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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.258.4675" ID-GBIF-Dataset="59fc613f-73d2-4bd2-91b4-a886654a6619" ID-PMC="PMC3591754" ID-Pensoft-Pub="1313-2970-258-17" ID-PubMed="23653497" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-258-17" ModsDocOrigin="ZooKeys 258" ModsDocTitle="A redescription of Rhysida celeris (Humbert &amp; Saussure, 1870), with a proposal of eight new synonyms (Scolopendromorpha, Scolopendridae, Otostigminae)" checkinTime="1451247782064" checkinUser="pensoft" docAuthor="Chagas-Junior, Amazonas" docDate="2013" docId="301B6C5E6DD81E1C34B35884B11F3071" docLanguage="en" docName="ZooKeys 258: 17-29" docOrigin="ZooKeys 258" docSource="http://dx.doi.org/10.3897/zookeys.258.4675" docTitle="Rhysida celeris Humbert &amp; Saussure 1870" docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="B1551411FFA35413E6093D3A4F3FFFE8" masterDocTitle="A redescription of Rhysida celeris (Humbert &amp; Saussure, 1870), with a proposal of eight new synonyms (Scolopendromorpha, Scolopendridae, Otostigminae)" masterLastPageNumber="29" masterPageNumber="17" pageNumber="18" updateTime="1668155144758" updateUser="ExternalLinkService">
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<mods:title>A redescription of Rhysida celeris (Humbert &amp; Saussure, 1870), with a proposal of eight new synonyms (Scolopendromorpha, Scolopendridae, Otostigminae)</mods:title>
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<mods:namePart>Chagas-Junior, Amazonas</mods:namePart>
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<mods:date>2013</mods:date>
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<mods:number>258</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.258.4675</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.258.4675</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3509143" ID-GBIF-Taxon="152039650" ID-Zenodo-Dep="3509143" LSID="urn:lsid:plazi:treatment:301B6C5E6DD81E1C34B35884B11F3071" httpUri="http://treatment.plazi.org/id/301B6C5E6DD81E1C34B35884B11F3071" lastPageId="5" lastPageNumber="22" pageId="1" pageNumber="18">
<subSubSection pageId="1" pageNumber="18" type="nomenclature">
<paragraph pageId="1" pageNumber="18">
<taxonomicName LSID="http://species-id.net/wiki/Rhysida_celeris" authority="Humbert &amp; Saussure, 1870" authorityName="Humbert &amp; Saussure" authorityYear="1870" class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="celeris">Rhysida celeris (Humbert &amp; Saussure, 1870)</taxonomicName>
Figures 1-10
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="18" type="reference_group">
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Branchiostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiostoma celer" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="celer">Branchiostoma celer</taxonomicName>
Humbert &amp; Saussure, 1870: 202;
<bibRefCitation pageId="1" pageNumber="18">Saussure and Humbert 1872</bibRefCitation>
: 122;
<bibRefCitation author="Kohlrausch, E" journalOrPublisher="Archiv fur Naturgeschichte" pageId="8" pageNumber="25" pagination="50 - 132" title="Gattungen und Arten der Scolopendriden." volume="47" year="1881">Kohlrausch 1881</bibRefCitation>
: 69;
<bibRefCitation author="Meinert, F" journalOrPublisher="Proceedings of the American Philosophical Society" pageId="9" pageNumber="26" pagination="161 - 232" title="Myriapoda Musei Cantabrigensis, Mass. Part I. Chilopoda." volume="28" year="1886">Meinert 1886</bibRefCitation>
: 183.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="celeris">Rhysida celeris</taxonomicName>
Pocock, 1896: 27;
<bibRefCitation author="Kraepelin, K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pageId="8" pageNumber="25" pagination="1 - 276" title="Revision der Scolopendriden." url="10.1080/00222938500770151" volume="20" year="1903">Kraepelin 1903</bibRefCitation>
: 149 (+
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida aspera" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="aspera">Rhysida aspera</taxonomicName>
);
<bibRefCitation pageId="1" pageNumber="18">Attems 1930</bibRefCitation>
: 188;
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="251 - 372" title="Catalogo dos quilopodos da zona neotropica." volume="15" year="1942">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
1942
</bibRefCitation>
: 315; 1974: 119.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Trematoptychus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trematoptychus celeris" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="celeris">Trematoptychus celeris</taxonomicName>
Chamberlin, 1914: 181.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris subsp. andina" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="celeris" subSpecies="andina">Rhysida celeris andina</taxonomicName>
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
, 1953: 112; 1974: 119. Syn. n.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida caripensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="caripensis">Rhysida caripensis</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 2002: 51. Syn. n.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida neoespartana" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="neoespartana">Rhysida neoespartana</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 2002: 52. Syn. n.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida guayanica" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="guayanica">Rhysida guayanica</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 2002: 53. Syn. n.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida maritima" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="maritima">Rhysida maritima</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 2002: 54. Syn. n.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida monaguensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="monaguensis">Rhysida monaguensis</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 2002: 55. Syn. n.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida porlamarensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="porlamarensis">Rhysida porlamarensis</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 2002: 56. Syn. n.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida sucupanensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="sucupanensis">Rhysida sucupanensis</taxonomicName>
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
, 2002: 58. Syn. n.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="18" type="type material examined">
<paragraph pageId="1" pageNumber="18">Type material examined.</paragraph>
<paragraph pageId="1" pageNumber="18">
Venezuela -
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida caripensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="caripensis">Rhysida caripensis</taxonomicName>
- Holotype (MIZA0015958, ex. MAGS 680a) and paratype (MIZA0015958, ex. MAGS 680b), Edo. Monagas, Caripe, Caripe (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.2">10°12'00&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-63.5">63°30'00&quot;W</geoCoordinate>
), ix.1991, Dora
<normalizedToken originalValue="Padrón">Padron</normalizedToken>
de
<normalizedToken originalValue="García">Garcia</normalizedToken>
leg.;
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida neoespartana" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="neoespartana">Rhysida neoespartana</taxonomicName>
- Holotype (MIZA0015954, ex. MAGS 319a) and paratype (MIZA0015954, ex. MAGS 319b), Edo. Nueva Esparta, La
<normalizedToken originalValue="Asunción">Asuncion</normalizedToken>
, Marcano (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="11.033334">11°02'00&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-63.862778">63°51'46&quot;W</geoCoordinate>
), 1988
<normalizedToken originalValue="1989">-1989-</normalizedToken>
1991, Dora
<normalizedToken originalValue="Padrón">Padron</normalizedToken>
de Garcia leg.;
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida guayanica" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="guayanica">Rhysida guayanica</taxonomicName>
- Holotype (MIZA0015961, ex. MAGS 738a) and paratype (MIZA0015961, ex. MAGS 738b), Edo.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
, Los Pijiguaos,
<normalizedToken originalValue="Cedeño">Cedeno</normalizedToken>
(
<geoCoordinate direction="north" orientation="latitude" precision="15" value="6.588889">06°35'20&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-66.753334">66°45'12&quot;W</geoCoordinate>
), 01.v.1992,
<normalizedToken originalValue="José">Jose</normalizedToken>
Manuel Ayala leg.;
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida maritima" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="maritima">Rhysida maritima</taxonomicName>
- Holotype (MIZA0015957, ex. MAGS 580a), Edo. Carabobo, Goaigoaza, Puerto Cabello (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.493334">10°29'36&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-68.04667">68°02'48&quot;W</geoCoordinate>
) in 19.iii.1990, and paratype (MIZA0015963, ex. MAGS 1002), Edo. Carabobo, San Esteban, Puerto Cabello, in 18.x.1998, both collected by A. R. Delgado de
<normalizedToken originalValue="González">Gonzalez</normalizedToken>
&amp; Manuel
<normalizedToken originalValue="Ángel">Angel</normalizedToken>
<normalizedToken originalValue="González">Gonzalez</normalizedToken>
Sponga;
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida monaguensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="monaguensis">Rhysida monaguensis</taxonomicName>
- Holotype (MIZA0016084, ex. MAGS 970a) and paratype (MIZA0016084, ex. MAGS 970b), Edo. Monagas, Uverito, Sotillo (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.702778">08°42'10&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-63.366665">63°22'00&quot;W</geoCoordinate>
), 30.iv.1998, Oswaldo Fuentes leg.;
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida porlamarensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="porlamarensis">Rhysida porlamarensis</taxonomicName>
- Holotype (MIZA0015953, ex. MAGS 315a and paratype MIZA0015953, ex. MAGS 315b), Edo. Nueva Esparta, Porlamar, Calle San Rafael, Porlamar (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.958333">10°57'30&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-63.85">63°51'00&quot;W</geoCoordinate>
), 30.iii.1988, Carlos Contreras leg.;
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida sucupanensis" order="Scolopendromorpha" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="sucupanensis">Rhysida sucupanensis</taxonomicName>
- Holotype (MIZA0015908, ex. MAGS 63a) and paratype (MIZA0015908, ex. MAGS 63b), Edo. Delta Amaruco, Sucupana, Casacoima (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.683333">08°41'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-61.8">61°48'W</geoCoordinate>
), 16.ix.1987, Pedro Delgado leg.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19" type="additional material examined">
<paragraph pageId="2" pageNumber="19">
<pageBreakToken pageId="2" pageNumber="19" start="start">Additional</pageBreakToken>
material examined.
</paragraph>
<paragraph lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19">
Jamaica - BMNH - 1 specimen; Dominican Republic - ZMH - 4 specimens,
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
, 5.vii.1905, C. Gagzol; ZMH - 1 specimen,
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
, Bay V.
<normalizedToken originalValue="Samaná">Samana</normalizedToken>
, xii.1894, Bock; Haiti - ZMH - 4 specimens, Port au Prince, 29.v.1901, Dr. Rouch; Montserrat - BMNH - 1 specimen, B.W.I., ii.1932; Costa Rica - ZMH - 1 specimen, Puerto
<normalizedToken originalValue="Simón">Simon</normalizedToken>
, 19.xi.1899, R. Mull; Suriname - ZMH - 2 specimens, Paramaribo, 28.v.1910, C. Hellesl; Colombia - IAVH - 1 specimen, Vichada, Cumaribo, Cgto. Santa Rita, PNN El Tuparro, 14-16-II-2004, I. Quintero, E. Gonzalez; IAVH - 1 specimen, Vichada, Cumaribo, Selva de Mataven, 31-III-02-IV-2007, L.E. Franco; IAVH - 1 specimen, Vichada, Gaviotas, 31-VI-1995; IAVH - 1 specimen,
<normalizedToken originalValue="Caquetá">Caqueta</normalizedToken>
, Solano, PNN Chiribiquete, 24-26-II- 2000, M. Ospina &amp; E.
<normalizedToken originalValue="González">Gonzalez</normalizedToken>
; ICN-M.Ch-0005, 1 specimen, Amazonas, Leticia, Via Torame, X-2000, Sist. Animal; ICN-M.Ch-0035, 1 specimen, Sucre, San Onofre, Boca cerrada, canal del dique, 1-IV-2000, E. Ulloa; ICN-M.Ch-0037, 1 specimen, Sucre, Galeras, Vereda corozera, 14-I-1999, E.
<normalizedToken originalValue="Hernández">Hernandez</normalizedToken>
; ICN-M.Ch-0039, 1 specimen, Meta, Vereda
<pageBreakToken pageId="3" pageNumber="20" start="start">Apiay</pageBreakToken>
, sector el bosque villa
<normalizedToken originalValue="Lolé">Lole</normalizedToken>
, 23-XII-2001, M. Rojas; ICN-M.Ch-0097, 1 specimen, Casanare, Aguazul, Vda. El Charte, Finca Namaste, 20-IX-1996, Estudiantes Biol UN; ICN-M.Ch-0100, 1 specimen, Tolima, San Luis, 13-VI-1992, A. Castillo; ICN-M.Ch-0136, 1 specimen, Meta, San Martin, 1-15 -IV-2011, W. Galvis; ICN-M.Ch-0138, 1 specimen,
<normalizedToken originalValue="Vaupés">Vaupes</normalizedToken>
, Est. Biol.
<normalizedToken originalValue="Caparù">Caparu</normalizedToken>
, 2002-2004, J.
<normalizedToken originalValue="Pinzón">Pinzon</normalizedToken>
; Ecuador - QCAZ - 3 specimens, Francisco Orelanna, La Joya de Los Sachas, 3.vi.2006, J. Mideros; QCAZ - 1 specimen, Francisco Orelanna, PN Yasumi, M.J. Tamariz; QCAZ - 1 specimen, Napo, Tena, 19.ix.2004; Argentina - ZMUC - 11 specimens, Riacho del Oro, 17.06.1899, W. Sorensen; ZMUC - 2 specimens, 19.08.1895.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="20" type="diagnosis">
<paragraph pageId="3" pageNumber="20">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="20">
General body color light blue or olive green, sternites and legs light blue or yellowish; prefemur and femur of the ultimate legs light blue, sometimes tibiae and tarsi are pale. Antennae with 17 to 21 articles, first two articles, dorsal surface and
<normalizedToken originalValue="¾">3/4</normalizedToken>
of ventral surface of third articles glabrous. Cephalic plate smooth, without sutures or depressions; tooth plates wider than high, 4+4, 4+5 or 5+5 teeth. Tergites smooth; complete paramedian sutures present from tergites 3-5 to 19-20, margination from tergites 5-9 to 21. Posterior border of tergite 21 ending in an obtuse angle. A pair of spiracles at 7th leg -bearing segment. Sternites 2, 3 or 4 to 19 with anterior incomplete paramedian sutures, but without depressions. Coxopleuron not prolonged, very short process with two small apical spines. Legs 1 to 16 (or 17) or 2 to 18 (sometimes 17) with two tarsal spurs, 19 (or 20) with one and 21 without; ultimate legs long, prefemur without spines.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20" type="description">
<paragraph pageId="3" pageNumber="20">Redescription.</paragraph>
<paragraph lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20">
Body length from 40 to 70 mm. General body color light blue or olive green, sternites and legs light blue or yellowish; prefemur and femur of the ultimate legs light or dark blue, but sometimes tibia and tarsi are pale or light blue. Antennae with 17 to 21 articles, first two and dorsal surface and
<normalizedToken originalValue="¾">3/4</normalizedToken>
of the ventral surface of the third article are glabrous; antennae reaching the posterior margin of tergite 5. Cephalic plate smooth, with a median sulcus but without sutures or depressions (Fig. 1), wider than long. Anterior margin of cephalic plate right-angled, with four ocelli in each side, posterior margin slightly rounded (Fig. 1). Forcipular coxosternum without depressions (Fig. 2); tooth plates with 4+4, 4+5 or 5+5 teeth; the inner two teeth are closer to each other than to the external teeth (Fig. 3). Each tooth plate with a long seta in the center; the tooth plates with obtuse angled basal suture. Trochanteropreforal process well-developed, long and ending as a point, with three to five denticles laterally, these sometimes not visible (Fig. 4). Tergites smooth, wider than long. Tergite 1 without sutures, its anterior border overlapping the posterior border of the cephalic plate. Tergites 3-5 to 19-20 with complete paramedian sutures (Fig. 5). Tergites 5 to 21 marginate, usually from tergite 11 (Fig. 5), but sometimes also from tergite 16. The margination is clearly visible in the anterior part of the tergites; tergite 21 with a slight posterior depression and triangular posterior margin (Fig. 7). A pair of spiracles on the 7th leg -bearing segment. Sternites smooth, wider than long. Sternites 2 (4)-19 with short incomplete anterior sutures (Fig. 6). However, sometimes, they are overlapped by the posterior margin of the previous sternite and visible only at sternites 5-18. Sternite 21 longer than wide; converging caudad and with straight or slightly concave posterior margin (Fig. 8). Coxopleural pores of several sizes, and numerous; the pore-field occupies nearly all surface of
<pageBreakToken pageId="4" pageNumber="21" start="start">coxopleuron</pageBreakToken>
except for its dorsal and posterior parts, which are free of pores, posterior part of coxopleuron with a longitudinal depression (Fig. 9); very short coxopleural process with two apical spines (Fig. 10); apical spines may be combined as 1+2 or 2+3. Leg 1 with one femoral spur and legs 1 and 2 with one tibial spurs; legs 1 to 16, 17 or from 2 to 18 with two tarsal spurs, 19 and 20 with one and 21 without. Sometimes legs 20 lack spurs. Pretarsi of all legs with two accessory spines. Ultimate legs long and slender (12 mm to 22 mm). Prefemur without spines.
</paragraph>
<caption pageId="4" pageNumber="21">
<paragraph pageId="4" pageNumber="21">
Figure 1-4. 1
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="celeris">Rhysida celeris</taxonomicName>
from Ecuador. Cephalic plate 2 Forcipular Coxosternum 3 Tooth plates 4 Forcipular trochanteroprefemur process. Scale bars 1 mm.
</paragraph>
</caption>
<caption pageId="4" pageNumber="21">
<paragraph pageId="4" pageNumber="21">Figure 5-8. 5 Tergites 11, 12 and 13 6 Sternites 4, 5 and 6 7 Tergite 21 8 Segment 21 showing sternite 21 and coxopleuron. Scale bars 1 mm.</paragraph>
</caption>
<caption pageId="4" pageNumber="21">
<paragraph pageId="4" pageNumber="21">Figure 9-10. 9 Segment 21 showing the coxopleuron 10 Detail of the terminal part of the coxopleuron showing the spines. Scale bar 1 mm.</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21" type="taxonomic remarks">
<paragraph pageId="4" pageNumber="21">Taxonomic remarks.</paragraph>
<paragraph pageId="4" pageNumber="21">
Except for
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida maritima" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="maritima">Rhysida maritima</taxonomicName>
where the holotype and paratype were separated and given different numbers (e.g., holotype, MAGS 580a, and, paratype, MAGS 1002), the type specimens of all other species described by
<bibRefCitation author="Gonzalez-Sponga, MA" journalOrPublisher="Aula Y Ambiente" pageId="8" pageNumber="25" pagination="49 - 60" title="Miriapodos de Venezuela. Descripcion de siete nuevas especies del genero Rhysida y redescripcion de Rhysida longipes Newport, 1845 (Chilopoda: Scolopendridae)." volume="4" year="2002">
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
(2002)
</bibRefCitation>
were left united and were labeled with identical numbers (e.g., holotype, MAGS7400, and a paratype, MAGS740b). The acronym MAGS in the collection numbers stands for 'Manuel
<normalizedToken originalValue="Ángel">Angel</normalizedToken>
Gonzalez
<normalizedToken originalValue="Sponga">Sponga'</normalizedToken>
and indicates that the specimens belong to his private collection, which after his death was transferred to the myriapodological collection of Museo del Intituto de
<normalizedToken originalValue="Zoología">Zoologia</normalizedToken>
<normalizedToken originalValue="Agrícola">Agricola</normalizedToken>
Francisco
<normalizedToken originalValue="Fernández">Fernandez</normalizedToken>
<normalizedToken originalValue="Yèpez">Yepez</normalizedToken>
, Maracay, Venezuela. The type series of
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida caripensis" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="caripensis">Rhysida caripensis</taxonomicName>
,
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida neoespartana" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="neoespartana">Rhysida neoespartana</taxonomicName>
,
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida monaguensis" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="monaguensis">Rhysida monaguensis</taxonomicName>
and
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida porlamarensis" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="porlamarensis">Rhysida porlamarensis</taxonomicName>
consists of more than two specimens. The recognition of the holotype is only possible by measuring the total body length, cephalic plate and ultimate legs and by checking them against the original publication (
<bibRefCitation author="Gonzalez-Sponga, MA" journalOrPublisher="Aula Y Ambiente" pageId="8" pageNumber="25" pagination="49 - 60" title="Miriapodos de Venezuela. Descripcion de siete nuevas especies del genero Rhysida y redescripcion de Rhysida longipes Newport, 1845 (Chilopoda: Scolopendridae)." volume="4" year="2002">
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
2002
</bibRefCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="21">
The results of the review of the type-material described by
<bibRefCitation author="Gonzalez-Sponga, MA" journalOrPublisher="Aula Y Ambiente" pageId="8" pageNumber="25" pagination="49 - 60" title="Miriapodos de Venezuela. Descripcion de siete nuevas especies del genero Rhysida y redescripcion de Rhysida longipes Newport, 1845 (Chilopoda: Scolopendridae)." volume="4" year="2002">
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
(2002)
</bibRefCitation>
are summarized in Table 1. The type of
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris subsp. andina" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="celeris" subSpecies="andina">Rhysida celeris andina</taxonomicName>
was not examined, but all characters (number of antennal articles, complete paramedian sutures and the margination of the tergites, sternite sutures, forcipular tooth-plates and tarsal spurs) fall within the intraspecific variation found in
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="celeris">Rhysida celeris</taxonomicName>
. Therefore,
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris subsp. andina" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="celeris" subSpecies="andina">Rhysida celeris andina</taxonomicName>
was also included in this revision, but the proposed synonymy is based solely on critical analysis of the published information (
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="109 - 151" title="Quilopodos, aranhas e escorpioes enviados ao Instituto Butantan para determinacao." volume="25" year="1953">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
1953
</bibRefCitation>
).
</paragraph>
<caption ID-Table-UUID="4A0CE33CBEE47ADF4466C5AD42D7107A" httpUri="http://table.plazi.org/id/4A0CE33CBEE47ADF4466C5AD42D7107A" pageId="4" pageNumber="21">
<paragraph pageId="4" pageNumber="21">
Table 1. Characters of the species considered in this study. Information based on personal examination of types and original data (in parentheses) as published by
<bibRefCitation author="Gonzalez-Sponga, MA" journalOrPublisher="Aula Y Ambiente" pageId="8" pageNumber="25" pagination="49 - 60" title="Miriapodos de Venezuela. Descripcion de siete nuevas especies del genero Rhysida y redescripcion de Rhysida longipes Newport, 1845 (Chilopoda: Scolopendridae)." volume="4" year="2002">
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
(2002)
</bibRefCitation>
and
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="43 - 362" title="Os quilopodos do Brasil." volume="13" year="1939">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
(1939
</bibRefCitation>
,
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="109 - 151" title="Quilopodos, aranhas e escorpioes enviados ao Instituto Butantan para determinacao." volume="25" year="1953">1953</bibRefCitation>
).? = not described.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="21">
<table pageId="4" pageNumber="21">
<tr pageId="4" pageNumber="21">
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida caripensis" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="caripensis">Rhysida caripensis</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida neoespartana" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="neoespartana">Rhysida neoespartana</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida guayanica" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="guayanica">Rhysida guayanica</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida maritima" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="maritima">Rhysida maritima</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida monaguensis" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="monaguensis">Rhysida monaguensis</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida porlamarensis" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="porlamarensis">Rhysida porlamarensis</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida socupanensis" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="socupanensis">Rhysida socupanensis</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="21" rowspan="1">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris subsp. andina" order="Scolopendromorpha" pageId="4" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="celeris" subSpecies="andina">Rhysida celeris andina</taxonomicName>
</th>
</tr>
</table>
</paragraph>
<paragraph pageId="5" pageNumber="22">
<pageBreakToken pageId="5" pageNumber="22" start="start">His</pageBreakToken>
descriptions are based only on the holotypes, and he did not describe the variation of some characters, such as the number of antennal articles, the distribution of paramedian sutures and the margination of tergites and the number of teeth of forcipular tooth-plates. For example, complete paramedian sutures are present in all of the
<normalizedToken originalValue="González-Spongas">Gonzalez-Sponga's</normalizedToken>
(2002) species, however, the distribution of sutures vary. The complete paramedian sutures on tergite 3 are not as evident as on tergite 4, but the sutures always end at tergite 20. The number of teeth in forcipular tooth-plates also vary from 3+3 to 5+5.
</paragraph>
<paragraph pageId="5" pageNumber="22">
Two other important characters that were omitted or were poorly described by
<bibRefCitation author="Gonzalez-Sponga, MA" journalOrPublisher="Aula Y Ambiente" pageId="8" pageNumber="25" pagination="49 - 60" title="Miriapodos de Venezuela. Descripcion de siete nuevas especies del genero Rhysida y redescripcion de Rhysida longipes Newport, 1845 (Chilopoda: Scolopendridae)." volume="4" year="2002">
<normalizedToken originalValue="González-Sponga">Gonzalez-Sponga</normalizedToken>
(2002)
</bibRefCitation>
are the incomplete paramedian sutures on the sternites and the distribution of tarsal spurs on the legs. The incomplete paramedian sutures were not mentioned in his paper, although they are clearly visible from sternites 1 to 18, being more evident on sternites 6-15. The tarsal spurs were only described for
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida guayanica" order="Scolopendromorpha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="guayanica">Rhysida guayanica</taxonomicName>
,
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida monaguensis" order="Scolopendromorpha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="monaguensis">Rhysida monaguensis</taxonomicName>
and
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida porlamarensis" order="Scolopendromorpha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="porlamarensis">Rhysida porlamarensis</taxonomicName>
, but their distribution were not indicated.
</paragraph>
<paragraph pageId="5" pageNumber="22">
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris" order="Scolopendromorpha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="celeris">Rhysida celeris</taxonomicName>
was originally described from
<normalizedToken originalValue="Carolina">'Carolina'</normalizedToken>
(
<bibRefCitation author="Humbert, A" journalOrPublisher="Revue et Magasin de Zoologie" pageId="8" pageNumber="25" pagination="196 - 205" title="Myriapoda nova Americana, Series 2." volume="22" year="1870">Humbert and Saussure 1870</bibRefCitation>
,
<bibRefCitation pageId="5" pageNumber="22">Saussure and Humbert 1872</bibRefCitation>
,
<bibRefCitation author="Underwood, LM" journalOrPublisher="Entomologica Americana" pageId="9" pageNumber="26" pagination="61 - 65" title="The Scolopendridae of the United States." volume="3" year="1887">Underwood 1887</bibRefCitation>
) and later on recorded also from Georgia, USA (
<bibRefCitation author="Pocock, RI" journalOrPublisher="Journal of Linnean Society of London, Zoology" pageId="9" pageNumber="26" pagination="454 - 473" title="Contributions to our knowledge of the arthropod fauna of the West Indies. Part II. Chilopoda." volume="24" year="1893">Pocock 1893</bibRefCitation>
,
<bibRefCitation author="Kraepelin, K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pageId="8" pageNumber="25" pagination="1 - 276" title="Revision der Scolopendriden." url="10.1080/00222938500770151" volume="20" year="1903">Kraepelin 1903</bibRefCitation>
,
<bibRefCitation pageId="5" pageNumber="22">Attems 1930</bibRefCitation>
).
<bibRefCitation author="Crabill, RE" journalOrPublisher="Natural Museum" pageId="8" pageNumber="25" pagination="1 - 15" title="A new American genus of cryptopid centipede, with an annotated key to the scolopendromorph genera from America north of Mexico. Proceedings of U. S." volume="111" year="1960">Crabill (1960)</bibRefCitation>
stated that the North American records were not corroborated, and
<bibRefCitation author="Shelley, RM" journalOrPublisher="Virginia Museum of Natural History Memoir" pageId="9" pageNumber="26" pagination="1 - 108" title="A synopsis of the North American centipedes of the order Scolopendromorpha (Chilopoda)." volume="5" year="2002">Shelley (2002)</bibRefCitation>
deleted
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris" order="Scolopendromorpha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="celeris">Rhysida celeris</taxonomicName>
from the list of
<taxonomicName genus="Scolopendromorpha" lsidName="Scolopendromorpha" pageId="5" pageNumber="22" rank="genus">Scolopendromorpha</taxonomicName>
of United States. Carolina is a common toponym in several countries in the Neotropical region. It exists in Puerto Rico, Cuba, Colombia (in the departments of Cesar, Antioquia and Magdalena), in Ecuador (in the departments of Guayas and Imbabura) and in Brazil. There is also a state of Carolina in the U.S. Virgin Islands, and also &quot;La Carolina&quot; in both, Mexico and Argentina. I did not examine the holotype of
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Rhysida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhysida celeris" order="Scolopendromorpha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="celeris">Rhysida celeris</taxonomicName>
, but it is hosted (in alcohol) at the Myriapodological collection of Museum of Natural History of
<normalizedToken originalValue="Genève">Geneve</normalizedToken>
, Switzerland. The specimen is labeled &quot;
<taxonomicName class="Chilopoda" family="Scolopendridae" genus="Branchiostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiostoma celer" order="Scolopendromorpha" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="celer">Branchiostoma celer</taxonomicName>
Humbert &amp; Saussure, type, from Caroline&quot;. A second label reads &quot;revision R. E. Crabill, back 1970&quot; (Peter Schwendinger, personal communication). I do not know any other additional information on the type locality, therefore, the exact whereabouts of the type locality remains uncertain. However, it is more likely that the specimen had been collected somewhere in Antilles, Central or South America rather than in the USA.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="distribution">
<paragraph pageId="5" pageNumber="22">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="22">
Jamaica, Dominican Republic, Haiti, Nicaragua (
<bibRefCitation author="Meinert, F" journalOrPublisher="Proceedings of the American Philosophical Society" pageId="9" pageNumber="26" pagination="161 - 232" title="Myriapoda Musei Cantabrigensis, Mass. Part I. Chilopoda." volume="28" year="1886">Meinert 1886</bibRefCitation>
), Venezuela (
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="251 - 372" title="Catalogo dos quilopodos da zona neotropica." volume="15" year="1942">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
1942
</bibRefCitation>
,
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="233 - 241" title="Chilopoden von Venezuela (II)." volume="22" year="1959">1959</bibRefCitation>
), Mexico (
<bibRefCitation author="Kraepelin, K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pageId="8" pageNumber="25" pagination="1 - 276" title="Revision der Scolopendriden." url="10.1080/00222938500770151" volume="20" year="1903">Kraepelin 1903</bibRefCitation>
,
<bibRefCitation pageId="5" pageNumber="22">Attems 1930</bibRefCitation>
,
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="251 - 372" title="Catalogo dos quilopodos da zona neotropica." volume="15" year="1942">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
1942
</bibRefCitation>
), Bolivia (
<bibRefCitation author="Silvestri, F" journalOrPublisher="Universita ̀ di Torino" pageId="9" pageNumber="26" pagination="1 - 11" title="Viaggio Del Dott. Alfredo Borelli nel Chaco Boliviano e nella Republica Argentina. IV. Chilopodi e Diplopodi. Bollettino dei Musei di zoologia ed anatomia comparata della R." volume="12" year="1897">Silvestri 1897</bibRefCitation>
), Argentina (
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="251 - 372" title="Catalogo dos quilopodos da zona neotropica." volume="15" year="1942">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
1942
</bibRefCitation>
), Brazil (
<bibRefCitation author="Kraepelin, K" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pageId="8" pageNumber="25" pagination="1 - 276" title="Revision der Scolopendriden." url="10.1080/00222938500770151" volume="20" year="1903">Kraepelin 1903</bibRefCitation>
,
<bibRefCitation pageId="5" pageNumber="22">Attems 1930</bibRefCitation>
,
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="251 - 372" title="Catalogo dos quilopodos da zona neotropica." volume="15" year="1942">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
1942
</bibRefCitation>
,
<bibRefCitation pageId="5" pageNumber="22">Schileyko 2002</bibRefCitation>
), Colombia (
<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Occasional Papers of the Museum of Zoology" pageId="8" pageNumber="25" pagination="1 - 28" title="Results of the Bryant Walker Expeditions of the University of the Michigan to Colombia, 1913, and British Guiana, 1914." volume="97" year="1921">Chamberlin 1921</bibRefCitation>
), Ecuador, Peru (
<bibRefCitation author="Buecherl, W" journalOrPublisher="Memorias do Instituto Butantan" pageId="8" pageNumber="25" pagination="173 - 186" title="Quilopodos da Peru II." volume="22" year="1950">
<normalizedToken originalValue="Bücherl">Buecherl</normalizedToken>
1950
</bibRefCitation>
,
<bibRefCitation pageId="5" pageNumber="22">Kraus 1957</bibRefCitation>
), Suriname and Paraguay (
<bibRefCitation author="Silvestri, F" journalOrPublisher="Bolletino dei Musei di Zoologia ed Anatomia Comparata" pageId="9" pageNumber="26" pagination="1 - 12" title="Viaggio deo dottor Alfredo Borelli nella Republica Argentina e nel Paraguay XIV. Chilopodi e Diplopodi." volume="10" year="1895">Silvestri 1895</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>