="C1B94D59EA532675FB92D16E5108FAB9" authorityName="Gery, Planquette &amp; Le Bail" authorityYear="1995" box="[1035,1142,1375,1398]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="moisae">
<emphasis id="34CDEAC8EA532675FB92D16E5108FAB9" box="[1035,1142,1375,1398]" italics="true" pageId="2" pageNumber="588">M. moisae</emphasis>
</taxonomicName>
from 
<bibRefCitation id="62284B2BEA532675FB21D16F5013FAB9" author="Gery, J. &amp; Planquette, P. &amp; Le Bail, P. Y." box="[1208,1389,1374,1398]" pageId="2" pageNumber="588" pagination="67 - 70" refId="ref6473" refString="Gery, J., Planquette, P. &amp; Le Bail, P. Y. (1995) Une espece nouvelle de Moenkhausia de la Guyane (Teleostei, Ostariophysi, Characidae), a ecailles nombreuses. Revue francaise d'aquariologie, 22 (3 - 4), 67 - 70." type="journal article" year="1995">
Géry 
<emphasis id="34CDEAC8EA532675FB6BD16E505FFAB9" box="[1266,1313,1374,1398]" italics="true" pageId="2" pageNumber="588">et al</emphasis>
. (1995
</bibRefCitation>
; n = 60).
</paragraph>
</caption>
<subSubSection id="4EA36551EA532675FF5ED1DD57A5F85C" pageId="2" pageNumber="588" type="diagnosis">
<paragraph id="060636DAEA532675FF5ED1DD57A5F85C" blockId="2.[151,1437,1516,2011]" pageId="2" pageNumber="588">
<emphasis id="34CDEAC8EA532675FF5ED1DD543CF9C9" bold="true" box="[199,322,1516,1542]" pageId="2" pageNumber="588">Diagnosis.</emphasis>
<taxonomicName id="C1B94D59EA532675FED7D1DD572EF9C9" box="[334,592,1516,1542]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA532675FED7D1DD572EF9C9" box="[334,592,1516,1542]" italics="true" pageId="2" pageNumber="588">Moenkhausia guaruba</emphasis>
</taxonomicName>
is distinguished from its congeners, except 
<taxonomicName id="C1B94D59EA532675FBF9D1DD54C1F9E4" authority="Gery, Planquette &amp; Le Bail 1995" authorityName="Gery, Planquette &amp; Le Bail" authorityYear="1995" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="moisae">
<emphasis id="34CDEAC8EA532675FBF9D1DD502CF9C9" box="[1120,1362,1516,1542]" italics="true" pageId="2" pageNumber="588">Moenkhausia moisae</emphasis>
<bibRefCitation id="62284B2BEA532675FAC6D1DC54C1F9E4" author="Gery, J. &amp; Planquette, P. &amp; Le Bail, P. Y." pageId="2" pageNumber="588" pagination="67 - 70" refId="ref6473" refString="Gery, J., Planquette, P. &amp; Le Bail, P. Y. (1995) Une espece nouvelle de Moenkhausia de la Guyane (Teleostei, Ostariophysi, Characidae), a ecailles nombreuses. Revue francaise d'aquariologie, 22 (3 - 4), 67 - 70." type="journal article" year="1995">Géry, Planquette &amp; Le Bail 1995</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="C1B94D59EA532675FE6AD2215113F9E4" authority="Zanata, Birindelli &amp; Moreira 2010" authorityName="Zanata, Birindelli &amp; Moreira" authorityYear="2010" box="[499,1133,1552,1579]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="pirauba">
<emphasis id="34CDEAC8EA532675FE6AD2215797F9E5" box="[499,745,1552,1578]" italics="true" pageId="2" pageNumber="588">Moenkhausia pirauba</emphasis>
<bibRefCitation id="62284B2BEA532675FD76D2205113F9E4" author="Zanata, A. M. &amp; Birindelli, J. L. O. &amp; Moreira, C. R." box="[751,1133,1552,1579]" pageId="2" pageNumber="588" pagination="2615 - 2628" refId="ref8466" refString="Zanata, A. M., Birindelli, J. L. O. &amp; Moreira, C. R. (2010) New species of Moenkhausia Eigenmann (Characiformes: Characidae) from Rio Xingu and Rio Tapajos basins, Brazil, with comments on a putative case of polymorphic Batesian mimicry. Journal of Fish Biology, 75 (10), 2615 - 2628. https: // doi. org / 10.1111 / j. 1095 - 8649.2009.02455. x" type="journal article" year="2010">Zanata, Birindelli &amp; Moreira 2010</bibRefCitation>
</taxonomicName>
by having a higher number of scales in the longitudinal series (43–46 
<emphasis id="34CDEAC8EA532675FDEED20457F0F981" box="[631,654,1589,1614]" italics="true" pageId="2" pageNumber="588">vs</emphasis>
. 
<quantity id="C1419B3FEA532675FD07D204567CF981" box="[670,770,1588,1615]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" metricValueMax="10.414" metricValueMin="5.842" pageId="2" pageNumber="588" unit="in" value="32.0" valueMax="41.0" valueMin="23.0">23–41 in</quantity>
other 
<taxonomicName id="C1B94D59EA532675FCD2D205513EF981" box="[843,1088,1588,1614]" pageId="2" pageNumber="588">
<emphasis id="34CDEAC8EA532675FCD2D205569CF981" box="[843,994,1588,1614]" italics="true" pageId="2" pageNumber="588">Moenkhausia</emphasis>
species
</taxonomicName>
). The new species differs from 
<taxonomicName id="C1B94D59EA532675FF0ED2685473F9BD" authorityName="Gery, Planquette &amp; Le Bail" authorityYear="1995" box="[151,269,1625,1651]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="moisae">
<emphasis id="34CDEAC8EA532675FF0ED26855D0F9BD" box="[151,174,1625,1650]" italics="true" pageId="2" pageNumber="588">M</emphasis>
. 
<emphasis id="34CDEAC8EA532675FF27D2685473F9BD" box="[190,269,1625,1650]" italics="true" pageId="2" pageNumber="588">moisae</emphasis>
</taxonomicName>
by having fewer branched anal-fin rays (21–25, modal 23 
<emphasis id="34CDEAC8EA532675FC2BD26856B7F9BD" box="[946,969,1625,1650]" italics="true" pageId="2" pageNumber="588">vs</emphasis>
. 25–29, modal 
<quantity id="C1419B3FEA532675FBE7D26851C7F9BD" box="[1150,1209,1624,1651]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="2" pageNumber="588" unit="in" value="27.0">27 in</quantity>
<taxonomicName id="C1B94D59EA532675FB5BD2685046F9BD" authorityName="Gery, Planquette &amp; Le Bail" authorityYear="1995" box="[1218,1336,1625,1650]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="moisae">
<emphasis id="34CDEAC8EA532675FB5BD2685046F9BD" box="[1218,1336,1625,1650]" italics="true" pageId="2" pageNumber="588">M. moisae</emphasis>
</taxonomicName>
; 
<figureCitation id="9E822A5FEA532675FAD0D26950F0F9BC" box="[1353,1422,1624,1651]" captionStart="FIGURE 2" captionStartId="2.[151,250,1338,1362]" captionTargetBox="[388,1200,774,1306]" captionTargetId="figure-106@2.[378,1210,768,1314]" captionTargetPageId="2" captionText="FIGURE 2. Number of branched anal-fin rays in Moenkahusia guaruba (blue) and Moenkhausia moisae (red). Data of M. guaruba from the type series (MZUSP 128235 and MZUSP 119389; n = 30), and of M. moisae from Géry et al. (1995; n = 60)." figureDoi="http://doi.org/10.5281/zenodo.8255382" httpUri="https://zenodo.org/record/8255382/files/figure.png" pageId="2" pageNumber="588">Fig. 2</figureCitation>
), a complete and regularly arranged series of predorsal scales (
<emphasis id="34CDEAC8EA532675FCCBD24C560EF959" box="[850,880,1661,1686]" italics="true" pageId="2" pageNumber="588">vs.</emphasis>
irregular arranged of scales at predorsal region), and by having a single, vertically elongated and relatively wide humeral blotch (
<emphasis id="34CDEAC8EA532675FBA1D2905131F975" box="[1080,1103,1697,1722]" italics="true" pageId="2" pageNumber="588">vs</emphasis>
. two humeral blotches in 
<taxonomicName id="C1B94D59EA532675FAE7D2905598F911" authorityName="Gery, Planquette &amp; Le Bail" authorityYear="1995" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="moisae">
<emphasis id="34CDEAC8EA532675FAE7D2905598F911" italics="true" pageId="2" pageNumber="588">M. moisae</emphasis>
</taxonomicName>
; see Discussion for further details). 
<taxonomicName id="C1B94D59EA532675FDEFD2F5560CF911" box="[630,882,1732,1758]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA532675FDEFD2F5560CF911" box="[630,882,1732,1758]" italics="true" pageId="2" pageNumber="588">Moenkhausia guaruba</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="C1B94D59EA532675FB9BD2F45101F911" authorityName="Zanata, Birindelli &amp; Moreira" authorityYear="2010" box="[1026,1151,1732,1759]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="pirauba">
<emphasis id="34CDEAC8EA532675FB9BD2F45167F911" box="[1026,1049,1733,1758]" italics="true" pageId="2" pageNumber="588">M</emphasis>
. 
<emphasis id="34CDEAC8EA532675FBBFD2F55101F911" box="[1062,1151,1732,1758]" italics="true" pageId="2" pageNumber="588">pirauba</emphasis>
</taxonomicName>
by having a conspicuous, rounded, and symmetrical dark blotch located at the posterior limit of the caudal peduncle and base of caudal-fin rays (
<emphasis id="34CDEAC8EA532675FE99D33C5469F8E9" box="[256,279,1805,1830]" italics="true" pageId="2" pageNumber="588">vs</emphasis>
. caudal blotch horizontally elongated, asymmetrical, continuous anteriorly with midlateral stripe and extending posteriorly to margins of four or five middle caudal-fin rays in 
<taxonomicName id="C1B94D59EA532675FC4FD300512BF885" authorityName="Zanata, Birindelli &amp; Moreira" authorityYear="2010" box="[982,1109,1840,1866]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="pirauba">
<emphasis id="34CDEAC8EA532675FC4FD300512BF885" box="[982,1109,1840,1866]" italics="true" pageId="2" pageNumber="588">M. pirauba</emphasis>
</taxonomicName>
), and a thin longitudinal line formed by dark pigmentation running along horizontal septum of body (
<emphasis id="34CDEAC8EA532675FC26D36456A0F8A1" box="[959,990,1877,1902]" italics="true" pageId="2" pageNumber="588">vs.</emphasis>
dark longitudinal line wide, forming an elongated blotch at caudal peduncle in 
<taxonomicName id="C1B94D59EA532675FDD5D34857B4F85D" authorityName="Zanata, Birindelli &amp; Moreira" authorityYear="2010" box="[588,714,1912,1938]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="pirauba">
<emphasis id="34CDEAC8EA532675FDD5D34857B4F85D" box="[588,714,1912,1938]" italics="true" pageId="2" pageNumber="588">M. pirauba</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="4EA36551EA532671FF5ED3AD5675FAB7" lastPageId="6" lastPageNumber="592" pageId="2" pageNumber="588" type="description">
<paragraph id="060636DAEA532674FF5ED3AD5798FEAA" blockId="2.[151,1437,1516,2011]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="589" pageId="2" pageNumber="588">
<emphasis id="34CDEAC8EA532675FF5ED3AD5427F879" bold="true" box="[199,345,1948,1974]" pageId="2" pageNumber="588">Description.</emphasis>
Morphometric data of 
<taxonomicName id="C1B94D59EA532675FDC0D3AD5628F879" box="[601,854,1948,1974]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="588" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA532675FDC0D3AD5628F879" box="[601,854,1948,1974]" italics="true" pageId="2" pageNumber="588">Moenkhausia guaruba</emphasis>
</taxonomicName>
in 
<tableCitation id="4B3B0361EA532675FCEED3AD56B4F878" box="[887,970,1948,1975]" captionStart="TABLE 1" captionStartId="3.[151,239,404,428]" captionText="TABLE 1. Morphometric data for Moenkhausia guaruba, new species, n = 30, range includes holotype. SD = standard deviation." pageId="2" pageNumber="588">Table 1</tableCitation>
. Largest examined specimen 
<quantity id="C1419B3FEA532675FA8AD3AD5009F878" box="[1299,1399,1948,1975]" metricMagnitude="-2" metricUnit="m" metricValue="9.69" pageId="2" pageNumber="588" unit="mm" value="96.9">96.9 mm</quantity>
SL (
<typeStatus id="D9028878EA532675FF06D3F15476F815" box="[159,264,1984,2010]" pageId="2" pageNumber="588">holotype</typeStatus>
). Body compressed and moderately deep (
<figureCitation id="9E822A5FEA532675FD79D3F1565FF814" box="[736,801,1984,2011]" captionStart="FIGURE 1" captionStartId="2.[151,250,684,708]" captionTargetBox="[378,1209,309,660]" captionTargetId="figure-81@2.[378,1209,309,660]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Moenkhausia guaruba, MZUSP 128235, 96.9 mm SL, Brazil, Pará, Novo Progresso, rio Braço Norte, rio Tapajós basin." figureDoi="http://doi.org/10.5281/zenodo.8255380" httpUri="https://zenodo.org/record/8255380/files/figure.png" pageId="2" pageNumber="588">Fig. 1</figureCitation>
). Greatest body depth at dorsal-fin origin. Dorsal profile of head convex from snout tip to vertical through nostrils, and straight from that point to tip of supraoccipital spine. Dorsal profile of body convex from tip of supraoccipital spine to dorsal-fin origin, straight along dorsal-fin base, straight to slightly convex from dorsal-fin terminus to adipose fin origin, and concave from that point to anteriormost dorsal caudal-fin ray. Ventral profile of body convex from tip of lower jaw to pelvic-fin origin, approximately straight from pelvic-fin origin to urogenital papilla, straight along anal-fin base, and concave between last anal-fin ray to anteriormost ventral caudal-fin procurrent ray.
</paragraph>
<caption id="52C66652EA522674FF0ED5A5547CFE1F" pageId="3" pageNumber="589" startId="3.[151,239,404,428]" targetType="table">
<paragraph id="060636DAEA522674FF0ED5A5547CFE1F" blockId="3.[151,1436,404,464]" pageId="3" pageNumber="589">
<emphasis id="34CDEAC8EA522674FF0ED5A55475FE63" bold="true" box="[151,267,404,428]" pageId="3" pageNumber="589">TABLE 1.</emphasis>
Morphometric data for 
<taxonomicName id="C1B94D59EA522674FD96D5A5567EFE63" authority="Lima &amp; Vita &amp; Dutra &amp; Ohara &amp; Pastana, 2023" authorityName="Lima &amp; Vita &amp; Dutra &amp; Ohara &amp; Pastana" authorityYear="2023" box="[527,768,404,428]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="589" phylum="Chordata" rank="species" species="guaruba" status="new species">
<emphasis id="34CDEAC8EA522674FD96D5A5567EFE63" box="[527,768,404,428]" italics="true" pageId="3" pageNumber="589">Moenkhausia guaruba</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="2FFE57B3EA522674FC96D5A456EDFE63" box="[783,915,404,428]" pageId="3" pageNumber="589" rank="species">new species</taxonomicNameLabel>
, n = 30, range includes holotype. SD = standard deviation.
</paragraph>
</caption>
<paragraph id="060636DAEA522674FD72D5D05021F990" pageId="3" pageNumber="589">
<table id="74B9C47AEA52D988FF3AD5D15021F990" box="[163,1375,480,1631]" gridcols="5" gridrows="29" pageId="3" pageNumber="589">
<tr id="B8893498EA52D988FF3AD5D15021FE36" box="[163,1375,480,505]" gridrow="0" pageId="3" pageNumber="589" rowspan-0="1">
<th id="FB585DE4EA52D988FD72D5D15630FE36" box="[747,846,480,505]" gridcol="1" gridrow="0" pageId="3" pageNumber="589">
<emphasis id="34CDEAC8EA522674FD72D5D05630FE36" bold="true" box="[747,846,481,505]" pageId="3" pageNumber="589">Holotype</emphasis>
</th>
<th id="FB585DE4EA52D988FC35D5D15155FE36" box="[940,1067,480,505]" gridcol="2" gridrow="0" pageId="3" pageNumber="589">
<emphasis id="34CDEAC8EA522674FC50D5D05170FE36" bold="true" box="[969,1038,481,505]" pageId="3" pageNumber="589">Range</emphasis>
</th>
<th id="FB585DE4EA52D988FB09D5D151B0FE36" box="[1168,1230,480,505]" gridcol="3" gridrow="0" pageId="3" pageNumber="589">
<emphasis id="34CDEAC8EA522674FB09D5D051B0FE36" bold="true" box="[1168,1230,481,505]" pageId="3" pageNumber="589">Mean</emphasis>
</th>
<th id="FB585DE4EA52D988FAA6D5D15021FE36" box="[1343,1375,480,505]" gridcol="4" gridrow="0" pageId="3" pageNumber="589">
<emphasis id="34CDEAC8EA522674FAA6D5D15021FE37" bold="true" box="[1343,1375,480,504]" pageId="3" pageNumber="589">SD</emphasis>
</th>
</tr>
<tr id="B8893498EA52D988FF3AD6385021FDEE" box="[163,1375,521,545]" gridrow="1" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD638573DFDEE" box="[163,579,521,545]" gridcol="0" gridrow="1" pageId="3" pageNumber="589">Standard length (mm)</th>
<td id="FB585DE4EA52D988FD72D6385630FDEE" box="[747,846,521,545]" gridcol="1" gridrow="1" pageId="3" pageNumber="589">96.9</td>
<td id="FB585DE4EA52D988FC35D6385155FDEE" box="[940,1067,521,545]" gridcol="2" gridrow="1" pageId="3" pageNumber="589">44.4–96.9</td>
<td id="FB585DE4EA52D988FB09D63851B0FDEE" box="[1168,1230,521,545]" gridcol="3" gridrow="1" pageId="3" pageNumber="589">-</td>
<td id="FB585DE4EA52D988FAA6D6385021FDEE" box="[1343,1375,521,545]" gridcol="4" gridrow="1" pageId="3" pageNumber="589">-</td>
</tr>
<tr id="B8893498EA52D988FF3AD6015021FD86" box="[163,1375,560,585]" gridrow="2" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD6015021FD86" box="[163,1375,560,585]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="3" pageNumber="589">
<emphasis id="34CDEAC8EA522674FDEAD60056BFFD86" bold="true" box="[627,961,560,585]" pageId="3" pageNumber="589">Percentages of standard length</emphasis>
</th>
</tr>
<tr id="B8893498EA52D988FF3AD6685021FDBE" box="[163,1375,601,625]" gridrow="3" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD668573DFDBE" box="[163,579,601,625]" gridcol="0" gridrow="3" pageId="3" pageNumber="589">Depth at dorsal-fin origin</th>
<td id="FB585DE4EA52D988FD72D6685630FDBE" box="[747,846,601,625]" gridcol="1" gridrow="3" pageId="3" pageNumber="589">42.7</td>
<td id="FB585DE4EA52D988FC35D6685155FDBE" box="[940,1067,601,625]" gridcol="2" gridrow="3" pageId="3" pageNumber="589">37.8–46.1</td>
<td id="FB585DE4EA52D988FB09D66851B0FDBE" box="[1168,1230,601,625]" gridcol="3" gridrow="3" pageId="3" pageNumber="589">41.3</td>
<td id="FB585DE4EA52D988FAA6D6685021FDBE" box="[1343,1375,601,625]" gridcol="4" gridrow="3" pageId="3" pageNumber="589">2.0</td>
</tr>
<tr id="B8893498EA52D988FF3AD6B05021FD56" box="[163,1375,641,665]" gridrow="4" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD6B0573DFD56" box="[163,579,641,665]" gridcol="0" gridrow="4" pageId="3" pageNumber="589">Snout to dorsal-fin origin</th>
<td id="FB585DE4EA52D988FD72D6B05630FD56" box="[747,846,641,665]" gridcol="1" gridrow="4" pageId="3" pageNumber="589">53.1</td>
<td id="FB585DE4EA52D988FC35D6B05155FD56" box="[940,1067,641,665]" gridcol="2" gridrow="4" pageId="3" pageNumber="589">50.0–55.1</td>
<td id="FB585DE4EA52D988FB09D6B051B0FD56" box="[1168,1230,641,665]" gridcol="3" gridrow="4" pageId="3" pageNumber="589">51.7</td>
<td id="FB585DE4EA52D988FAA6D6B05021FD56" box="[1343,1375,641,665]" gridcol="4" gridrow="4" pageId="3" pageNumber="589">1.3</td>
</tr>
<tr id="B8893498EA52D988FF3AD6985021FD0D" box="[163,1375,681,706]" gridrow="5" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD698573DFD0D" box="[163,579,681,706]" gridcol="0" gridrow="5" pageId="3" pageNumber="589">Snout to pectoral-fin origin</th>
<td id="FB585DE4EA52D988FD72D6985630FD0D" box="[747,846,681,706]" gridcol="1" gridrow="5" pageId="3" pageNumber="589">28.9</td>
<td id="FB585DE4EA52D988FC35D6985155FD0D" box="[940,1067,681,706]" gridcol="2" gridrow="5" pageId="3" pageNumber="589">26.4–30.9</td>
<td id="FB585DE4EA52D988FB09D69851B0FD0D" box="[1168,1230,681,706]" gridcol="3" gridrow="5" pageId="3" pageNumber="589">28.9</td>
<td id="FB585DE4EA52D988FAA6D6985021FD0D" box="[1343,1375,681,706]" gridcol="4" gridrow="5" pageId="3" pageNumber="589">1.1</td>
</tr>
<tr id="B8893498EA52D988FF3AD6E35021FD25" box="[163,1375,722,746]" gridrow="6" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD6E3573DFD25" box="[163,579,722,746]" gridcol="0" gridrow="6" pageId="3" pageNumber="589">Snout to pelvic-fin origin</th>
<td id="FB585DE4EA52D988FD72D6E35630FD25" box="[747,846,722,746]" gridcol="1" gridrow="6" pageId="3" pageNumber="589">49.9</td>
<td id="FB585DE4EA52D988FC35D6E35155FD25" box="[940,1067,722,746]" gridcol="2" gridrow="6" pageId="3" pageNumber="589">46.2–53.0</td>
<td id="FB585DE4EA52D988FB09D6E351B0FD25" box="[1168,1230,722,746]" gridcol="3" gridrow="6" pageId="3" pageNumber="589">49.3</td>
<td id="FB585DE4EA52D988FAA6D6E35021FD25" box="[1343,1375,722,746]" gridcol="4" gridrow="6" pageId="3" pageNumber="589">1.4</td>
</tr>
<tr id="B8893498EA52D988FF3AD6CB5021FCDD" box="[163,1375,762,786]" gridrow="7" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD6CB573DFCDD" box="[163,579,762,786]" gridcol="0" gridrow="7" pageId="3" pageNumber="589">Snout to anal-fin origin</th>
<td id="FB585DE4EA52D988FD72D6CB5630FCDD" box="[747,846,762,786]" gridcol="1" gridrow="7" pageId="3" pageNumber="589">68.4</td>
<td id="FB585DE4EA52D988FC35D6CB5155FCDD" box="[940,1067,762,786]" gridcol="2" gridrow="7" pageId="3" pageNumber="589">63.6–68.5</td>
<td id="FB585DE4EA52D988FB09D6CB51B0FCDD" box="[1168,1230,762,786]" gridcol="3" gridrow="7" pageId="3" pageNumber="589">66.0</td>
<td id="FB585DE4EA52D988FAA6D6CB5021FCDD" box="[1343,1375,762,786]" gridcol="4" gridrow="7" pageId="3" pageNumber="589">1.3</td>
</tr>
<tr id="B8893498EA52D988FF3AD7135021FCF5" box="[163,1375,802,826]" gridrow="8" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD713573DFCF5" box="[163,579,802,826]" gridcol="0" gridrow="8" pageId="3" pageNumber="589">Caudal peduncle depth</th>
<td id="FB585DE4EA52D988FD72D7135630FCF5" box="[747,846,802,826]" gridcol="1" gridrow="8" pageId="3" pageNumber="589">10.9</td>
<td id="FB585DE4EA52D988FC35D7135155FCF5" box="[940,1067,802,826]" gridcol="2" gridrow="8" pageId="3" pageNumber="589">10.2–12.2</td>
<td id="FB585DE4EA52D988FB09D71351B0FCF5" box="[1168,1230,802,826]" gridcol="3" gridrow="8" pageId="3" pageNumber="589">11.4</td>
<td id="FB585DE4EA52D988FAA6D7135021FCF5" box="[1343,1375,802,826]" gridcol="4" gridrow="8" pageId="3" pageNumber="589">0.5</td>
</tr>
<tr id="B8893498EA52D988FF3AD77B5021FCAC" box="[163,1375,842,867]" gridrow="9" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD77B573DFCAC" box="[163,579,842,867]" gridcol="0" gridrow="9" pageId="3" pageNumber="589">Caudal peduncle length</th>
<td id="FB585DE4EA52D988FD72D77B5630FCAC" box="[747,846,842,867]" gridcol="1" gridrow="9" pageId="3" pageNumber="589">13.1</td>
<td id="FB585DE4EA52D988FC35D77B5155FCAC" box="[940,1067,842,867]" gridcol="2" gridrow="9" pageId="3" pageNumber="589">10.6–13.8</td>
<td id="FB585DE4EA52D988FB09D77B51B0FCAC" box="[1168,1230,842,867]" gridcol="3" gridrow="9" pageId="3" pageNumber="589">12.4</td>
<td id="FB585DE4EA52D988FAA6D77B5021FCAC" box="[1343,1375,842,867]" gridcol="4" gridrow="9" pageId="3" pageNumber="589">0.8</td>
</tr>
<tr id="B8893498EA52D988FF3AD7425021FC44" box="[163,1375,883,907]" gridrow="10" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD742573DFC44" box="[163,579,883,907]" gridcol="0" gridrow="10" pageId="3" pageNumber="589">Pectoral-fin length</th>
<td id="FB585DE4EA52D988FD72D7425630FC44" box="[747,846,883,907]" gridcol="1" gridrow="10" pageId="3" pageNumber="589">19.8</td>
<td id="FB585DE4EA52D988FC35D7425155FC44" box="[940,1067,883,907]" gridcol="2" gridrow="10" pageId="3" pageNumber="589">19.6–22.7</td>
<td id="FB585DE4EA52D988FB09D74251B0FC44" box="[1168,1230,883,907]" gridcol="3" gridrow="10" pageId="3" pageNumber="589">20.9</td>
<td id="FB585DE4EA52D988FAA6D7425021FC44" box="[1343,1375,883,907]" gridcol="4" gridrow="10" pageId="3" pageNumber="589">0.9</td>
</tr>
<tr id="B8893498EA52D988FF3AD7AA5021FC7C" box="[163,1375,923,947]" gridrow="11" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD7AA573DFC7C" box="[163,579,923,947]" gridcol="0" gridrow="11" pageId="3" pageNumber="589">Pelvic-fin length</th>
<td id="FB585DE4EA52D988FD72D7AA5630FC7C" box="[747,846,923,947]" gridcol="1" gridrow="11" pageId="3" pageNumber="589">15.7</td>
<td id="FB585DE4EA52D988FC35D7AA5155FC7C" box="[940,1067,923,947]" gridcol="2" gridrow="11" pageId="3" pageNumber="589">12.0–17.6</td>
<td id="FB585DE4EA52D988FB09D7AA51B0FC7C" box="[1168,1230,923,947]" gridcol="3" gridrow="11" pageId="3" pageNumber="589">16.2</td>
<td id="FB585DE4EA52D988FAA6D7AA5021FC7C" box="[1343,1375,923,947]" gridcol="4" gridrow="11" pageId="3" pageNumber="589">1.1</td>
</tr>
<tr id="B8893498EA52D988FF3AD7F25021FC14" box="[163,1375,963,987]" gridrow="12" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD7F2573DFC14" box="[163,579,963,987]" gridcol="0" gridrow="12" pageId="3" pageNumber="589">Pelvic-fin insertion to anal-fin origin</th>
<td id="FB585DE4EA52D988FD72D7F25630FC14" box="[747,846,963,987]" gridcol="1" gridrow="12" pageId="3" pageNumber="589">20.1</td>
<td id="FB585DE4EA52D988FC35D7F25155FC14" box="[940,1067,963,987]" gridcol="2" gridrow="12" pageId="3" pageNumber="589">17.1–21.6</td>
<td id="FB585DE4EA52D988FB09D7F251B0FC14" box="[1168,1230,963,987]" gridcol="3" gridrow="12" pageId="3" pageNumber="589">19.0</td>
<td id="FB585DE4EA52D988FAA6D7F25021FC14" box="[1343,1375,963,987]" gridcol="4" gridrow="12" pageId="3" pageNumber="589">1.1</td>
</tr>
<tr id="B8893498EA52D988FF3AD7DA5021FBCB" box="[163,1375,1003,1028]" gridrow="13" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD7DA573DFBCB" box="[163,579,1003,1028]" gridcol="0" gridrow="13" pageId="3" pageNumber="589">Dorsal-fin origin to anal-fin origin</th>
<td id="FB585DE4EA52D988FD72D7DA5630FBCB" box="[747,846,1003,1028]" gridcol="1" gridrow="13" pageId="3" pageNumber="589">42.5</td>
<td id="FB585DE4EA52D988FC35D7DA5155FBCB" box="[940,1067,1003,1028]" gridcol="2" gridrow="13" pageId="3" pageNumber="589">38.7–46.2</td>
<td id="FB585DE4EA52D988FB09D7DA51B0FBCB" box="[1168,1230,1003,1028]" gridcol="3" gridrow="13" pageId="3" pageNumber="589">42.6</td>
<td id="FB585DE4EA52D988FAA6D7DA5021FBCB" box="[1343,1375,1003,1028]" gridcol="4" gridrow="13" pageId="3" pageNumber="589">1.9</td>
</tr>
<tr id="B8893498EA52D988FF3AD0255021FBE3" box="[163,1375,1044,1068]" gridrow="14" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD025573DFBE3" box="[163,579,1044,1068]" gridcol="0" gridrow="14" pageId="3" pageNumber="589">Dorsal-fin origin to pelvic-fin insertion</th>
<td id="FB585DE4EA52D988FD72D0255630FBE3" box="[747,846,1044,1068]" gridcol="1" gridrow="14" pageId="3" pageNumber="589">43.0</td>
<td id="FB585DE4EA52D988FC35D0255155FBE3" box="[940,1067,1044,1068]" gridcol="2" gridrow="14" pageId="3" pageNumber="589">37.6–46.3</td>
<td id="FB585DE4EA52D988FB09D02551B0FBE3" box="[1168,1230,1044,1068]" gridcol="3" gridrow="14" pageId="3" pageNumber="589">41.4</td>
<td id="FB585DE4EA52D988FAA6D0255021FBE3" box="[1343,1375,1044,1068]" gridcol="4" gridrow="14" pageId="3" pageNumber="589">2.1</td>
</tr>
<tr id="B8893498EA52D988FF3AD00D5021FB9B" box="[163,1375,1084,1108]" gridrow="15" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD00D573DFB9B" box="[163,579,1084,1108]" gridcol="0" gridrow="15" pageId="3" pageNumber="589">Dorsal-fin origin to pectoral-fin insertion</th>
<td id="FB585DE4EA52D988FD72D00D5630FB9B" box="[747,846,1084,1108]" gridcol="1" gridrow="15" pageId="3" pageNumber="589">46.1</td>
<td id="FB585DE4EA52D988FC35D00D5155FB9B" box="[940,1067,1084,1108]" gridcol="2" gridrow="15" pageId="3" pageNumber="589">39.6–46.1</td>
<td id="FB585DE4EA52D988FB09D00D51B0FB9B" box="[1168,1230,1084,1108]" gridcol="3" gridrow="15" pageId="3" pageNumber="589">42.9</td>
<td id="FB585DE4EA52D988FAA6D00D5021FB9B" box="[1343,1375,1084,1108]" gridcol="4" gridrow="15" pageId="3" pageNumber="589">1.6</td>
</tr>
<tr id="B8893498EA52D988FF3AD0555021FBB3" box="[163,1375,1124,1148]" gridrow="16" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD055573DFBB3" box="[163,579,1124,1148]" gridcol="0" gridrow="16" pageId="3" pageNumber="589">Dorsal-fin length</th>
<td id="FB585DE4EA52D988FD72D0555630FBB3" box="[747,846,1124,1148]" gridcol="1" gridrow="16" pageId="3" pageNumber="589">25.0</td>
<td id="FB585DE4EA52D988FC35D0555155FBB3" box="[940,1067,1124,1148]" gridcol="2" gridrow="16" pageId="3" pageNumber="589">24.4–28.7</td>
<td id="FB585DE4EA52D988FB09D05551B0FBB3" box="[1168,1230,1124,1148]" gridcol="3" gridrow="16" pageId="3" pageNumber="589">26.4</td>
<td id="FB585DE4EA52D988FAA6D0555021FBB3" box="[1343,1375,1124,1148]" gridcol="4" gridrow="16" pageId="3" pageNumber="589">1.1</td>
</tr>
<tr id="B8893498EA52D988FF3AD0BD5021FB6A" box="[163,1375,1164,1189]" gridrow="17" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD0BD573DFB6A" box="[163,579,1164,1189]" gridcol="0" gridrow="17" pageId="3" pageNumber="589">Dorsal-fin base length</th>
<td id="FB585DE4EA52D988FD72D0BD5630FB6A" box="[747,846,1164,1189]" gridcol="1" gridrow="17" pageId="3" pageNumber="589">13.3</td>
<td id="FB585DE4EA52D988FC35D0BD5155FB6A" box="[940,1067,1164,1189]" gridcol="2" gridrow="17" pageId="3" pageNumber="589">11.5–15.7</td>
<td id="FB585DE4EA52D988FB09D0BD51B0FB6A" box="[1168,1230,1164,1189]" gridcol="3" gridrow="17" pageId="3" pageNumber="589">14.0</td>
<td id="FB585DE4EA52D988FAA6D0BD5021FB6A" box="[1343,1375,1164,1189]" gridcol="4" gridrow="17" pageId="3" pageNumber="589">0.9</td>
</tr>
<tr id="B8893498EA52D988FF3AD0855021FB02" box="[163,1375,1204,1229]" gridrow="18" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD085573DFB02" box="[163,579,1204,1229]" gridcol="0" gridrow="18" pageId="3" pageNumber="589">Anal-fin length</th>
<td id="FB585DE4EA52D988FD72D0855630FB02" box="[747,846,1204,1229]" gridcol="1" gridrow="18" pageId="3" pageNumber="589">15.1</td>
<td id="FB585DE4EA52D988FC35D0855155FB02" box="[940,1067,1204,1229]" gridcol="2" gridrow="18" pageId="3" pageNumber="589">14.6–19.3</td>
<td id="FB585DE4EA52D988FB09D08551B0FB02" box="[1168,1230,1204,1229]" gridcol="3" gridrow="18" pageId="3" pageNumber="589">17.2</td>
<td id="FB585DE4EA52D988FAA6D0855021FB02" box="[1343,1375,1204,1229]" gridcol="4" gridrow="18" pageId="3" pageNumber="589">1.3</td>
</tr>
<tr id="B8893498EA52D988FF3AD0EC5021FB3A" box="[163,1375,1245,1269]" gridrow="19" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD0EC573DFB3A" box="[163,579,1245,1269]" gridcol="0" gridrow="19" pageId="3" pageNumber="589">Anal-fin base length</th>
<td id="FB585DE4EA52D988FD72D0EC5630FB3A" box="[747,846,1245,1269]" gridcol="1" gridrow="19" pageId="3" pageNumber="589">31.0</td>
<td id="FB585DE4EA52D988FC35D0EC5155FB3A" box="[940,1067,1245,1269]" gridcol="2" gridrow="19" pageId="3" pageNumber="589">28.7–31.7</td>
<td id="FB585DE4EA52D988FB09D0EC51B0FB3A" box="[1168,1230,1245,1269]" gridcol="3" gridrow="19" pageId="3" pageNumber="589">30.3</td>
<td id="FB585DE4EA52D988FAA6D0EC5021FB3A" box="[1343,1375,1245,1269]" gridcol="4" gridrow="19" pageId="3" pageNumber="589">0.8</td>
</tr>
<tr id="B8893498EA52D988FF3AD1345021FAD1" box="[163,1375,1285,1310]" gridrow="20" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD134573DFAD1" box="[163,579,1285,1310]" gridcol="0" gridrow="20" pageId="3" pageNumber="589">Eye to dorsal-fin origin</th>
<td id="FB585DE4EA52D988FD72D1345630FAD1" box="[747,846,1285,1310]" gridcol="1" gridrow="20" pageId="3" pageNumber="589">40.9</td>
<td id="FB585DE4EA52D988FC35D1345155FAD1" box="[940,1067,1285,1310]" gridcol="2" gridrow="20" pageId="3" pageNumber="589">35.9–41.6</td>
<td id="FB585DE4EA52D988FB09D13451B0FAD1" box="[1168,1230,1285,1310]" gridcol="3" gridrow="20" pageId="3" pageNumber="589">38.4</td>
<td id="FB585DE4EA52D988FAA6D1345021FAD1" box="[1343,1375,1285,1310]" gridcol="4" gridrow="20" pageId="3" pageNumber="589">1.3</td>
</tr>
<tr id="B8893498EA52D988FF3AD11C5021FA8A" box="[163,1375,1325,1349]" gridrow="21" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD11C573DFA8A" box="[163,579,1325,1349]" gridcol="0" gridrow="21" pageId="3" pageNumber="589">Dorsal-fin origin to caudal-fin base</th>
<td id="FB585DE4EA52D988FD72D11C5630FA8A" box="[747,846,1325,1349]" gridcol="1" gridrow="21" pageId="3" pageNumber="589">55.5</td>
<td id="FB585DE4EA52D988FC35D11C5155FA8A" box="[940,1067,1325,1349]" gridcol="2" gridrow="21" pageId="3" pageNumber="589">52.7–58.3</td>
<td id="FB585DE4EA52D988FB09D11C51B0FA8A" box="[1168,1230,1325,1349]" gridcol="3" gridrow="21" pageId="3" pageNumber="589">55.4</td>
<td id="FB585DE4EA52D988FAA6D11C5021FA8A" box="[1343,1375,1325,1349]" gridcol="4" gridrow="21" pageId="3" pageNumber="589">1.6</td>
</tr>
<tr id="B8893498EA52D988FF3AD1645021FAA2" box="[163,1375,1365,1389]" gridrow="22" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD1645021FAA2" box="[163,1375,1365,1389]" colspan="5" colspanRight="4" gridcol="0" gridrow="22" pageId="3" pageNumber="589">
<emphasis id="34CDEAC8EA522674FD10D16456D4FAA2" bold="true" box="[649,938,1365,1389]" pageId="3" pageNumber="589">Percentages of head length</emphasis>
</th>
</tr>
<tr id="B8893498EA52D988FF3AD14C5021FA59" box="[163,1375,1405,1430]" gridrow="23" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD14C573DFA59" box="[163,579,1405,1430]" gridcol="0" gridrow="23" pageId="3" pageNumber="589">Head length</th>
<td id="FB585DE4EA52D988FD72D14C5630FA59" box="[747,846,1405,1430]" gridcol="1" gridrow="23" pageId="3" pageNumber="589">28.4</td>
<td id="FB585DE4EA52D988FC35D14C5155FA59" box="[940,1067,1405,1430]" gridcol="2" gridrow="23" pageId="3" pageNumber="589">26.4–30.3</td>
<td id="FB585DE4EA52D988FB09D14C51B0FA59" box="[1168,1230,1405,1430]" gridcol="3" gridrow="23" pageId="3" pageNumber="589">28.4</td>
<td id="FB585DE4EA52D988FAA6D14C5021FA59" box="[1343,1375,1405,1430]" gridcol="4" gridrow="23" pageId="3" pageNumber="589">1.0</td>
</tr>
<tr id="B8893498EA52D988FF3AD1945021FA71" box="[163,1375,1445,1470]" gridrow="24" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD194573DFA71" box="[163,579,1445,1470]" gridcol="0" gridrow="24" pageId="3" pageNumber="589">Head depth</th>
<td id="FB585DE4EA52D988FD72D1945630FA71" box="[747,846,1445,1470]" gridcol="1" gridrow="24" pageId="3" pageNumber="589">125.5</td>
<td id="FB585DE4EA52D988FC35D1945155FA71" box="[940,1067,1445,1470]" gridcol="2" gridrow="24" pageId="3" pageNumber="589">107.9–126.2</td>
<td id="FB585DE4EA52D988FB09D19451B0FA71" box="[1168,1230,1445,1470]" gridcol="3" gridrow="24" pageId="3" pageNumber="589">115.4</td>
<td id="FB585DE4EA52D988FAA6D1945021FA71" box="[1343,1375,1445,1470]" gridcol="4" gridrow="24" pageId="3" pageNumber="589">4.4</td>
</tr>
<tr id="B8893498EA52D988FF3AD1FF5021FA29" box="[163,1375,1486,1510]" gridrow="25" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD1FF573DFA29" box="[163,579,1486,1510]" gridcol="0" gridrow="25" pageId="3" pageNumber="589">Horizontal eye diameter</th>
<td id="FB585DE4EA52D988FD72D1FF5630FA29" box="[747,846,1486,1510]" gridcol="1" gridrow="25" pageId="3" pageNumber="589">32.1</td>
<td id="FB585DE4EA52D988FC35D1FF5155FA29" box="[940,1067,1486,1510]" gridcol="2" gridrow="25" pageId="3" pageNumber="589">32.1–40.0</td>
<td id="FB585DE4EA52D988FB09D1FF51B0FA29" box="[1168,1230,1486,1510]" gridcol="3" gridrow="25" pageId="3" pageNumber="589">35.7</td>
<td id="FB585DE4EA52D988FAA6D1FF5021FA29" box="[1343,1375,1486,1510]" gridcol="4" gridrow="25" pageId="3" pageNumber="589">2.3</td>
</tr>
<tr id="B8893498EA52D988FF3AD1C75021F9C1" box="[163,1375,1526,1550]" gridrow="26" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD1C7573DF9C1" box="[163,579,1526,1550]" gridcol="0" gridrow="26" pageId="3" pageNumber="589">Snout length</th>
<td id="FB585DE4EA52D988FD72D1C75630F9C1" box="[747,846,1526,1550]" gridcol="1" gridrow="26" pageId="3" pageNumber="589">25.2</td>
<td id="FB585DE4EA52D988FC35D1C75155F9C1" box="[940,1067,1526,1550]" gridcol="2" gridrow="26" pageId="3" pageNumber="589">19.9–26.3</td>
<td id="FB585DE4EA52D988FB09D1C751B0F9C1" box="[1168,1230,1526,1550]" gridcol="3" gridrow="26" pageId="3" pageNumber="589">23.2</td>
<td id="FB585DE4EA52D988FAA6D1C75021F9C1" box="[1343,1375,1526,1550]" gridcol="4" gridrow="26" pageId="3" pageNumber="589">1.8</td>
</tr>
<tr id="B8893498EA52D988FF3AD22F5021F9F8" box="[163,1375,1566,1591]" gridrow="27" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD22F573DF9F8" box="[163,579,1566,1591]" gridcol="0" gridrow="27" pageId="3" pageNumber="589">Interorbital width</th>
<td id="FB585DE4EA52D988FD72D22F5630F9F8" box="[747,846,1566,1591]" gridcol="1" gridrow="27" pageId="3" pageNumber="589">36.7</td>
<td id="FB585DE4EA52D988FC35D22F5155F9F8" box="[940,1067,1566,1591]" gridcol="2" gridrow="27" pageId="3" pageNumber="589">32.0–39.1</td>
<td id="FB585DE4EA52D988FB09D22F51B0F9F8" box="[1168,1230,1566,1591]" gridcol="3" gridrow="27" pageId="3" pageNumber="589">35.8</td>
<td id="FB585DE4EA52D988FAA6D22F5021F9F8" box="[1343,1375,1566,1591]" gridcol="4" gridrow="27" pageId="3" pageNumber="589">1.6</td>
</tr>
<tr id="B8893498EA52D988FF3AD2765021F990" box="[163,1375,1607,1631]" gridrow="28" pageId="3" pageNumber="589">
<th id="FB585DE4EA52D988FF3AD276573DF990" box="[163,579,1607,1631]" gridcol="0" gridrow="28" pageId="3" pageNumber="589">Upper jaw length</th>
<td id="FB585DE4EA52D988FD72D2765630F990" box="[747,846,1607,1631]" gridcol="1" gridrow="28" pageId="3" pageNumber="589">47.6</td>
<td id="FB585DE4EA52D988FC35D2765155F990" box="[940,1067,1607,1631]" gridcol="2" gridrow="28" pageId="3" pageNumber="589">42.8–49.6</td>
<td id="FB585DE4EA52D988FB09D27651B0F990" box="[1168,1230,1607,1631]" gridcol="3" gridrow="28" pageId="3" pageNumber="589">46.0</td>
<td id="FB585DE4EA52D988FAA6D2765021F990" box="[1343,1375,1607,1631]" gridcol="4" gridrow="28" pageId="3" pageNumber="589">1.7</td>
</tr>
</table>
</paragraph>
<paragraph id="060636DAEA522674FF5ED2855797F891" blockId="3.[151,1437,1716,1995]" pageId="3" pageNumber="589">
Mouth terminal, posterior end of maxilla not reaching vertical through the middle of the orbit. Premaxillary teeth arranged into two rows (
<figureCitation id="9E822A5FEA522674FE72D2E9574EF93C" box="[491,560,1752,1779]" captionStart="FIGURE 3" captionStartId="4.[151,250,932,956]" captionTargetBox="[414,1165,189,904]" captionTargetId="figure-19@4.[408,1179,181,908]" captionTargetPageId="4" captionText="FIGURE 3. Moenkhausia guaruba, paratype, MZUSP 119389, 70.9 mm SL. Lateral view of premaxilla, maxilla and dentary, left side." figureDoi="http://doi.org/10.5281/zenodo.8255384" httpUri="https://zenodo.org/record/8255384/files/figure.png" pageId="3" pageNumber="589">Fig. 3</figureCitation>
), outer row with 3(1), 4*(25) or 5(4) tricuspid* (29) or pentacuspid (1) teeth. Inner row with 4(1), 5* (27), 6(1) or 7(1) pentacuspid (21) or heptacuspid teeth* (9). Maxilla with 1(2), 2* (23), 3(4) or 4(1) tricuspid (20) or pentacuspid* (10) teeth. Dentary teeth with 4*(25) large pentacuspid (23) or heptacuspid (2) teeth followed by 12(2) conical to tricuspid teeth.
</paragraph>
<paragraph id="060636DAEA522674FF5ED35957BCF805" blockId="3.[151,1437,1716,1995]" pageId="3" pageNumber="589">Hypobranchial with 1(1), or 3(1) rakers, ceratobranchials with 10(2) rakers, epibranchials with 8(1), or 9(1) rakers, and one raker between those last two elements. Branchiostegals 4(2), three articulating with the anterior ceratohyal, and one with the posterior ceratohyal.</paragraph>
<caption id="52C66652EA552673FF0ED7955591FC2F" ID-DOI="http://doi.org/10.5281/zenodo.8255384" ID-Zenodo-Dep="8255384" httpUri="https://zenodo.org/record/8255384/files/figure.png" pageId="4" pageNumber="590" startId="4.[151,250,932,956]" targetBox="[414,1165,189,904]" targetPageId="4" targetType="figure">
<paragraph id="060636DAEA552673FF0ED7955591FC2F" blockId="4.[151,1436,932,992]" pageId="4" pageNumber="590">
<emphasis id="34CDEAC8EA552673FF0ED795546DFC73" bold="true" box="[151,275,932,956]" pageId="4" pageNumber="590">FIGURE 3.</emphasis>
<taxonomicName id="C1B94D59EA552673FE82D795577EFC73" authorityName="Lima &amp; Vita &amp; Dutra &amp; Ohara &amp; Pastana" authorityYear="2023" box="[283,512,932,956]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="590" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA552673FE82D795577EFC73" box="[283,512,932,956]" italics="true" pageId="4" pageNumber="590">Moenkhausia guaruba</emphasis>
</taxonomicName>
, paratype, MZUSP 119389, 70.9 mm SL. Lateral view of premaxilla, maxilla and dentary, left side.
</paragraph>
</caption>
<caption id="52C66652EA552673FF0ED15E5488FA64" ID-DOI="http://doi.org/10.5281/zenodo.8255386" ID-Zenodo-Dep="8255386" httpUri="https://zenodo.org/record/8255386/files/figure.png" pageId="4" pageNumber="590" startId="4.[151,250,1391,1415]" targetBox="[411,1177,1021,1363]" targetPageId="4" targetType="figure">
<paragraph id="060636DAEA552673FF0ED15E5488FA64" blockId="4.[151,1436,1391,1451]" pageId="4" pageNumber="590">
<emphasis id="34CDEAC8EA552673FF0ED15E546AFA48" bold="true" box="[151,276,1391,1415]" pageId="4" pageNumber="590">FIGURE 4.</emphasis>
Live specimen of 
<taxonomicName id="C1B94D59EA552673FE4AD15E5664FA48" authority=", MZUSP" authorityName="MZUSP" box="[467,794,1391,1415]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="590" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA552673FE4AD15E57C7FA48" box="[467,697,1391,1415]" italics="true" pageId="4" pageNumber="590">Moenkhausia guaruba</emphasis>
, MZUSP
</taxonomicName>
119389, paratype, SL uncertain, Brazil, Pará, Novo Progresso, rio Braço Norte, rio Tapajós basin.
</paragraph>
</caption>
<paragraph id="060636DAEA552673FF5ED1CC54EDF967" blockId="4.[151,1437,1533,1991]" pageId="4" pageNumber="590">
Scales cycloid, without 
<emphasis id="34CDEAC8EA552673FE54D1CC576BF9D8" box="[461,533,1533,1559]" italics="true" pageId="4" pageNumber="590">circuli</emphasis>
on exposed portion of scales and with seven divergent 
<emphasis id="34CDEAC8EA552673FBE9D1CC51D8F9D8" box="[1136,1190,1533,1559]" italics="true" pageId="4" pageNumber="590">radii</emphasis>
reaching the posterior margin of the scale. Longitudinal line with 43*(7), 44(10), 45(2), or 46(2) perforated scales. Scale rows above lateral line 8(7) or 9*(23). Scale rows below lateral line 7*(27) or 8(2). Predorsal scales 12(7), 13(8), or 14*(1). Circumpeduncular scales 16*(18) or 17(1). Scales on caudal fin rays restricted to base of upper and lower lobes. Axillary scale present.
</paragraph>
<paragraph id="060636DAEA552673FF5ED2805672F808" blockId="4.[151,1437,1533,1991]" pageId="4" pageNumber="590">
Pectoral-fin rays i,11(1), i,12(19), or i,13*(10), longest ray reaching vertical through pelvic-fin insertion. Pelvic-fin rays i,7(26) or i,8*(4), distal tip of pelvic fin nearly straight, not reaching anal-fin origin. Dorsal-fin rays ii,8(5), ii,9*(24), or ii,10(1). First dorsal-fin pterygiophore inserted posteriorly to ninth neural spine (2). Anal-fin rays iv,21(1), iv,22(7), iv,23(10), iii,24*(9), or iv, 25(3), first unbranched ray covered by skin and last unbranched and first branched anal-fin ray the longest. First anal-fin pterygiophore inserted posteriorly to 18 
<superScript id="F1CC9B92EA552673FB69D3705183F880" attach="left" box="[1264,1277,1857,1871]" fontSize="6" pageId="4" pageNumber="590">th</superScript>
haemal spine (2). Caudal fin forked, with lobes equal in size, slightly pointed. Caudal-fin rays i,17,i*(29). Dorsal caudal-fin procurrent rays 11(1), or 12(1). Ventral caudal-fin procurrent rays 9(1), or 13(1). Adipose fin present. Supraneurals 4(2). Precaudal vertebrae 18(2). Caudal vertebrae 18(2).
</paragraph>
<caption id="52C66652EA542672FF0ED0685031FBBE" ID-DOI="http://doi.org/10.5281/zenodo.8255388" ID-Zenodo-Dep="8255388" box="[151,1359,1113,1137]" httpUri="https://zenodo.org/record/8255388/files/figure.png" pageId="5" pageNumber="591" startId="5.[151,250,1113,1137]" targetBox="[157,1432,186,1086]" targetPageId="5" targetType="figure">
<paragraph id="060636DAEA542672FF0ED0685031FBBE" blockId="5.[151,1359,1113,1137]" box="[151,1359,1113,1137]" pageId="5" pageNumber="591">
<emphasis id="34CDEAC8EA542672FF0ED068546DFBBE" bold="true" box="[151,275,1113,1137]" pageId="5" pageNumber="591">FIGURE 5.</emphasis>
Distribution of 
<taxonomicName id="C1B94D59EA542672FE2FD06857E4FBBE" box="[438,666,1113,1137]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="591" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA542672FE2FD06857E4FBBE" box="[438,666,1113,1137]" italics="true" pageId="5" pageNumber="591">Moenkhausia guaruba</emphasis>
</taxonomicName>
at Pará State, Brazil. Red star = holotype, white circles = paratypes.
</paragraph>
</caption>
<caption id="52C66652EA542672FF0ED20F5446F951" ID-DOI="http://doi.org/10.5281/zenodo.8255390" ID-Zenodo-Dep="8255390" httpUri="https://zenodo.org/record/8255390/files/figure.png" pageId="5" pageNumber="591" startId="5.[151,250,1598,1622]" targetBox="[379,1209,1161,1574]" targetPageId="5" targetType="figure">
<paragraph id="060636DAEA542672FF0ED20F5446F951" blockId="5.[151,1436,1598,1694]" pageId="5" pageNumber="591">
<emphasis id="34CDEAC8EA542672FF0ED20F546DF999" bold="true" box="[151,275,1598,1622]" pageId="5" pageNumber="591">FIGURE 6.</emphasis>
Type locality of 
<taxonomicName id="C1B94D59EA542672FE27D20F57DDF999" box="[446,675,1598,1622]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="591" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA542672FE27D20F57DDF999" box="[446,675,1598,1622]" italics="true" pageId="5" pageNumber="591">Moenkhausia guaruba</emphasis>
</taxonomicName>
at up 
<emphasis id="34CDEAC8EA542672FD43D20F5796F999" bold="true" box="[730,744,1598,1622]" pageId="5" pageNumber="591">p</emphasis>
er rio Braço Norte at Serra do Cachimbo, tributary of rio Teles Pires, rio Tapajós basin, Pará State, Brazil: (a) waterfall upstream, substrate composed mainly by rocks; (b) sandy beach downstream to the waterfall.
</paragraph>
</caption>
<paragraph id="060636DAEA542672FF5ED2D55172F836" blockId="5.[151,1437,1763,2041]" pageId="5" pageNumber="591">
Laterosensory system fully developed, tubules opening in individual pores (
<emphasis id="34CDEAC8EA542672FBD0D2D55112F932" box="[1097,1132,1764,1789]" italics="true" pageId="5" pageNumber="591">i.e.</emphasis>
never forming subpores). Supraorbital canal associated with nasal, frontal, and parietal bones. Epiphyseal and parietal branches of supraorbital canal present. Supraorbital connecting to infraorbital and otic canal near posterolateral margin of frontal bone. Infraorbital canal fully developed. Otic canal present, associated with anterior portion of pterotic. Postotic canal running along pterotic, extrascapular, post-temporal and supracleithrum. Supratemporal canal associated with extrascapula and posterior margin of parietal bone, running dorsomedially and opening in a pore near cranial fontanel. Preopercular canal extending along entire extension of pre-opercular bone, continuous with mandibular canal. Mandibular canal associated with the anguloarticular and dentary bones.
</paragraph>
<paragraph id="060636DAEA572671FF5ED4A75682FD03" blockId="6.[151,1437,150,1544]" pageId="6" pageNumber="592">
<emphasis id="34CDEAC8EA572671FF5ED4A754F0FF7F" bold="true" box="[199,398,150,176]" pageId="6" pageNumber="592">Color in alcohol.</emphasis>
Ground body coloration pale brown to tan (
<figureCitation id="9E822A5FEA572671FCE4D4A756C0FF7F" box="[893,958,150,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,684,708]" captionTargetBox="[378,1209,309,660]" captionTargetId="figure-81@2.[378,1209,309,660]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Moenkhausia guaruba, MZUSP 128235, 96.9 mm SL, Brazil, Pará, Novo Progresso, rio Braço Norte, rio Tapajós basin." figureDoi="http://doi.org/10.5281/zenodo.8255380" httpUri="https://zenodo.org/record/8255380/files/figure.png" pageId="6" pageNumber="592">Fig. 1</figureCitation>
). Dorsal portion of head dark brown, with high concentration of chromatophores. Lateral and ventral portion of head pale brown. Guanine pigmentation and few large chromatophores present on infraorbitals, preopercle and opercle. Body tan on dorsal and dorsolateral portion, becoming cream ventrally, usually with few and sparse chromatophores. Humeral region with a single vertically elongated and relatively narrow dark blotch with diffuse margins, bordered anteriorly and posteriorly by unpigmented areas. Humeral blotch encompassing eight to 10 scales vertically and two to three scales horizontally. Humeral blotch wider and darker above lateral-line series of scales, becoming narrower, and fainter towards its ventral limit. Thin longitudinal line formed by dark pigmentation running along horizontal septum of body. Longitudinal line starting on vertical through first to third branched dorsal-fin rays, and extending to caudal peduncle. Broad brown to black blotch with diffuse margins located on posterior limit of caudal peduncle and base of middle caudal-fin rays. Caudal blotch pigmentation restricted to base of principal caudal-fin rays, usually not continuing onto middle caudal-fin rays to posterior margin of the fin. Caudal fin mostly dusky due to presence of scattered chromatophores along ray margins, and on inter-radial membranes. Dorsal, pectoral, and anal fins uniformly dusky, with dark chromatophores more concentrated along leading margin of rays and at interradial membrane. Pelvic fin pale to translucid, with few dark chromatophores distributed along ray margin, unbranched rays more pigmented than remaining ones. Adipose fin pale brown, with scattered chromatophores.
</paragraph>
<paragraph id="060636DAEA572671FF5ED6E757CEFBFB" blockId="6.[151,1437,150,1544]" pageId="6" pageNumber="592">
<emphasis id="34CDEAC8EA572671FF5ED6E7541FFD3E" bold="true" box="[199,353,726,753]" pageId="6" pageNumber="592">Color in life.</emphasis>
Ground coloration silvery, with golden to olive-green hue (
<figureCitation id="9E822A5FEA572671FB97D6E7512BFD3F" box="[1038,1109,726,752]" captionStart="FIGURE 4" captionStartId="4.[151,250,1391,1415]" captionTargetBox="[411,1177,1021,1363]" captionTargetId="figure-42@4.[407,1181,1016,1367]" captionTargetPageId="4" captionText="FIGURE 4. Live specimen of Moenkhausia guaruba, MZUSP 119389, paratype, SL uncertain, Brazil, Pará, Novo Progresso, rio Braço Norte, rio Tapajós basin." figureDoi="http://doi.org/10.5281/zenodo.8255386" httpUri="https://zenodo.org/record/8255386/files/figure.png" pageId="6" pageNumber="592">Fig. 4</figureCitation>
). Body darker on dorsal and dorsolateral regions and lighter on abdominal region. Head dark dorsally. Snout, jaws, first and second infraorbitals and gular area yellow to light orange. Infraorbitals three to six and opercular series silvery, with guanine pigmentation. Eyes golden yellow dorsally and silvery ventrally. Body mostly silvery on flanks, with olive-green hue dorsally and yellow to golden hues on lateral and ventral regions. Bright silvery to greenish stripe present, starting immediately posterior to humeral region and extending onto caudal blotch. Fins mostly yellow, coloration more pronounced on dorsal, adipose and caudal fins; paler on paired and anal fins. Anal fin more pigmented along first to sixth rays and inter-radial membrane; seventh to last anal-fin rays with faint yellow pigmentation restricted to basal portion of rays, inter-radial membranes dusky. Tip of unbranched and first branched dorsal and anal-fin rays lacking pigmentation. Black round blotch visible at base of caudal fin.
</paragraph>
<paragraph id="060636DAEA572671FF5ED00F579DFBB3" blockId="6.[151,1437,150,1544]" pageId="6" pageNumber="592">
<emphasis id="34CDEAC8EA572671FF5ED00F54C9FB97" bold="true" box="[199,439,1086,1112]" pageId="6" pageNumber="592">Sexual dimorphism.</emphasis>
The examined specimens of 
<taxonomicName id="C1B94D59EA572671FD67D00F5685FB97" box="[766,1019,1086,1112]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="592" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA572671FD67D00F5685FB97" box="[766,1019,1086,1112]" italics="true" pageId="6" pageNumber="592">Moenkhausia guaruba</emphasis>
</taxonomicName>
did not exhibit any secondary sexual dimorphism. Bony hooks on fins were not observed.
</paragraph>
<paragraph id="060636DAEA572671FF5ED0B7514BFB0A" blockId="6.[151,1437,150,1544]" pageId="6" pageNumber="592">
<emphasis id="34CDEAC8EA572671FF5ED0B7577DFB6F" bold="true" box="[199,515,1158,1184]" pageId="6" pageNumber="592">Geographical distribution.</emphasis>
<taxonomicName id="C1B94D59EA572671FD90D0B7567AFB6F" box="[521,772,1158,1184]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="592" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA572671FD90D0B7567AFB6F" box="[521,772,1158,1184]" italics="true" pageId="6" pageNumber="592">Moenkhausia guaruba</emphasis>
</taxonomicName>
is known from rio Braço Norte, a tributary of rio Teles Pires draining from the Serra do Cachimbo, rio Tapajós basin, Pará state, 
<collectingCountry id="7EAE764AEA572671FC16D09B56ADFB0B" box="[911,979,1194,1220]" name="Brazil" pageId="6" pageNumber="592">Brazil</collectingCountry>
(
<figureCitation id="9E822A5FEA572671FC7AD09B5159FB0A" box="[995,1063,1194,1221]" captionStart="FIGURE 5" captionStartId="5.[151,250,1113,1137]" captionTargetBox="[157,1432,186,1086]" captionTargetId="figure-21@5.[151,1436,181,1088]" captionTargetPageId="5" captionText="FIGURE 5. Distribution of Moenkhausia guaruba at Pará State, Brazil. Red star = holotype, white circles = paratypes." figureDoi="http://doi.org/10.5281/zenodo.8255388" httpUri="https://zenodo.org/record/8255388/files/figure.png" pageId="6" pageNumber="592">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="060636DAEA572671FF5ED0FF5675FAB7" blockId="6.[151,1437,150,1544]" pageId="6" pageNumber="592">
<emphasis id="34CDEAC8EA572671FF5ED0FF54F1FB27" bold="true" box="[199,399,1230,1256]" pageId="6" pageNumber="592">Ecological notes.</emphasis>
<materialsCitation id="B6D13C87EA572671FE0ED0FF5679FAB7" collectionCode="CPBV" elevation="445" location="Serra do Cachimbo" municipality="Campo de Provas Brigadeiro Velloso" pageId="6" pageNumber="592" specimenCount="1">
The 
<typeStatus id="D9028878EA572671FE52D0FE5485FB26" box="[459,507,1231,1257]" pageId="6" pageNumber="592">type</typeStatus>
locality of 
<taxonomicName id="C1B94D59EA572671FDE7D0FF5602FB27" box="[638,892,1230,1256]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="592" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA572671FDE7D0FF5602FB27" box="[638,892,1230,1256]" italics="true" pageId="6" pageNumber="592">Moenkhausia guaruba</emphasis>
</taxonomicName>
is at an altitude of 
<elevation id="8D94D1E9EA572671FBC0D0FF502BFB27" box="[1113,1365,1230,1257]" metricMagnitude="2" metricUnit="m" metricValue="4.45" pageId="6" pageNumber="592" unit="m" value="445.0">
<quantity id="C1419B3FEA572671FBC0D0FF51DFFB26" box="[1113,1185,1230,1257]" metricMagnitude="2" metricUnit="m" metricValue="4.45" pageId="6" pageNumber="592" unit="m" value="445.0">445 m</quantity>
above sea level
</elevation>
at the 
<location id="03666001EA572671FF0ED0C3540CFAC3" LSID="urn:lsid:plazi:treatment:8E1087CCEA502671FF0ED24F546AF9C7:03666001EA572671FF0ED0C3540CFAC3" box="[151,370,1266,1292]" municipality="Campo de Provas Brigadeiro Velloso" name="Serra do Cachimbo" pageId="6" pageNumber="592">Serra do Cachimbo</location>
, within the 
<collectingMunicipality id="E662ACA0EA572671FE6AD0C356EFFAC3" box="[499,913,1266,1293]" pageId="6" pageNumber="592">Campo de Provas Brigadeiro Velloso</collectingMunicipality>
(
<collectionCode id="60A8AE1FEA572671FC06D0C35690FAC3" box="[927,1006,1266,1292]" pageId="6" pageNumber="592">CPBV</collectionCode>
), an area under the management of the Brazilian Air Force. The sampled stretch was immediately downstream to a 3-meter waterfall (
<figureCitation id="9E822A5FEA572671FB5FD1275066FAFF" box="[1222,1304,1302,1328]" captionStart="FIGURE 6" captionStartId="5.[151,250,1598,1622]" captionTargetBox="[379,1209,1161,1574]" captionTargetId="figure-42@5.[379,1209,1161,1574]" captionTargetPageId="5" captionText="FIGURE 6. Type locality of Moenkhausia guaruba at upper rio Braço Norte at Serra do Cachimbo, tributary of rio Teles Pires, rio Tapajós basin, Pará State, Brazil: (a) waterfall upstream, substrate composed mainly by rocks; (b) sandy beach downstream to the waterfall." figureDoi="http://doi.org/10.5281/zenodo.8255390" httpUri="https://zenodo.org/record/8255390/files/figure.png" pageId="6" pageNumber="592">Fig. 6a</figureCitation>
), where the river is 50 wide and up to 
<quantity id="C1419B3FEA572671FE26D10B5494FA9A" box="[447,490,1338,1365]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="592" unit="m" value="4.0">
<elevation id="8D94D1E9EA572671FE26D10B5494FA9A" box="[447,490,1338,1365]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="592" unit="m" value="4.0">4 m</elevation>
</quantity>
deep (
<figureCitation id="9E822A5FEA572671FDAFD10B57F4FA9B" box="[566,650,1338,1364]" captionStart="FIGURE 6" captionStartId="5.[151,250,1598,1622]" captionTargetBox="[379,1209,1161,1574]" captionTargetId="figure-42@5.[379,1209,1161,1574]" captionTargetPageId="5" captionText="FIGURE 6. Type locality of Moenkhausia guaruba at upper rio Braço Norte at Serra do Cachimbo, tributary of rio Teles Pires, rio Tapajós basin, Pará State, Brazil: (a) waterfall upstream, substrate composed mainly by rocks; (b) sandy beach downstream to the waterfall." figureDoi="http://doi.org/10.5281/zenodo.8255390" httpUri="https://zenodo.org/record/8255390/files/figure.png" pageId="6" pageNumber="592">Fig. 6b</figureCitation>
). The substrate was composed mainly by sand and rocks and the river is bordered by Cerrado (Brazilian savannah) vegetation
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4EA36551EA572671FF5ED1B3546AF9C7" pageId="6" pageNumber="592" type="etymology">
<paragraph id="060636DAEA572671FF5ED1B3546AF9C7" blockId="6.[151,1437,150,1544]" pageId="6" pageNumber="592">
<emphasis id="34CDEAC8EA572671FF5ED1B35430FA53" bold="true" box="[199,334,1410,1436]" pageId="6" pageNumber="592">Etymology.</emphasis>
The specific name 
<taxonomicName id="C1B94D59EA572671FDA9D1B357F1FA53" box="[560,655,1410,1436]" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="592" phylum="Chordata" rank="species" species="guaruba">
<emphasis id="34CDEAC8EA572671FDA9D1B357F1FA53" box="[560,655,1410,1436]" italics="true" pageId="6" pageNumber="592">guaruba</emphasis>
</taxonomicName>
refers to the Brazilian popular name for 
<emphasis id="34CDEAC8EA572671FBFFD1B3503EFA53" box="[1126,1344,1410,1436]" italics="true" pageId="6" pageNumber="592">Guaruba guarouba</emphasis>
Gmelin 1788, also known as the Golden Parakeet, a medium-sized golden-yellow Neotropical parrot native to the Brazilian Amazon domain. The name alludes to the intense yellow present on all fins of the new species. A noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>