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<mods:title id="80B14F8D6DF8441E79F3B32BFEB93034">New olenelline trilobites from the Northwest Territories, Canada, and the phylogenetic placement of Judomia absita Fritz, 1973</mods:title>
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<mods:namePart id="15B8C7CECFB1C05D313E0A1E31912987">Gapp, Wesley</mods:namePart>
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<mods:namePart id="FF35E403E342381997C65BDEC4F73D99">Lieberman, Bruce S.</mods:namePart>
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<mods:namePart id="AC444189F2AEABBE43F88062DCA75139">Pope, Michael C.</mods:namePart>
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<mods:namePart id="E89022AC6F79CBF867D98154D1820D2A">Dilliard, Kelly A.</mods:namePart>
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<mods:date id="BA5BB4C06FC45A65C97BE3B00CD097F3">2011</mods:date>
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<treatment id="8E041C6CFFB4444FFF0689D895D6F942" ID-DOI="http://doi.org/10.5281/zenodo.6189867" ID-GBIF-Taxon="119411551" ID-Zenodo-Dep="6189867" LSID="urn:lsid:plazi:treatment:8E041C6CFFB4444FFF0689D895D6F942" httpUri="http://treatment.plazi.org/id/8E041C6CFFB4444FFF0689D895D6F942" lastPageNumber="23" pageId="8" pageNumber="23">
<subSubSection id="4EB7FEF1FFB4444FFF0689D8940BFD3D" pageId="8" pageNumber="23" type="nomenclature">
<paragraph id="0612AD7AFFB4444FFF0689D8972EFD5E" blockId="8.[151,583,655,714]" box="[151,583,655,681]" pageId="8" pageNumber="23">
<heading id="5D5A1A16FFB4444FFF0689D8972EFD5E" bold="true" box="[151,583,655,681]" fontSize="11" level="1" pageId="8" pageNumber="23" reason="1">
<emphasis id="34D97168FFB4444FFF0689D8972EFD5E" bold="true" box="[151,583,655,681]" pageId="8" pageNumber="23">
<emphasis id="34D97168FFB4444FFF0689D895CAFD5E" bold="true" box="[151,163,655,681]" italics="true" pageId="8" pageNumber="23">?</emphasis>
<taxonomicName id="C1ADD6F9FFB4444FFF3289D8972EFD5E" authority="McMenamin, 1987" authorityName="McMenamin" authorityYear="1987" box="[163,583,655,681]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="34D97168FFB4444FFF3289D8943FFD5E" bold="true" box="[163,342,655,681]" italics="true" pageId="8" pageNumber="23">Nevadia ovalis</emphasis>
<bibRefCitation id="623CD08BFFB4444FFECC89D8972EFD5E" author="McMenamin" box="[349,583,655,681]" pageId="8" pageNumber="23" refString="McMenamin, M. A. S. (1987) Lower Cambrian trilobites, zonation, and correlation of the Puerto Blanco Formation, Sonora, Mexico. Journal of Paleontology, 61, 738 - 749." type="journal article" year="1987">McMenamin, 1987</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="0612AD7AFFB4444FFF0689E5940BFD3D" blockId="8.[151,583,655,714]" box="[151,354,690,714]" pageId="8" pageNumber="23">(Fig. 6.3, 6.5, 6.6)</paragraph>
</subSubSection>
<subSubSection id="4EB7FEF1FFB4444FFF0689AD96AEFCE7" box="[151,967,762,784]" pageId="8" pageNumber="23" type="reference_group">
<paragraph id="0612AD7AFFB4444FFF0689AD96AEFCE7" blockId="8.[151,967,762,784]" box="[151,967,762,784]" pageId="8" pageNumber="23">
<emphasis id="34D97168FFB4444FFF0689AD9452FCE7" box="[151,315,762,784]" italics="true" pageId="8" pageNumber="23">
?
<taxonomicName id="C1ADD6F9FFB4444FFF3189AD9452FCE7" authority="McMenamin, 1987" authorityName="McMenamin" authorityYear="1987" box="[160,315,762,784]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="24" phylum="Arthropoda" rank="species" species="ovalis">Nevadia ovalis</taxonomicName>
</emphasis>
<bibRefCitation id="623CD08BFFB4444FFED089AC9767FCE7" author="MCMENAMIN" box="[321,526,763,784]" pageId="8" pageNumber="23" refString="McMenamin, M. A. S. (1987) Lower Cambrian trilobites, zonation, and correlation of the Puerto Blanco Formation, Sonora, Mexico. Journal of Paleontology, 61, 738 - 749." type="journal article" year="1987">MCMENAMIN, 1987</bibRefCitation>
, p. 744, figs. 5.2, 5.5, 5.6, 6.1, 6.3, 6.4, 6.6.
</paragraph>
</subSubSection>
<subSubSection id="4EB7FEF1FFB4444FFF06886A94F4FC6A" pageId="8" pageNumber="23" type="materials_examined">
<paragraph id="0612AD7AFFB4444FFF06886A94F4FC6A" blockId="8.[151,1436,829,1717]" pageId="8" pageNumber="23">
<emphasis id="34D97168FFB4444FFF06886A94EDFCA1" bold="true" box="[151,388,829,854]" pageId="8" pageNumber="23">Material examined.</emphasis>
PWNHC-2009.20.22 and KUMIP-320712-320713 from
<taxonomicName id="C1ADD6F9FFB4444FFB8A886A91E4FCA2" box="[1051,1165,829,853]" class="Trilobita" family="Nevadiidae" genus="Nevadella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="34D97168FFB4444FFB8A886A91E4FCA2" box="[1051,1165,829,853]" italics="true" pageId="8" pageNumber="23">Nevadella</emphasis>
</taxonomicName>
zone, Early Cambrian, Sekwi Formation, Mackenzie Mountains, Northwest Territories,
<collectingCountry id="7EBAEDEAFFB4444FFCEA883696A6FC8E" box="[891,975,865,889]" name="Canada" pageId="8" pageNumber="23">Canada</collectingCountry>
(Section 1, 199.5
<quantity id="C155009FFFB4444FFB3B8836919DFC8E" box="[1194,1268,865,889]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="23" unit="m" value="300.0">300 m</quantity>
above base of formation and in float).
</paragraph>
</subSubSection>
<subSubSection id="4EB7FEF1FFB4444FFF5688FE95D6F942" pageId="8" pageNumber="23" type="discussion">
<paragraph id="0612AD7AFFB4444FFF5688FE9604FB16" blockId="8.[151,1436,829,1717]" pageId="8" pageNumber="23">
<emphasis id="34D97168FFB4444FFF5688FE9425FC35" bold="true" box="[199,332,937,962]" pageId="8" pageNumber="23">Discussion.</emphasis>
These specimens may be referable to
<taxonomicName id="C1ADD6F9FFB4444FFD6888FE9111FC36" authority="McMenamin 1987" authorityName="McMenamin" authorityYear="1987" box="[761,1144,937,961]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="34D97168FFB4444FFD6888FE96C8FC36" box="[761,929,937,961]" italics="true" pageId="8" pageNumber="23">Nevadia ovalis</emphasis>
<bibRefCitation id="623CD08BFFB4444FFC3988FE9111FC36" author="McMenamin" box="[936,1144,937,961]" pageId="8" pageNumber="23" refString="McMenamin, M. A. S. (1987) Lower Cambrian trilobites, zonation, and correlation of the Puerto Blanco Formation, Sonora, Mexico. Journal of Paleontology, 61, 738 - 749." type="journal article" year="1987">McMenamin 1987</bibRefCitation>
</taxonomicName>
originally described from the middle shaley interval of unit 3 of the Puerto Blanco Formation of Sonora,
<collectingCountry id="7EBAEDEAFFB4444FFB89889A9118FC12" box="[1048,1137,973,997]" name="Mexico" pageId="8" pageNumber="23">Mexico</collectingCountry>
. For instance, the material from the Sekwi Formation appears to match that species in possessing the anterior and posterior cephalic borders much wider than in other species of
<taxonomicName id="C1ADD6F9FFB4444FFDA68F4297FDFBDA" box="[567,660,1045,1069]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="34D97168FFB4444FFDA68F4297FDFBDA" box="[567,660,1045,1069]" italics="true" pageId="8" pageNumber="23">Nevadia</emphasis>
</taxonomicName>
, and the extraocular area much narrower, as well as a cephalon that has an ovate shape along its anterolateral margins. However, because neither McMenamins (1987) original material nor the new material from the Sekwi Formation is particularly well preserved, our assignment is considered tentative, and awaits discovery and identification of new material from each of these localities. The material from the Sekwi appears distinct from the aforementioned early ontogenetic stages of cephala
<bibRefCitation id="623CD08BFFB4444FFBE98FF29196FB4A" author="Fritz" box="[1144,1279,1189,1213]" pageId="8" pageNumber="23" refString="Fritz, W. H. (1973) Medial Lower Cambrian trilobites from the Mackenzie Mountains, northwestern Canada. Geological Survey of Canada, Paper, 73 - 24, 43 pp." type="book" year="1973">Fritz (1973)</bibRefCitation>
referred to as
<taxonomicName id="C1ADD6F9FFB4444FFF068F9E959DFB16" box="[151,244,1225,1249]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="34D97168FFB4444FFF068F9E959DFB16" box="[151,244,1225,1249]" italics="true" pageId="8" pageNumber="23">Nevadia</emphasis>
</taxonomicName>
? sp., differing principally in the outline of the cephalon.
</paragraph>
<paragraph id="0612AD7AFFB4444FFF568FBA91BBF9F6" blockId="8.[151,1436,829,1717]" pageId="8" pageNumber="23">
McMenamin assigned this species to the genus
<taxonomicName id="C1ADD6F9FFB4444FFD768FBA962DFAF2" box="[743,836,1261,1285]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="34D97168FFB4444FFD768FBA962DFAF2" box="[743,836,1261,1285]" italics="true" pageId="8" pageNumber="23">Nevadia</emphasis>
</taxonomicName>
based on a character of the “anterior end of the glabella well forward of the palpebral lobes.” This species remains assigned to the “nevadiids” based on the presence of characters discussed by Lieberman (1999, 2001) such as the width (tr.) of the interocular area being approximately equal to the width of the ocular lobe at its midlength and S3 conjoined medially. This is in contrast with the monophyletic Judomiodea, which is grouped by characters such as the width (tr.) of the interocular area being equal to two to three times the width of the ocular lobe at its midlength and S3 not being conjoined medially. In addition, the thoracic pleural spines are relatively short, and although this may not be typical for all members of
<taxonomicName id="C1ADD6F9FFB4444FFF068EBE959DF9F6" box="[151,244,1513,1537]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="34D97168FFB4444FFF068EBE959DF9F6" box="[151,244,1513,1537]" italics="true" pageId="8" pageNumber="23">Nevadia</emphasis>
</taxonomicName>
and
<taxonomicName id="C1ADD6F9FFB4444FFEBA8EBE94F4F9F6" box="[299,413,1513,1537]" class="Trilobita" family="Nevadiidae" genus="Nevadella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="34D97168FFB4444FFEBA8EBE94F4F9F6" box="[299,413,1513,1537]" italics="true" pageId="8" pageNumber="23">Nevadella</emphasis>
</taxonomicName>
it is known from other &quot;nevadiids&quot;, for instance,
<taxonomicName id="C1ADD6F9FFB4444FFC578EBE91A7F9F6" authority="Blaker, 1988" authorityName="Blaker" authorityYear="1988" box="[966,1230,1513,1537]" class="Trilobita" family="Nevadiidae" genus="Buenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="34D97168FFB4444FFC578EBE915DF9F6" box="[966,1076,1513,1537]" italics="true" pageId="8" pageNumber="23">Buenellus</emphasis>
<bibRefCitation id="623CD08BFFB4444FFBAD8EBE91A7F9F6" author="Blaker" box="[1084,1230,1513,1537]" pageId="8" pageNumber="23" refString="Blaker, M. R. (1988) A new genus of nevadiid trilobite from the Buen Formation (Early Cambrian) of Peary Land, central North Greenland. Gronlands geologiska Undersogelse Rapports, 137, 33 - 41." type="journal article" year="1988">Blaker, 1988</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="0612AD7AFFB4444FFF568D5995D6F942" blockId="8.[151,1436,829,1717]" pageId="8" pageNumber="23">
Most olenellinid species tend to have very limited geographic ranges and are often endemic to narrowly defined areas. One of the rare exceptions is
<taxonomicName id="C1ADD6F9FFB4444FFDEE8D66914AF9BE" authority="Walcott 1910" authorityName="Walcott" authorityYear="1910" box="[639,1059,1585,1609]" class="Trilobita" family="Wanneriidae" genus="Elliptocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="species" species="logani">
<emphasis id="34D97168FFB4444FFDEE8D66961AF9BE" box="[639,883,1585,1609]" italics="true" pageId="8" pageNumber="23">Elliptocephala logani</emphasis>
(
<bibRefCitation id="623CD08BFFB4444FFC118D669172F9BE" author="Walcott" box="[896,1051,1585,1609]" pageId="8" pageNumber="23" refString="Walcott, C. D. (1910) Olenellus and other genera of the Mesonacidae. Smithsonian Miscellaneous Collections, 53 (6), 231 - 422." type="journal article" year="1910">Walcott 1910</bibRefCitation>
)
</taxonomicName>
, which straddles both the western and eastern parts of North
<collectingCountry id="7EBAEDEAFFB4444FFE508D029748F99A" box="[449,545,1621,1645]" name="United States of America" pageId="8" pageNumber="23">America</collectingCountry>
(Lieberman 1999). Therefore,
<taxonomicName id="C1ADD6F9FFB4444FFCEC8D02914CF99A" box="[893,1061,1621,1645]" class="Trilobita" family="Nevadiidae" genus="Nevadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="23" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="34D97168FFB4444FFCEC8D02914CF99A" box="[893,1061,1621,1645]" italics="true" pageId="8" pageNumber="23">Nevadia ovalis</emphasis>
</taxonomicName>
, from southwestern North
<collectingCountry id="7EBAEDEAFFB4444FFAC58D0195D3F966" name="United States of America" pageId="8" pageNumber="23">America</collectingCountry>
, may comprise another such exception, if it is ultimately confirmed to be present in northwestern North
<collectingCountry id="7EBAEDEAFFB4444FFAC28D2D95D2F942" name="United States of America" pageId="8" pageNumber="23">America</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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