treatments-xml/data/64/13/F3/6413F378FFE06A7DF28B7739FDAAF8A4.xml
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<document id="90BFAAC913F33FB330777085DA58C74F" ID-DOI="10.11646/zootaxa.2462.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629676798216" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Ponder, Winston F." docDate="2010" docId="6413F378FFE06A7DF28B7739FDAAF8A4" docLanguage="en" docName="zt02462p148.pdf" docOrigin="Zootaxa 2462 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Dendrotrochus (Santotrochus) eva subsp. eva eva (Pfeiffer 1854" docType="treatment" docVersion="4" lastPageNumber="120" masterDocId="982A8B00FF976A0AF21C766BFFEBFF8B" masterDocTitle="A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462" masterLastPageNumber="148" masterPageNumber="1" pageNumber="120" updateTime="1699094437759" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E5311EE6F87C513DE6A9C9D584E77214">A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462</mods:title>
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<mods:namePart id="EFBE5D508A3B9DCCE4CD940A383C3D0F">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="4D9D5CC26EB2F5FAEC84A77FE7E72CB3">Ponder, Winston F.</mods:namePart>
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<mods:title id="F266CC7727D8D5F14767868F6A8FC7E7">Zootaxa</mods:title>
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<mods:date id="B3BDA86A31FD74B82FFC0AA7556C9327">2010</mods:date>
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<mods:number id="6223C71A938DB40C8C71EAAF9FBEBF1C">2010-05-14</mods:number>
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<mods:number id="F0EAA43E7C4FC2F6D3B0F820C8CA6250">2462</mods:number>
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<treatment id="6413F378FFE06A7DF28B7739FDAAF8A4" ID-DOI="http://doi.org/10.5281/zenodo.5321149" ID-Zenodo-Dep="5321149" LSID="urn:lsid:plazi:treatment:6413F378FFE06A7DF28B7739FDAAF8A4" httpUri="http://treatment.plazi.org/id/6413F378FFE06A7DF28B7739FDAAF8A4" lastPageNumber="120" pageId="119" pageNumber="120">
<subSubSection id="A4A011E5FFE06A7DF28B7739FCC3FEE7" box="[151,808,338,364]" pageId="119" pageNumber="120" type="nomenclature">
<paragraph id="EC05426EFFE06A7DF28B7739FCC3FEE7" blockId="119.[151,808,338,399]" box="[151,808,338,364]" pageId="119" pageNumber="120">
<heading id="B74DF502FFE06A7DF28B7739FCC3FEE7" bold="true" box="[151,808,338,364]" fontSize="11" level="1" pageId="119" pageNumber="120" reason="1">
<taxonomicName id="2BBA39EDFFE06A7DF28B7739FCC3FEE7" ID-CoL="8RXSC" authority="eva (Pfeiffer, 1854)" authorityName="eva (Pfeiffer" authorityYear="1854" baseAuthorityName="Pfeiffer" baseAuthorityYear="1854" box="[151,808,338,364]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="subSpecies" species="eva" subGenus="Santotrochus" subSpecies="eva">
<emphasis id="DECE9E7CFFE06A7DF28B7739FCC3FEE7" bold="true" box="[151,808,338,364]" pageId="119" pageNumber="120">
<emphasis id="DECE9E7CFFE06A7DF28B7739FEA6FEE7" bold="true" box="[151,333,338,364]" italics="true" pageId="119" pageNumber="120">Dendrotrochus</emphasis>
(
<emphasis id="DECE9E7CFFE06A7DF3427739FE14FEE7" bold="true" box="[350,511,338,364]" italics="true" pageId="119" pageNumber="120">Santotrochus</emphasis>
)
<emphasis id="DECE9E7CFFE06A7DF00C7739FD82FEE7" bold="true" box="[528,617,338,364]" italics="true" pageId="119" pageNumber="120">eva eva</emphasis>
(Pfeiffer, 1854)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFE06A7DF28B771EFE11FE04" box="[151,506,373,399]" pageId="119" pageNumber="120" type="description">
<paragraph id="EC05426EFFE06A7DF28B771EFE11FE04" blockId="119.[151,808,338,399]" box="[151,506,373,399]" pageId="119" pageNumber="120">
<figureCitation id="74815EEBFFE06A7DF28B771EFEC0FE04" box="[151,299,373,399]" captionStart="FIGURE 33" captionStartId="87.[151,255,1800,1824]" captionTargetBox="[302,1298,193,1767]" captionTargetId="figure-146@87.[271,1323,175,1789]" captionTargetPageId="87" captionText="FIGURE 33. Penis interiors. A. Erepta odontina, AM C205302. B. Kalidos balstoni, AM C205303. C. Madagascan species MA1, AM C205304. D. Sheldonia poeppigii, NMSA L5563. E. Elisolimax flavescens, NMSA V9710. F. Parmarion martensi, BPBM 252738. G. Coneuplecta pampini, AM C438751. H. Hiona pilsbryi, BPBM 206801 &amp; 20701. I. Trochomorpha rubens, FMNH 175247. J. Videna planorbis, FMNH 160957. K. Dendrotrochus eva eva, FMNH 175196. L. Orpiella concavum, BMNH no registration number. Scale bars 1 mm (A, CD, FH, J, M), 2 mm (B, E, I, K). Abbreviations: bc, bursa copulatrix; c, caecum; ep, epiphallus; fov, free oviduct; ovi, ovipositer; p, penis; pp, penial pilaster; prm, penis retractor muscle; ppu, penial pustules; pv, penial verge." figureDoi="http://doi.org/10.5281/zenodo.5312800" httpUri="https://zenodo.org/record/5312800/files/figure.png" pageId="119" pageNumber="120">Figures 33K</figureCitation>
,
<figureCitation id="74815EEBFFE06A7DF32B771EFE8EFE04" box="[311,357,373,399]" captionStart="FIGURE 35" captionStartId="91.[151,255,1089,1113]" captionTargetBox="[276,1312,513,1059]" captionTargetId="figure-352@91.[264,1324,499,1076]" captionTargetPageId="91" captionText="FIGURE 35. External morphology of caudal regions. A. Kalidos balstoni, AM C205303. B. Macrochlamys petrosa, AM C205303. C. Everettia sp., BMNH 20020188. D. Greenwoodoconcha nux, AM C433375. E. Videna planorbis, FMNH 160957. F. Dendrotrochus eva eva, FMNH 160957. Scale bars 1 mm (D), 2 mm (EF), 5 mm (AC)." figureDoi="http://doi.org/10.5281/zenodo.5312806" httpUri="https://zenodo.org/record/5312806/files/figure.png" pageId="119" pageNumber="120">35F</figureCitation>
,
<figureCitation id="74815EEBFFE06A7DF36D771EFE48FE04" box="[369,419,373,399]" captionStart="FIGURE 41" captionStartId="114.[151,255,1795,1819]" captionTargetBox="[243,1348,266,1770]" captionTargetId="figure-131@114.[240,1348,262,1771]" captionTargetPageId="114" captionText="FIGURE 41. Reproductive systems. A. Lamprocystis guttula, FMNH 174940. B. Trochomorpha rubens, FMNH 175247. C. Videna planorbis, FMNH 160957. D. Orpiella concavum, BMNH no registration number. E. Dendrotrochus eva eva, FMNH 175196. Scale bars 1 mm (A), 2 mm (BD, F), 5 mm (E). Abbreviations: alb, albumen gland; bc, bursa copulatrix; ca, carrefour; e, eggs / embryos; ep, epiphallus; fov, free oviduct; her, hermaphrodite duct; ov, ovotestis; p, penis; pr, prostate; prm, penis retractor muscle; puo, post-uterine oviduct; t, talon; ut, uterus; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5312822" httpUri="https://zenodo.org/record/5312822/files/figure.png" pageId="119" pageNumber="120">41E</figureCitation>
,
<figureCitation id="74815EEBFFE06A7DF3B3771EFE11FE04" box="[431,506,373,399]" captionStart="FIGURE 42" captionStartId="118.[151,255,1168,1192]" captionTargetBox="[257,1330,194,1144]" captionTargetId="figure-236@118.[257,1330,194,1144]" captionTargetPageId="118" captionText="FIGURE 42. Radulae. AC. Trochomorpha rubens, FMNH 175247. DF. Videna planorbis, FMNH 160957. GI. Dendrotrochus eva eva, FMNH 15719. JL. Orpiella concavum, BMNH no registration number. Scale bars 20 mm (AL)." figureDoi="http://doi.org/10.5281/zenodo.5312826" httpUri="https://zenodo.org/record/5312826/files/figure.png" pageId="119" pageNumber="120">42GI</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFE06A7DF28B77AEFBEBFD8C" pageId="119" pageNumber="120" type="materials_examined">
<paragraph id="EC05426EFFE06A7DF28B77AEFBEBFD8C" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
<emphasis id="DECE9E7CFFE06A7DF28B77AEFE60FE54" bold="true" box="[151,395,453,479]" pageId="119" pageNumber="120">Material examined.</emphasis>
<materialsCitation id="5CD24833FFE06A7DF38F77AEFC16FD8C" collectingDate="1972-11-22" collectionCode="FMNH" collectorName="L. Price" country="Vanuatu" elevation="240" location="Clems Hill" municipality="Nfate" pageId="119" pageNumber="120" specimenCode="FMNH 175196" specimenCount="1">
<emphasis id="DECE9E7CFFE06A7DF38F77AEFE15FE54" bold="true" box="[403,510,453,479]" pageId="119" pageNumber="120">
<collectingCountry id="94AD02FEFFE06A7DF38F77AEFE15FE54" box="[403,510,453,479]" name="Vanuatu" pageId="119" pageNumber="120">Vanuatu</collectingCountry>
</emphasis>
:
<specimenCode id="BC1CEA15FFE06A7DF01277AEFD28FE54" box="[526,707,453,479]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="119" pageNumber="120" type="Museum">FMNH 175196</specimenCode>
(
<specimenCount id="FABC89E7FFE06A7DF0CF77AEFC68FE54" box="[723,899,453,479]" pageId="119" pageNumber="120" type="generic">two specimens</specimenCount>
dissected, one radula examined),
<location id="E96514B5FFE06A7DF70877AEFA7CFE54" LSID="urn:lsid:plazi:treatment:6413F378FFE06A7DF28B7739FDAAF8A4:E96514B5FFE06A7DF70877AEFA7CFE54" box="[1300,1431,453,479]" country="Vanuatu" municipality="Nfate" name="Clems Hill" pageId="119" pageNumber="120">Clems Hill</location>
,
<locationDeviation id="FDB75B4CFFE06A7DF28B7786FE9FFD8C" box="[151,372,493,519]" location="Vila" pageId="119" pageNumber="120">
<quantity id="2B42EF8BFFE06A7DF28B7786FF0BFD8C" box="[151,224,493,519]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="119" pageNumber="120" unit="km" value="17.0">17 km</quantity>
NW of Vila
</locationDeviation>
,
<collectingMunicipality id="0C61D814FFE06A7DF3637786FE28FD8C" box="[383,451,493,519]" pageId="119" pageNumber="120">Nfate</collectingMunicipality>
, dense forest, alt.
<quantity id="2B42EF8BFFE06A7DF08A7786FD09FD8C" box="[662,738,493,519]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="119" pageNumber="120" unit="m" value="240.0">
<elevation id="6797A55DFFE06A7DF08A7786FD09FD8C" box="[662,738,493,519]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="119" pageNumber="120" unit="m" value="240.0">240 m</elevation>
</quantity>
,
<date id="980464AEFFE06A7DF0F37786FC7BFD8C" box="[751,912,493,519]" pageId="119" pageNumber="120" value="1972-11-22">
<collectingDate id="88409D46FFE06A7DF0F37786FC7BFD8C" box="[751,912,493,519]" pageId="119" pageNumber="120" value="1972-11-22">22 Nov. 1972</collectingDate>
</date>
,
<collectorName id="414F27B8FFE06A7DF1807786FC16FD8C" box="[924,1021,493,519]" pageId="119" pageNumber="120">L. Price</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFE06A7DF2DA747EFDF1FAFC" pageId="119" pageNumber="120" type="description">
<paragraph id="EC05426EFFE06A7DF2DA747EFF09FD7C" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
<emphasis id="DECE9E7CFFE06A7DF2DA747EFEB5FDA4" bold="true" box="[198,350,533,559]" pageId="119" pageNumber="120">Description.</emphasis>
<emphasis id="DECE9E7CFFE06A7DF37A747EFDB4FDA5" box="[358,607,533,558]" italics="true" pageId="119" pageNumber="120">External morphology</emphasis>
: Shell of about 5.5 whorls, white, glossy, spire and apex raised. Protoconch smooth; teleoconch sculptured with strong radial growth lines. Whorl profile rounded above and below keeled periphery. Internal walls of early whorls complete; umbilicus open, very narrow U-shape. Animal (
<figureCitation id="74815EEBFFE06A7DF775740EFF1FFD2C" captionStart="FIGURE 35" captionStartId="91.[151,255,1089,1113]" captionTargetBox="[276,1312,513,1059]" captionTargetId="figure-352@91.[264,1324,499,1076]" captionTargetPageId="91" captionText="FIGURE 35. External morphology of caudal regions. A. Kalidos balstoni, AM C205303. B. Macrochlamys petrosa, AM C205303. C. Everettia sp., BMNH 20020188. D. Greenwoodoconcha nux, AM C433375. E. Videna planorbis, FMNH 160957. F. Dendrotrochus eva eva, FMNH 160957. Scale bars 1 mm (D), 2 mm (EF), 5 mm (AC)." figureDoi="http://doi.org/10.5281/zenodo.5312806" httpUri="https://zenodo.org/record/5312806/files/figure.png" pageId="119" pageNumber="120">Figure 35F</figureCitation>
) light yellow-brown. Mantle laps and lobes absent. Sole of foot tripartite; caudal apparatus formed from curled up sole (
<bibRefCitation id="882B3F9FFFE06A7DF39274DEFDA0FD44" author="Muratov, I. V." box="[398,587,693,719]" pageId="119" pageNumber="120" pagination="5 - 26" refId="ref75844" refString="Muratov, I. V. (1999) Analysis of the phylogenetic relationships and their systematic implications in the Limacoinei (= Zonitinia) infraorder (Gastropoda, Pulmonata, Geophila). Ruthenica, 9, 5 - 26." type="journal article" year="1999">Muratovs 1999</bibRefCitation>
helicarionid
<typeStatus id="3301FCCCFFE06A7DF0F474DEFCF4FD44" box="[744,799,693,719]" pageId="119" pageNumber="120">type</typeStatus>
); caudal horn medium size; caudal foss horizontal slit in tail.
</paragraph>
<paragraph id="EC05426EFFE06A7DF2DF756EFC49FC94" blockId="119.[151,1437,453,1839]" box="[195,930,773,799]" pageId="119" pageNumber="120">
<emphasis id="DECE9E7CFFE06A7DF2DF756EFE88FC95" box="[195,355,773,798]" italics="true" pageId="119" pageNumber="120">Mantle cavity</emphasis>
: As for family. Pigmentation of tiny white spots.
</paragraph>
<paragraph id="EC05426EFFE06A7DF2DF7546FD29FCCC" blockId="119.[151,1437,453,1839]" box="[195,706,813,839]" pageId="119" pageNumber="120">
<emphasis id="DECE9E7CFFE06A7DF2DF7546FE6CFCCD" box="[195,391,813,838]" italics="true" pageId="119" pageNumber="120">Digestive system</emphasis>
: Oesophageal crop absent.
</paragraph>
<paragraph id="EC05426EFFE06A7DF2DF753EFBF8FB0C" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
<emphasis id="DECE9E7CFFE06A7DF2DF753EFE99FCE5" box="[195,370,853,878]" italics="true" pageId="119" pageNumber="120">Genital system</emphasis>
(
<figureCitation id="74815EEBFFE06A7DF398753EFDF7FCE4" box="[388,540,853,879]" captionStart="FIGURE 33" captionStartId="87.[151,255,1800,1824]" captionTargetBox="[302,1298,193,1767]" captionTargetId="figure-146@87.[271,1323,175,1789]" captionTargetPageId="87" captionText="FIGURE 33. Penis interiors. A. Erepta odontina, AM C205302. B. Kalidos balstoni, AM C205303. C. Madagascan species MA1, AM C205304. D. Sheldonia poeppigii, NMSA L5563. E. Elisolimax flavescens, NMSA V9710. F. Parmarion martensi, BPBM 252738. G. Coneuplecta pampini, AM C438751. H. Hiona pilsbryi, BPBM 206801 &amp; 20701. I. Trochomorpha rubens, FMNH 175247. J. Videna planorbis, FMNH 160957. K. Dendrotrochus eva eva, FMNH 175196. L. Orpiella concavum, BMNH no registration number. Scale bars 1 mm (A, CD, FH, J, M), 2 mm (B, E, I, K). Abbreviations: bc, bursa copulatrix; c, caecum; ep, epiphallus; fov, free oviduct; ovi, ovipositer; p, penis; pp, penial pilaster; prm, penis retractor muscle; ppu, penial pustules; pv, penial verge." figureDoi="http://doi.org/10.5281/zenodo.5312800" httpUri="https://zenodo.org/record/5312800/files/figure.png" pageId="119" pageNumber="120">Figures 33K</figureCitation>
,
<figureCitation id="74815EEBFFE06A7DF036753EFDB5FCE4" box="[554,606,853,879]" captionStart="FIGURE 41" captionStartId="114.[151,255,1795,1819]" captionTargetBox="[243,1348,266,1770]" captionTargetId="figure-131@114.[240,1348,262,1771]" captionTargetPageId="114" captionText="FIGURE 41. Reproductive systems. A. Lamprocystis guttula, FMNH 174940. B. Trochomorpha rubens, FMNH 175247. C. Videna planorbis, FMNH 160957. D. Orpiella concavum, BMNH no registration number. E. Dendrotrochus eva eva, FMNH 175196. Scale bars 1 mm (A), 2 mm (BD, F), 5 mm (E). Abbreviations: alb, albumen gland; bc, bursa copulatrix; ca, carrefour; e, eggs / embryos; ep, epiphallus; fov, free oviduct; her, hermaphrodite duct; ov, ovotestis; p, penis; pr, prostate; prm, penis retractor muscle; puo, post-uterine oviduct; t, talon; ut, uterus; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5312822" httpUri="https://zenodo.org/record/5312822/files/figure.png" pageId="119" pageNumber="120">41E</figureCitation>
): As for family. Free oviduct of medium length; internal longitudinal pilasters present. Bursa copulatrix inserted at junction of penis and free oviduct; duct of bursa copulatrix wide, distinguishable from bursa copulatrix, internally with irregular pustules. Vagina absent. Penis moderately long; epiphallus enters penis through short verge; penis internally smooth, one circular penis pilaster present, penial diverticulum absent. Penial sheath present, enclosing only penis; penis retractor muscle attached to epiphallus. Epiphallus much longer than penis, simple internally. Epiphallic retractor caecum absent; diverticulum at junction of epiphallus and vas deferens in the form of a short lime-sac (thin-walled, no axial filament). Spermatophore long soft-walled tube closed at both ends, with no sculpture.
</paragraph>
<paragraph id="EC05426EFFE06A7DF2DF72FEFDF1FAFC" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
<taxonomicName id="2BBA39EDFFE06A7DF2DF72FEFEFCFB25" box="[195,279,1173,1198]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" order="Porellales" pageId="119" pageNumber="120" phylum="Marchantiophyta" rank="genus">
<emphasis id="DECE9E7CFFE06A7DF2DF72FEFEFCFB25" box="[195,279,1173,1198]" italics="true" pageId="119" pageNumber="120">Radula</emphasis>
</taxonomicName>
(
<figureCitation id="74815EEBFFE06A7DF33572FEFE39FB24" box="[297,466,1173,1199]" captionStart="FIGURE 42" captionStartId="118.[151,255,1168,1192]" captionTargetBox="[257,1330,194,1144]" captionTargetId="figure-236@118.[257,1330,194,1144]" captionTargetPageId="118" captionText="FIGURE 42. Radulae. AC. Trochomorpha rubens, FMNH 175247. DF. Videna planorbis, FMNH 160957. GI. Dendrotrochus eva eva, FMNH 15719. JL. Orpiella concavum, BMNH no registration number. Scale bars 20 mm (AL)." figureDoi="http://doi.org/10.5281/zenodo.5312826" httpUri="https://zenodo.org/record/5312826/files/figure.png" pageId="119" pageNumber="120">Figure 42GI</figureCitation>
): Relatively long and narrow. Central tooth absent from single radula examined; first two lateral teeth with small ectocones; mesocones lanceolate; shorter than tooth base. Lateral and marginal tooth fields distinguishable. Lateral teeth with endocone absent; ectocone equal in size to first lateral tooth ectocone; mesocone shorter than tooth base. Marginal teeth with endocones absent; ectocones approximately same length and breadth as mesocone, not subdivided into extra teeth. Radular formula (~104.19.1.19.~104) × 110 rows.
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFE06A7DF2DA73EEFDAAF8A4" pageId="119" pageNumber="120" type="discussion">
<paragraph id="EC05426EFFE06A7DF2DA73EEFED1F99C" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
<emphasis id="DECE9E7CFFE06A7DF2DA73EEFED4FA14" bold="true" box="[198,319,1413,1439]" pageId="119" pageNumber="120">Remarks.</emphasis>
The anatomy of the species examined here shows strong similarities to the
<typeStatus id="3301FCCCFFE06A7DF6DC73EEFB19FA14" box="[1216,1266,1413,1439]" pageId="119" pageNumber="120">type</typeStatus>
species of the subgenus
<taxonomicName id="2BBA39EDFFE06A7DF31173C6FE4DFA4D" authorityName="Solem" authorityYear="1959" box="[269,422,1453,1478]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="subGenus" subGenus="Santotrochus">
<emphasis id="DECE9E7CFFE06A7DF31173C6FE4DFA4D" box="[269,422,1453,1478]" italics="true" pageId="119" pageNumber="120">Santotrochus</emphasis>
</taxonomicName>
,
<taxonomicName id="2BBA39EDFFE06A7DF3A473C6FB96FA4C" authority="(Hartman, 1889) (Schileyko 2002 b)" authorityName="Schileyko" authorityYear="2002" baseAuthorityName="Hartman" baseAuthorityYear="1889" box="[440,1149,1453,1479]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="species" species="layardi">
<emphasis id="DECE9E7CFFE06A7DF3A473C6FD2EFA4D" box="[440,709,1453,1478]" italics="true" pageId="119" pageNumber="120">Dendrotrochus layardi</emphasis>
(Hartman, 1889) (
<bibRefCitation id="882B3F9FFFE06A7DF1BA73C6FB9EFA4C" author="Schileyko, A. A." box="[934,1141,1453,1479]" pageId="119" pageNumber="120" refId="ref76904" refString="Schileyko, A. A. (2002 b) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 9: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko 2002b</bibRefCitation>
)
</taxonomicName>
. In particular, both species share the presence of a penial verge and a large circular penial pilaster (referred to as a stimulator by
<bibRefCitation id="882B3F9FFFE06A7DF74273BEFEC6F99C" author="Schileyko, A. A." pageId="119" pageNumber="120" refId="ref76904" refString="Schileyko, A. A. (2002 b) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 9: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko 2002b</bibRefCitation>
).
</paragraph>
<paragraph id="EC05426EFFE06A7DF2DF704EFDBCF904" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
<taxonomicName id="2BBA39EDFFE06A7DF2DF704EFEB7F9B5" authorityName="Solem" authorityYear="1959" box="[195,348,1573,1598]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="subGenus" subGenus="Santotrochus">
<emphasis id="DECE9E7CFFE06A7DF2DF704EFEB7F9B5" box="[195,348,1573,1598]" italics="true" pageId="119" pageNumber="120">Santotrochus</emphasis>
</taxonomicName>
includes 23 species and is limited to
<collectingCountry id="94AD02FEFFE06A7DF131704EFC66F9B4" box="[813,909,1573,1599]" name="Vanuatu" pageId="119" pageNumber="120">Vanuatu</collectingCountry>
(
<bibRefCitation id="882B3F9FFFE06A7DF182704EFB87F9B4" author="Schileyko, A. A." box="[926,1132,1573,1599]" pageId="119" pageNumber="120" refId="ref76904" refString="Schileyko, A. A. (2002 b) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 9: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko 2002b</bibRefCitation>
).
<bibRefCitation id="882B3F9FFFE06A7DF69D704EFAC9F9B4" author="Solem, A." box="[1153,1314,1573,1599]" pageId="119" pageNumber="120" pagination="1 - 238" refId="ref77542" refString="Solem, A. (1959) Systematics of the land and fresh-water Mollusca of the New Hebrides. Fieldiana Zoology, 43, 1 - 238." type="journal article" year="1959">Solem (1959)</bibRefCitation>
described
<taxonomicName id="2BBA39EDFFE06A7DF28B7026FEDBF9ED" authorityName="Solem" authorityYear="1959" box="[151,304,1613,1638]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="subGenus" subGenus="Santotrochus">
<emphasis id="DECE9E7CFFE06A7DF28B7026FEDBF9ED" box="[151,304,1613,1638]" italics="true" pageId="119" pageNumber="120">Santotrochus</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="2BBA39EDFFE06A7DF01F7026FD59F9ED" authorityName="Pilsbry" authorityYear="1894" box="[515,690,1613,1638]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="DECE9E7CFFE06A7DF01F7026FD59F9ED" box="[515,690,1613,1638]" italics="true" pageId="119" pageNumber="120">Dendrotrochus</emphasis>
</taxonomicName>
and this usage is followed in the current study, although
<bibRefCitation id="882B3F9FFFE06A7DF74A7026FEDFF904" author="Schileyko, A. A." pageId="119" pageNumber="120" refId="ref76904" refString="Schileyko, A. A. (2002 b) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 9: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko (2002b)</bibRefCitation>
raised it to generic rank.
</paragraph>
<paragraph id="EC05426EFFE06A7DF2DF70F6FEFBF954" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
The varying taxonomic placement of
<taxonomicName id="2BBA39EDFFE06A7DF0B670F6FCB2F93D" authorityName="Pilsbry" authorityYear="1894" box="[682,857,1693,1718]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="DECE9E7CFFE06A7DF0B670F6FCB2F93D" box="[682,857,1693,1718]" italics="true" pageId="119" pageNumber="120">Dendrotrochus</emphasis>
</taxonomicName>
is outlined above (Family
<taxonomicName id="2BBA39EDFFE06A7DF6DF70F6FA72F93C" box="[1219,1433,1693,1719]" class="Gastropoda" family="Trochomorphidae" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="family">Trochomorphidae</taxonomicName>
: Remarks).
</paragraph>
<paragraph id="EC05426EFFE06A7DF2DF7086FDAAF8A4" blockId="119.[151,1437,453,1839]" pageId="119" pageNumber="120">
<taxonomicName id="2BBA39EDFFE06A7DF2DF7086FE3FF88D" authority="eva" authorityName="eva (Pfeiffer" authorityYear="1854" box="[195,468,1773,1798]" class="Gastropoda" family="Trochomorphidae" genus="Dendrotrochus" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="subSpecies" species="eva" subSpecies="eva">
<emphasis id="DECE9E7CFFE06A7DF2DF7086FE3FF88D" box="[195,468,1773,1798]" italics="true" pageId="119" pageNumber="120">Dendrotrochus eva eva</emphasis>
</taxonomicName>
grouped with trochomorphids
<taxonomicName id="2BBA39EDFFE06A7DF1597086FBA1F88D" box="[837,1098,1773,1798]" class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="species" species="rubens">
<emphasis id="DECE9E7CFFE06A7DF1597086FBA1F88D" box="[837,1098,1773,1798]" italics="true" pageId="119" pageNumber="120">Trochomorpha rubens</emphasis>
</taxonomicName>
and
<taxonomicName id="2BBA39EDFFE06A7DF69A7086FABBF88D" box="[1158,1360,1773,1798]" class="Gastropoda" family="Trochomorphidae" genus="Videna" kingdom="Animalia" order="Stylommatophora" pageId="119" pageNumber="120" phylum="Mollusca" rank="species" species="planorbis">
<emphasis id="DECE9E7CFFE06A7DF69A7086FABBF88D" box="[1158,1360,1773,1798]" italics="true" pageId="119" pageNumber="120">Videna planorbis</emphasis>
</taxonomicName>
in the phylogenetic analysis (
<figureCitation id="74815EEBFFE06A7DF3BF717EFDD9F8A4" box="[419,562,1813,1839]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="13.[151,255,1878,1902]" captionStartId-1="14.[151,255,1728,1752]" captionStartId-2="15.[151,255,1916,1940]" captionTargetBox-0="[174,1417,215,1826]" captionTargetBox-1="[183,1411,208,1687]" captionTargetBox-2="[185,1397,280,1882]" captionTargetId-0="figure-53@13.[154,1439,197,1848]" captionTargetId-1="figure-109@14.[160,1425,194,1701]" captionTargetId-2="figure-69@15.[185,1410,274,1888]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURE 1. Majority-rule consensus tree (1,743 trees, length 117 steps) from the morphological analysis containing representatives of all groups. Families and subfamilies are shaded and are labelled in bold if the group is not monophyletic." captionText-1="FIGURE 2. Majority-rule consensus tree (646 trees, length 106 steps) from the morphological analysis with Dyakiidae and Urocyclidae excluded. Families and subfamilies are shaded and are labelled in bold if the group is not monophyletic." captionText-2="FIGURE 3. Majority-rule consensus tree (770 trees, length 117 steps) resulting from the morphological analysis with two additional mantle lobe characters. Families and subfamilies are shaded and are labelled in bold if the group is not monophyletic." figureDoi-0="http://doi.org/10.5281/zenodo.5312714" figureDoi-1="http://doi.org/10.5281/zenodo.5312716" figureDoi-2="http://doi.org/10.5281/zenodo.5312718" httpUri-0="https://zenodo.org/record/5312714/files/figure.png" httpUri-1="https://zenodo.org/record/5312716/files/figure.png" httpUri-2="https://zenodo.org/record/5312718/files/figure.png" pageId="119" pageNumber="120">Figures 13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>