207 lines
20 KiB
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207 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.97.59143" ID-GBIF-Dataset="83d5b180-4e0b-44d1-ad83-67ad55ee0355" ID-GBIF-Taxon="182236447" ID-Pensoft-Pub="1860-0743-1-1" ID-Pensoft-UUID="BD006A3AD2C6516AB596CDA05E0F375D" ID-ZooBank="1D5553BA24894F2C9527B7E46B9B432B" LSID="urn:lsid:zoobank.org:act:C6236997-4DFE-43BD-971B-29C5E071A89E" ModsDocID="1860-0743-97-1-1" checkinTime="1621423297205" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Pall-Gergely, Barna, Sutcharit, Chirasak & Tongkerd, Piyoros" docDate="2021" docId="77D9199D9ECA5F1D835AD07CC058731B" docLanguage="en" docName="ZoosystEvol 97(1): 1-20" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-01-05" docSource="http://dx.doi.org/10.3897/zse.97.59143" docTitle="Dicharax panhai Jirapatrasilp & Pall-Gergely 2021, sp. nov." docType="treatment" docUuid="C6236997-4DFE-43BD-971B-29C5E071A89E" docUuidSource="ZooBank" docVersion="3" id="BD006A3AD2C6516AB596CDA05E0F375D" lastPageNumber="1" masterDocId="BD006A3AD2C6516AB596CDA05E0F375D" masterDocTitle="The operculate micro land snail genus Dicharax Kobelt & Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species" masterLastPageNumber="20" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1632012559486" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The operculate micro land snail genus Dicharax Kobelt & Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9221-9293</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">piyorose@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-01-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>20</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.97.59143</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.59143</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-1</mods:identifier>
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<mods:identifier type="ZooBank">1D5553BA24894F2C9527B7E46B9B432B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BD006A3AD2C6516AB596CDA05E0F375D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182236447" LSID="urn:lsid:zoobank.org:act:C6236997-4DFE-43BD-971B-29C5E071A89E" httpUri="http://treatment.plazi.org/id/77D9199D9ECA5F1D835AD07CC058731B" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/C6236997-4DFE-43BD-971B-29C5E071A89E" authority="Jirapatrasilp & Páll-Gergely" authorityName="Jirapatrasilp & Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax panhai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="panhai" status="sp. nov.">
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Dicharax panhai Jirapatrasilp &
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">Figs 6C</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C. Dicharax panhai sp. nov. A. Holotype CUMZ 7429 / 1; B. Paratype CUMZ 7429 / 2; C. paratype SMF 331452 (paratype of " Alycaeus pratatensis Cy 003 " from Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand). D. Dicharax pongrati sp. nov., holotype CUMZ 7430 / 1 Close-up images of the aperture not to scale. Photo: B. Pall-Gergely, courtesy Ronald Janssen (C.)." figureDoi="10.3897/zse.97.59143.figure12" httpUri="https://binary.pensoft.net/fig/494376" pageId="0" pageNumber="1">, 12A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="Jirapatrasilp & Páll-Gergely & Sutcharit & Tongkerd, 2021" authorityName="Jirapatrasilp & Páll-Gergely & Sutcharit & Tongkerd" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax pratatensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pratatensis" status="sp. nov.">Dicharax pratatensis</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.981.53583" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 220" refId="B47" refString="Pall-Gergely, B, Sajan, S, Tripathy, B, Meng, K, Takahiro, A, Ablett, J, 2020b. Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1 - 220, DOI: https://doi.org/10.3897/zookeys.981.53583" title="Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue." url="https://doi.org/10.3897/zookeys.981.53583" volume="981" year="2020 b">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2020b
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</bibRefCitation>
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: 71, 72.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 7429/1 (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C. Dicharax panhai sp. nov. A. Holotype CUMZ 7429 / 1; B. Paratype CUMZ 7429 / 2; C. paratype SMF 331452 (paratype of " Alycaeus pratatensis Cy 003 " from Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand). D. Dicharax pongrati sp. nov., holotype CUMZ 7430 / 1 Close-up images of the aperture not to scale. Photo: B. Pall-Gergely, courtesy Ronald Janssen (C.)." figureDoi="10.3897/zse.97.59143.figure12" httpUri="https://binary.pensoft.net/fig/494376" pageId="0" pageNumber="1">12A</figureCitation>
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),
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paratypes</emphasis>
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CUMZ 7429/2 (16 shells; Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">6C</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C. Dicharax panhai sp. nov. A. Holotype CUMZ 7429 / 1; B. Paratype CUMZ 7429 / 2; C. paratype SMF 331452 (paratype of " Alycaeus pratatensis Cy 003 " from Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand). D. Dicharax pongrati sp. nov., holotype CUMZ 7430 / 1 Close-up images of the aperture not to scale. Photo: B. Pall-Gergely, courtesy Ronald Janssen (C.)." figureDoi="10.3897/zse.97.59143.figure12" httpUri="https://binary.pensoft.net/fig/494376" pageId="0" pageNumber="1">12B</figureCitation>
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), HNHM (2 shells), NHMUK 20200327 (2 shells), paratypes of "
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus pratatensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pratatensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus pratatensis</emphasis>
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</taxonomicName>
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Cy 003" FMNH (2 shells), SMF 331452 (2 shells; Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C. Dicharax panhai sp. nov. A. Holotype CUMZ 7429 / 1; B. Paratype CUMZ 7429 / 2; C. paratype SMF 331452 (paratype of " Alycaeus pratatensis Cy 003 " from Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand). D. Dicharax pongrati sp. nov., holotype CUMZ 7430 / 1 Close-up images of the aperture not to scale. Photo: B. Pall-Gergely, courtesy Ronald Janssen (C.)." figureDoi="10.3897/zse.97.59143.figure12" httpUri="https://binary.pensoft.net/fig/494376" pageId="0" pageNumber="1">12C</figureCitation>
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) and UMMZ (2 shells) from Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand,
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<geoCoordinate degrees="14" direction="north" minutes="23" orientation="latitude" precision="1" seconds="50.8" value="14.397445">14°23'50.8"N</geoCoordinate>
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,
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<geoCoordinate degrees="99" direction="east" minutes="04" orientation="longitude" precision="1" seconds="56.3" value="99.082306">99°04'56.3"E</geoCoordinate>
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, 230 a.s.l., 26 Oct. 1996, coll. S. Panha.
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</paragraph>
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<caption doi="10.3897/zse.97.59143.figure12" httpUri="https://binary.pensoft.net/fig/494376" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="1">A-C.</emphasis>
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<taxonomicName authorityName="Jirapatrasilp & Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax panhai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="panhai">
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<emphasis italics="true" pageId="0" pageNumber="1">Dicharax panhai</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">A.</emphasis>
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Holotype CUMZ 7429/1;
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<emphasis bold="true" pageId="0" pageNumber="1">B.</emphasis>
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Paratype CUMZ 7429/2;
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<emphasis bold="true" pageId="0" pageNumber="1">C.</emphasis>
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paratype SMF 331452 (paratype of "
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus pratatensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pratatensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus pratatensis</emphasis>
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</taxonomicName>
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Cy 003" from Tham Pratat, Erawan National Park, Kanchanaburi Province, Thailand).
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<emphasis bold="true" pageId="0" pageNumber="1">D.</emphasis>
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<taxonomicName authorityName="Jirapatrasilp & Tongkerd" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax pongrati" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pongrati">
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<emphasis italics="true" pageId="0" pageNumber="1">Dicharax pongrati</emphasis>
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</taxonomicName>
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sp. nov., holotype CUMZ 7430/1 Close-up images of the aperture not to scale. Photo: B.
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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, courtesy Ronald Janssen (
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<emphasis bold="true" pageId="0" pageNumber="1">C.</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell small (SH up to 2.4 mm, SW up to 3.1 mm), depressed-conical with triangular dorsal side and widened, sigmoid last whorl. Spire ca. 1/3 of shell height. R1 with regular, thread-like ribs; R2 twice shorter than R3; R2 with ca. 18 ribs. R3 with blunt central swelling. Aperture round. Inner peristome thick, protruding. Outer peristome expanded, not reflected. Umbilicus elliptical.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell small (SH up to 2.4 mm, SW up to 3.1 mm), depressed-conical with triangular dorsal side and widened, sigmoid last whorl; solid, worn out white; the exact color of non-weathered (fresh) shells unknown. Shell outline oval in apical view, spire ca. 1/3 of shell height. Whorls ca. 4 and 1/8. Protoconch low, ca. two whorls, glossy and smooth. R1 ca.
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<normalizedToken originalValue="1¾">13/4</normalizedToken>
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whorls with fine, regular, thread-like ribs; with ca. 14 ribs in
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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whorl adjoining R2, ribs sharper near suture and flatter near umbilicus. Boundary between R1 and R2 distinct as R2 contains more close-set ribs than R1; R2 with ca. 18 ribs that are curved towards aperture; R2 ca. 1/8 whorl, twice shorter than R3. Boundary between R2 and R3 distinct due to shallow constriction; R3 initially coils horizontally forming a flat base, but aperture strongly descent being strongly oblique to shell axis; R3 mostly smooth with indistinct growth lines that become stronger near aperture; with blunt central swelling. Aperture round; slightly oblique to shell axis. Peristome double with regular outer peristome. Inner peristome thick, protruding. Outer peristome slightly expanded, not reflected, multi-layered (visible mostly in lateral view). Umbilicus elliptical, open, slightly more than one third of shell width. Operculum unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">In honor of Prof. Somsak Panha, a prominent Thai malacologist and our beloved professor who initiated the taxonomic and systematic study of land snails and other terrestrial invertebrates in Thailand and surrounding areas of mainland Southeast Asia.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">The new species is known only from the type locality in Kanchanaburi Province, Thailand.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Similar to
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<taxonomicName genus="D." lsidName="D. pratatensis" pageId="0" pageNumber="1" rank="species" species="pratatensis">
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<emphasis italics="true" pageId="0" pageNumber="1">D. pratatensis</emphasis>
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</taxonomicName>
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from Doi Chiang Dao, Chiang Mai Province, Thailand in having a depressed-conical shell shape and R3 with blunt swelling, but differs in having a conical apical part on a wide shell base, less distinct ribs of R1 and shorter R2 with less ribs. The short R2 and breathing tube is also exhibited in
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<taxonomicName genus="D." lsidName="D. abdoui" pageId="0" pageNumber="1" rank="species" species="abdoui">
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<emphasis italics="true" pageId="0" pageNumber="1">D. abdoui</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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in
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Do, DS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331: 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al., 2017
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</bibRefCitation>
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from Khammouane Province, Laos and
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<taxonomicName genus="D." lsidName="D. stuparum" pageId="0" pageNumber="1" rank="species" species="stuparum">
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<emphasis italics="true" pageId="0" pageNumber="1">D. stuparum</emphasis>
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</taxonomicName>
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from Chiang Rai Province, Northern Thailand, but
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<taxonomicName genus="D." lsidName="D. panhai" pageId="0" pageNumber="1" rank="species" species="panhai">
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<emphasis italics="true" pageId="0" pageNumber="1">D. panhai</emphasis>
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</taxonomicName>
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sp. nov. is larger, comprises more whorls, has a more elevated spire and a larger last whorl (
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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,
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |