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<document id="63FFACCAB0B7E2AC1EC15789B39538ED" ID-CLB-Dataset="8766" ID-DOI="10.11646/zootaxa.4834.2.5" ID-GBIF-Dataset="f725a409-be04-4f11-b538-092af4454641" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403040" ID-ZooBank="C6EC3841-60F7-4A99-AC88-D4FC6310CB83" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609346204277" checkinUser="plazi" docAuthor="Martínez, María José &amp; Martín, Guillermo San" docDate="2020" docId="2634DB6FE671F65BFF14F137919FFB66" docLanguage="en" docName="zootaxa.4834.2.5.pdf" docOrigin="Zootaxa 4834 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Syllis hampirmenyatu Martínez &amp; Martín 2020, n. sp." docType="treatment" docUuid="A335743F-82B8-4CF2-B040-7E2A26E9288B" docUuidSource="ZooBank" docVersion="9" lastPageNumber="246" masterDocId="DA0DA317E67DF654FF83F525920FFFC7" masterDocTitle="Syllidae (Annelida) from East Timor and the Philippines (Pacific Ocean), with the description of three new species of Syllis Savigny in Lamarck, 1818" masterLastPageNumber="263" masterPageNumber="231" pageNumber="243" updateTime="1698802608034" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CE347E61816B2B2763730F1738B07AAC">Syllidae (Annelida) from East Timor and the Philippines (Pacific Ocean), with the description of three new species of Syllis Savigny in Lamarck, 1818</mods:title>
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<mods:namePart id="260A2527700F4C693F7654CBDE394CFB">Martínez, María José</mods:namePart>
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<mods:affiliation id="E9FEDA7C4289C89D7AF284CB611FC422">mariajosemtnez @ outlook. com; https: // orcid. org / 0000 - 0002 - 0467 - 0998 &amp; mariajosemtnez @ outlook. com; https: // orcid. org / 0000 - 0002 - 0467 - 0998</mods:affiliation>
<mods:nameIdentifier id="83D1ED2056E806F696ACB8B938000376" type="email">mariajosemtnez@outlook.com</mods:nameIdentifier>
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<mods:namePart id="F9E0DDC608B0D06F0FA28B0678EABF2C">Martín, Guillermo San</mods:namePart>
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<mods:affiliation id="E4B1B3923A3A5F75BE498A173CC4DA37">Universidad Autónoma de Madrid, Departamento de Biología (Zoología), Facultad de Ciencias, Centro de Investigación en Biodiversidad y Cambio global (CIBC-UAM), Cantoblanco, 28049, Madrid, Spain. &amp; guillermo. sanmartin @ uam. es; https: // orcid. org / 0000 - 0002 - 8360 - 6221</mods:affiliation>
<mods:nameIdentifier id="A1ED7EAE0F23E6DB7FC4848F65BA0A21" type="email">guillermo.sanmartin@uam.es</mods:nameIdentifier>
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<mods:title id="28C9E7580B3D3B0701DC2281B9B2CFFF">Zootaxa</mods:title>
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<mods:date id="1AF2573FE0647B457467DEEF359640A9">2020</mods:date>
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<mods:number id="EBDFFBA5DE1B4E4B5349E19906BA40CF">4834</mods:number>
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<treatment id="2634DB6FE671F65BFF14F137919FFB66" ID-DOI="http://doi.org/10.5281/zenodo.4452285" ID-GBIF-Taxon="177010351" ID-Zenodo-Dep="4452285" LSID="urn:lsid:zoobank.org:act:A335743F-82B8-4CF2-B040-7E2A26E9288B" httpUri="http://treatment.plazi.org/id/2634DB6FE671F65BFF14F137919FFB66" lastPageId="15" lastPageNumber="246" pageId="12" pageNumber="243">
<subSubSection id="E68739F2E671F658FF14F13793E5FBEB" box="[151,490,1042,1069]" pageId="12" pageNumber="243" type="nomenclature">
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<emphasis id="9CE9B66BE671F658FF14F13793E5FBEB" bold="true" box="[151,490,1042,1069]" pageId="12" pageNumber="243">
<taxonomicName id="699D11FAE671F658FF14F13793AFFBEA" ID-CoL="85S3L" authority="Martínez &amp; Martín, 2020" authorityName="Martínez &amp; Martín" authorityYear="2020" box="[151,416,1042,1069]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="243" phylum="Annelida" rank="species" species="hampirmenyatu" status="sp. nov.">
<emphasis id="9CE9B66BE671F658FF14F13793AFFBEA" bold="true" box="[151,416,1042,1069]" italics="true" pageId="12" pageNumber="243">Syllis hampirmenyatu</emphasis>
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<taxonomicNameLabel id="87DA0B10E671F658FE24F13793E5FBEB" box="[423,490,1042,1068]" pageId="12" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="E68739F2E671F658FF14F11391B9FB55" pageId="12" pageNumber="243" type="description">
<paragraph id="AE226A79E671F658FF14F1139313FB97" blockId="12.[151,490,1042,1105]" box="[151,284,1078,1105]" pageId="12" pageNumber="243">
<figureCitation id="36A676FCE671F658FF14F113930DFB96" box="[151,258,1078,1105]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figures 5</figureCitation>
,
<figureCitation id="36A676FCE671F658FE8DF1139313FB97" box="[270,284,1078,1104]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">6</figureCitation>
</paragraph>
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LSID:
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urn:lsid:zoobank.org:act:
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<subSubSection id="E68739F2E671F658FF14F1E490CDFAE4" pageId="12" pageNumber="243" type="materials_examined">
<paragraph id="AE226A79E671F658FF14F1E490CDFAE4" blockId="12.[151,1437,1217,2035]" pageId="12" pageNumber="243">
<emphasis id="9CE9B66BE671F658FF14F1E49375FB1C" bold="true" box="[151,378,1217,1243]" pageId="12" pageNumber="243">Material examined</emphasis>
.
<materialsCitation id="1EF56024E671F658FE0AF1E490B1FAE4" ID-GBIF-Occurrence="3019880301" collectingDate="2012-09-20" collectionCode="AM" collectorName="A. Murray &amp; P. A. Hutchings" country="Timor-Leste" elevation="16" latitude="-8.23" location="East of Atauro Island" longLatPrecision="21" longitude="125.615555" pageId="12" pageNumber="243" specimenCount="1" typeStatus="holotype">
<collectingCountry id="D68A2AE9E671F658FE0AF1E4903BFB1C" box="[393,564,1217,1243]" name="Timor-Leste" pageId="12" pageNumber="243">EAST TIMOR</collectingCountry>
.
<location id="AB423CA2E671F658FDBCF1E4913EFB1C" LSID="urn:lsid:plazi:treatment:2634DB6FE671F65BFF14F137919FFB66:AB423CA2E671F658FDBCF1E4913EFB1C" box="[575,817,1217,1243]" country="Timor-Leste" latitude="-8.23" longLatPrecision="21" longitude="125.615555" name="East of Atauro Island" pageId="12" pageNumber="243">East of Atauro Island</location>
,
<geoCoordinate id="CBA90CBEE671F658FCBEF1E491CEFB1C" box="[829,961,1217,1243]" degrees="08" direction="south" minutes="13" orientation="latitude" pageId="12" pageNumber="243" precision="15" seconds="48" value="-8.23">08°1348”S</geoCoordinate>
<geoCoordinate id="CBA90CBEE671F658FC4AF1E49650FB1B" box="[969,1119,1217,1244]" degrees="125" direction="east" minutes="36" orientation="longitude" pageId="12" pageNumber="243" precision="15" seconds="56" value="125.615555">125°3656”E</geoCoordinate>
, outer reef, coral reef slope, dead coral rubble,
<elevation id="25B08D4AE671F658FEE6F1C093EBFB38" box="[357,484,1253,1279]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="243" unit="m" value="16.0">
<quantity id="6965C79CE671F658FEE6F1C09392FB38" box="[357,413,1253,1279]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="243" unit="m" value="16.0">16 m</quantity>
depth
</elevation>
,
<date id="DA234CB9E671F658FE6DF1C09074FB38" box="[494,635,1253,1279]" pageId="12" pageNumber="243" value="2012-09-20">
<collectingDate id="CA67B551E671F658FE6DF1C09074FB38" box="[494,635,1253,1279]" pageId="12" pageNumber="243" value="2012-09-20">20 Sep 2012</collectingDate>
</date>
, coll.
<collectorName id="03680FAFE671F658FD38F1C0913EFB38" box="[699,817,1253,1279]" pageId="12" pageNumber="243">A. Murray</collectorName>
&amp;
<collectorName id="03680FAFE671F658FCD0F1C091F2FB38" box="[851,1021,1253,1279]" pageId="12" pageNumber="243">P.A. Hutchings</collectorName>
,
<typeStatus id="7126D4DBE671F658FB8BF1C09665FB38" box="[1032,1130,1253,1279]" pageId="12" pageNumber="243" type="holotype">holotype</typeStatus>
(
<collectionCode id="C88CF2BCE671F658FBF4F1C096A9FB38" box="[1143,1190,1253,1279]" country="Australia" name="Australian Museum" pageId="12" pageNumber="243" type="Museum">AM</collectionCode>
W.53024) and 3 paratypes (
<collectionCode id="C88CF2BCE671F658FF62F02C931FFAE4" box="[225,272,1289,1315]" country="Australia" name="Australian Museum" pageId="12" pageNumber="243" type="Museum">AM</collectionCode>
W.45645), one of them used for SEM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E68739F2E671F65BFF44F00896EAFE16" lastPageId="15" lastPageNumber="246" pageId="12" pageNumber="243" type="description">
<paragraph id="AE226A79E671F65BFF44F00896EAFE16" blockId="12.[151,1437,1217,2035]" lastBlockId="15.[151,1437,151,1185]" lastPageId="15" lastPageNumber="246" pageId="12" pageNumber="243">
<emphasis id="9CE9B66BE671F658FF44F008935EFA80" bold="true" box="[199,337,1325,1351]" pageId="12" pageNumber="243">Description</emphasis>
.
<typeStatus id="7126D4DBE671F658FEDCF00893C8FA80" box="[351,455,1325,1351]" pageId="12" pageNumber="243" type="holotype">Holotype</typeStatus>
and one
<typeStatus id="7126D4DBE671F658FDAEF0089082FA80" box="[557,653,1325,1351]" pageId="12" pageNumber="243" type="paratype">paratype</typeStatus>
complete specimens, other two
<typeStatus id="7126D4DBE671F658FC76F008966FFA80" box="[1013,1120,1325,1351]" pageId="12" pageNumber="243" type="paratype">paratypes</typeStatus>
incomplete.
<typeStatus id="7126D4DBE671F658FB71F0089752FA80" box="[1266,1373,1325,1351]" pageId="12" pageNumber="243" type="holotype">Holotype</typeStatus>
, largest complete specimen,
<quantity id="6965C79CE671F658FE2AF07493F6FAAC" box="[425,505,1361,1387]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="243" unit="mm" value="11.0">11 mm</quantity>
long,
<quantity id="6965C79CE671F658FDC0F0749094FAAC" box="[579,667,1361,1388]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="243" unit="mm" value="0.5">0.5 mm</quantity>
wide, with 84 chaetigers. Body elongated, thin, somewhat filiform (
<figureCitation id="36A676FCE671F658FF1CF05092E7FA57" box="[159,232,1397,1424]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5</figureCitation>
AC, 6A), without colour pattern. Prostomium oval, with four red eyes in trapezoidal arrangement (
<figureCitation id="36A676FCE671F658FAECF05092B2FA73" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Fig. 5A</figureCitation>
); eyespots not seen. Palps robust, elongated, somewhat longer than prostomium (
<figureCitation id="36A676FCE671F658FBCBF0BC96AAFA73" box="[1096,1189,1433,1460]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5A</figureCitation>
,
<figureCitation id="36A676FCE671F658FB32F0BC96DAFA73" box="[1201,1237,1433,1460]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">6B</figureCitation>
). Median antenna arising from middle of prostomium, between posterior eyes, approximately twice as long as the combined length of prostomium and palps, with 18 articles (
<figureCitation id="36A676FCE671F658FDFAF0C490D9FA3B" box="[633,726,1505,1532]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5A</figureCitation>
,
<figureCitation id="36A676FCE671F658FD61F0C49109FA3B" box="[738,774,1505,1532]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">6B</figureCitation>
); lateral antennae shorter than median one, with 15 articles (
<figureCitation id="36A676FCE671F658FF1CF32092F4F9E7" box="[159,251,1541,1568]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5A</figureCitation>
,
<figureCitation id="36A676FCE671F658FE85F3209324F9E7" box="[262,299,1541,1568]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">6B</figureCitation>
). Peristomium shorter than subsequent segments; well-visible nuchal organs (
<figureCitation id="36A676FCE671F658FB09F32096EFF9D8" box="[1162,1248,1541,1567]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">Fig. 6B</figureCitation>
, arrows, C); dorsal tentacular cirri similar in length to median antenna, with 1617 articles (
<figureCitation id="36A676FCE671F658FC61F30C9633F983" box="[994,1084,1577,1604]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Fig. 5A</figureCitation>
); ventral tentacular cirri similar to lateral antennae, with about 10 articles (
<figureCitation id="36A676FCE671F658FDFEF36890D8F9AF" box="[637,727,1613,1640]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">Fig. 6B</figureCitation>
). Dorsal cirri longer on anterior segments, with 17-19 articles, becoming shorter at midbody (7 articles) and posterior (5 articles) segments (
<figureCitation id="36A676FCE671F658FC79F354964FF94B" box="[1018,1088,1649,1676]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5</figureCitation>
AC, 6B, E); with slight variation in size, meaning that a short one is followed by a slightly longer one, and the next one is again short and the next one slightly longer, along the body; articles relatively shorter and thicker at bases, gradually decreasing thickness towards distal end, ending in rounded tips (
<figureCitation id="36A676FCE671F658FDF6F3F890B5F93F" box="[629,698,1757,1784]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5</figureCitation>
AC, 6B, E). Thin, digitiform ventral cirri, shorter than parapodial lobes (
<figureCitation id="36A676FCE671F658FF61F2249336F8DB" box="[226,313,1793,1820]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">Fig. 6E</figureCitation>
). Parapodial lobes with two lips, anterior one rounded, posterior one digitiform, longer than anterior one (
<figureCitation id="36A676FCE671F658FF53F2009346F8F8" box="[208,329,1829,1855]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">Fig. 6B, E</figureCitation>
). Compound chaetae heterogomph falcigers, with relatively short articles except those of anteriormost chaetae (
<figureCitation id="36A676FCE671F658FEB8F26C9395F8A3" box="[315,410,1865,1892]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5D</figureCitation>
,
<figureCitation id="36A676FCE671F658FE24F26C93E4F8A4" box="[423,491,1865,1891]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">6D, F</figureCitation>
), bidentate, with proximal tooth clearly smaller and thinner than distal one, almost absent in some posterior chaetae (
<figureCitation id="36A676FCE671F658FD93F2489069F84F" box="[528,614,1901,1928]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5F</figureCitation>
,
<figureCitation id="36A676FCE671F658FDF2F2489098F84F" box="[625,663,1901,1928]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">6K</figureCitation>
). Anterior parapodia (
<figureCitation id="36A676FCE671F658FC08F248960EF840" box="[907,1025,1901,1927]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">Fig. 6B, E</figureCitation>
), with 810 compound chaetae, with slightly elongated, bidentate articles, short, straight marginal spines; dorsal ones long and relatively thick, with articles ca.
<quantity id="6965C79CE671F658FF6DF2909339F808" box="[238,310,1973,1999]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="12" pageNumber="243" unit="mm" value="44.0">44 µm</quantity>
long; progressively shorter and narrower ventrally, most ventral articles ca.
<quantity id="6965C79CE671F658FBFCF29096C8F808" box="[1151,1223,1973,1999]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="243" unit="mm" value="17.0">17 µm</quantity>
long (
<figureCitation id="36A676FCE671F658FA8FF2909766F817" box="[1292,1385,1973,2000]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="12" pageNumber="243">Figs 5D</figureCitation>
,
<figureCitation id="36A676FCE671F658FAF0F29092A5F834" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="12" pageNumber="243">6D, F</figureCitation>
). Number of chaetae per parapodium progressively decreasing towards posterior segments. Midbody parapodia with 34 compound chaetae, with blades shorter and wider than those of anterior parapodia,
<quantity id="6965C79CE672F65BFB32F5B296F4FF76" box="[1201,1275,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="246" unit="mm" value="22.0">22 µm</quantity>
long dorsally,
<quantity id="6965C79CE672F65BFF14F59E92EEFF12" box="[151,225,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="246" unit="mm" value="16.0">16 µm</quantity>
ventrally; shafts of most dorsal chaeta rather thick, appearing fused with blade, still clearly distinguishable (
<figureCitation id="36A676FCE672F65BFF1CF5FA92F6FF3E" box="[159,249,222,249]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Figs 5E</figureCitation>
,
<figureCitation id="36A676FCE672F65BFE87F5FA931DFF3D" box="[260,274,223,250]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="15" pageNumber="246">6</figureCitation>
GJ). Posterior parapodia each with three compound chaetae, similar but shorter than those on midbody, not fused either; blades
<quantity id="6965C79CE672F65BFE20F42693E3FEDA" box="[419,492,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="246" unit="mm" value="16.0">16 µm</quantity>
dorsally,
<quantity id="6965C79CE672F65BFDDAF42690ADFEDA" box="[601,674,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="246" unit="mm" value="14.0">14 µm</quantity>
ventrally (
<figureCitation id="36A676FCE672F65BFC9FF426917DFEDA" box="[796,882,258,285]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Figs 5F</figureCitation>
,
<figureCitation id="36A676FCE672F65BFCFDF42691ABFED9" box="[894,932,259,286]" captionStart="FIGURE 6" captionStartId="14.[151,250,1830,1854]" captionTargetBox="[152,1435,183,1806]" captionTargetId="figure-24@14.[151,1436,181,1806]" captionTargetPageId="14" captionText="FIGURE 6. Syllis hampirmenyatu n. sp. Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium." figureDoi="http://doi.org/10.5281/zenodo.4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="15" pageNumber="246">6K</figureCitation>
). Dorsal and ventral simple capillary chaetae not seen. Anterior parapodia with four aciculae, one with rounded tip and three with oblique tips (
<figureCitation id="36A676FCE672F65BFB4FF402972AFE86" box="[1228,1317,294,321]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Fig. 5G</figureCitation>
); midbody parapodia with two aciculae, with oblique tips (
<figureCitation id="36A676FCE672F65BFD2EF46E9107FEA2" box="[685,776,330,357]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Fig. 5H</figureCitation>
); posterior parapodia with solitary, acuminate acicula (
<figureCitation id="36A676FCE672F65BFAECF46E92BEFE4E" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Fig. 5I</figureCitation>
). Pharynx long, extending along 10 segments; pharyngeal tooth, acute, on anterior pharyngeal margin (
<figureCitation id="36A676FCE672F65BFAB6F44A979FFE4E" box="[1333,1424,366,393]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Fig. 5A</figureCitation>
). Proventricle shorter and similarly wider than pharynx, extending along approximately five segments (
<figureCitation id="36A676FCE672F65BFB7CF4B69758FE6A" box="[1279,1367,402,429]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Fig. 5A</figureCitation>
), with about 32 muscle cell rows. Pygidium short, with two anal cirri with about 10-12 articles (
<figureCitation id="36A676FCE672F65BFBFCF49296D7FE16" box="[1151,1240,438,465]" captionStart="FIGURE 5" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[219,1368,195,1868]" captionTargetId="figure-17@13.[212,1375,181,1885]" captionTargetPageId="13" captionText="FIGURE 5. Syllis hampirmenyatu n. sp. Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm." figureDoi="http://doi.org/10.5281/zenodo.4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="15" pageNumber="246">Fig. 5C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="FAE23AF1E670F659FF14F25091EFF83E" ID-DOI="http://doi.org/10.5281/zenodo.4403050" ID-Zenodo-Dep="4403050" httpUri="https://zenodo.org/record/4403050/files/figure.png" pageId="13" pageNumber="244" startId="13.[151,250,1909,1933]" targetBox="[219,1368,195,1868]" targetPageId="13">
<paragraph id="AE226A79E670F659FF14F25091EFF83E" blockId="13.[151,1437,1909,2041]" pageId="13" pageNumber="244">
<emphasis id="9CE9B66BE670F659FF14F250931AF84A" bold="true" box="[151,277,1909,1933]" pageId="13" pageNumber="244">FIGURE 5.</emphasis>
<taxonomicName id="699D11FAE670F659FE9DF25093F3F84A" authority="Martínez &amp; Martín, 2020" authorityName="Martínez &amp; Martín" authorityYear="2020" box="[286,508,1909,1933]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="244" phylum="Annelida" rank="species" species="hampirmenyatu" status="sp. nov.">
<emphasis id="9CE9B66BE670F659FE9DF25093F3F84A" box="[286,508,1909,1933]" italics="true" pageId="13" pageNumber="244">Syllis hampirmenyatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="87DA0B10E670F659FD87F2509032F84B" box="[516,573,1909,1932]" pageId="13" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
Holotype (AM W.53024), drawn from preserved specimen. A) anterior end, dorsal view. B) midbody segments, dorsal view. C) pygidium and anal cirri, dorsal view. D) compound chaetae, anterior parapodium. E) compound chaetae, midbody parapodium. F) compound chaetae, posterior parapodium. G) aciculae, anterior parapodium. H) aciculae midbody parapodium. I) acicula, posterior parapodium. Scale bars: 20µm.
</paragraph>
</caption>
<caption id="FAE23AF1E673F65AFF14F203906CF809" ID-DOI="http://doi.org/10.5281/zenodo.4403052" ID-Zenodo-Dep="4403052" httpUri="https://zenodo.org/record/4403052/files/figure.png" pageId="14" pageNumber="245" startId="14.[151,250,1830,1854]" targetBox="[152,1435,183,1806]" targetPageId="14">
<paragraph id="AE226A79E673F65AFF14F203906CF809" blockId="14.[151,1437,1830,1998]" pageId="14" pageNumber="245">
<emphasis id="9CE9B66BE673F65AFF14F2039319F8F9" bold="true" box="[151,278,1830,1854]" pageId="14" pageNumber="245">FIGURE 6.</emphasis>
<taxonomicName id="699D11FAE673F65AFE9CF20393F1F8F9" authority="Martínez &amp; Martín, 2020" authorityName="Martínez &amp; Martín" authorityYear="2020" box="[287,510,1830,1854]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="245" phylum="Annelida" rank="species" species="hampirmenyatu" status="sp. nov.">
<emphasis id="9CE9B66BE673F65AFE9CF20393F1F8F9" box="[287,510,1830,1854]" italics="true" pageId="14" pageNumber="245">Syllis hampirmenyatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="87DA0B10E673F65AFD84F203904FF8FA" box="[519,576,1830,1853]" pageId="14" pageNumber="245" rank="species">n. sp.</taxonomicNameLabel>
Paratype (AM W.45645). Scanning Electron Micrographs. A) complete specimen, anterior end (right), posterior end (left), lateral view. B) prostomium, dorsal view. Nuchal organs indicated by arrows. C) detail of nuchal organ. D) compound chaetae, anterior parapodium. E) anterior parapodia, lateral view. F) detail of compound chaeta, anterior parapodium. G) detail of compound chaeta, midbody parapodium. H), I), J) compound chaetae, midbody parapodium. K) compound chaetae, posterior parapodium.
</paragraph>
</caption>
<subSubSection id="E68739F2E672F65BFF44F4FE9773FC0E" pageId="15" pageNumber="246" type="discussion">
<paragraph id="AE226A79E672F65BFF44F4FE9773FC0E" blockId="15.[151,1437,151,1185]" pageId="15" pageNumber="246">
<emphasis id="9CE9B66BE672F65BFF44F4FE933BFE32" bold="true" box="[199,308,475,501]" pageId="15" pageNumber="246">Remarks</emphasis>
.
<taxonomicName id="699D11FAE672F65BFEC1F4FE903AFE32" authority="Martínez &amp; Martín, 2020" authorityName="Martínez &amp; Martín" authorityYear="2020" box="[322,565,475,501]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="246" phylum="Annelida" rank="species" species="hampirmenyatu" status="sp. nov.">
<emphasis id="9CE9B66BE672F65BFEC1F4FE903AFE32" box="[322,565,475,501]" italics="true" pageId="15" pageNumber="246">Syllis hampirmenyatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="87DA0B10E672F65BFDB8F4FE9077FE32" box="[571,632,475,501]" pageId="15" pageNumber="246" rank="species">n. sp.</taxonomicNameLabel>
, is characterized by having an elongated thin body, short and non-markedly thickened cirri, formed by relatively few articles and with a rounded tip, decreasing in length towards posterior segments, and posterior parapodium with three chaetae with short and wide blades appearing to be fused with shafts, but still clearly distinguishable from each other, bidentate, with proximal tooth clearly smaller and thinner than distal one. It belongs to the same group as
<taxonomicName id="699D11FAE672F65BFD2CF74E91D0FD42" authority="Grube, 1840" authorityName="Grube" authorityYear="1840" box="[687,991,619,645]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="246" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="9CE9B66BE672F65BFD2CF74E9146FD42" box="[687,841,619,645]" italics="true" pageId="15" pageNumber="246">Syllis gracilis</emphasis>
<bibRefCitation id="CA0C1788E672F65BFCD3F74E91D0FD42" author="Grube, A. E." box="[848,991,619,645]" pageId="15" pageNumber="246" refId="ref18319" refString="Grube, A. E. (1840) Actinien, Echinodermen und Wurmer des Adriatischen- und Mittelmeers nach eigenen Sammlungen bes- chrieben. Verlag J. H. Bon, Konigsberg, 92 pp. https: // doi. org / 10.5962 / bhl. title. 23025" type="book" year="1840">Grube, 1840</bibRefCitation>
</taxonomicName>
, an Atlanto-Mediterranean species having only two very thick chaetae per parapodia at midbody, with completely fused shafts and blades, as well as much thicker dorsal cirri (
<bibRefCitation id="CA0C1788E672F65BFEF0F796903AFD0A" author="San Martin, G." box="[371,565,691,717]" pageId="15" pageNumber="246" refId="ref22060" refString="San Martin, G. (2003) Annelida Polychaeta II. Syllidae. In: Ramos Sanchez, M. A., Alba Tercedor, J., Belles i Ros, X., Gosalbez i Noguera, J., Guerra Sierra, A., Macpherson Mayol, F., Martin Piera, F., Serrano Marino, J., Templado Gonzalez, J. (Eds.), Fauna Iberica. Vol. 21. Museo Nacional de Ciencias Naturales, CSIC, Madrid, pp. 1 - 554." type="book" year="2003">San Martín 2003</bibRefCitation>
).
<taxonomicName id="699D11FAE672F65BFDC4F7969178FD0A" authority="(Kinberg 1865)" baseAuthorityName="Kinberg" baseAuthorityYear="1865" box="[583,887,690,717]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="246" phylum="Annelida" rank="species" species="picta">
<emphasis id="9CE9B66BE672F65BFDC4F79690CDFD0B" box="[583,706,691,717]" italics="true" pageId="15" pageNumber="246">Syllis picta</emphasis>
(
<bibRefCitation id="CA0C1788E672F65BFD53F7969160FD0A" author="Kinberg, J. G. H." box="[720,879,690,717]" pageId="15" pageNumber="246" pagination="559 - 574" refId="ref19553" refString="Kinberg, J. G. H. (1865) Annulata nova. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, Stockholm, 21, 559 - 574." type="journal article" year="1865">Kinberg 1865</bibRefCitation>
)
</taxonomicName>
, from the Indo-Pacific, also has three chaetae per parapodia at midbody, with thickened shafts, especially the most dorsal chaeta that has also fused shaft and blade still showing the line of fusion perceptible. Moreover, these two species have very short, unidentate blades and the midbody cirri are clearly thickened (
<bibRefCitation id="CA0C1788E672F65BFDA8F6059179FCFE" author="Alvarez-Campos, P. &amp; San Martin, G. &amp; Hutchings, P." box="[555,886,798,827]" pageId="15" pageNumber="246" pagination="297 - 331" refId="ref16026" refString="Alvarez-Campos, P., San Martin, G. &amp; Hutchings, P. (2015 a) The genus Syllis Lamark, 1818 (Annelida, Syllidae) from Australia. Molecular analysis and re-description of some poorly-known species. Zootaxa, 4052 (2), 297 - 331. https: // doi. org / 10.11646 / zootaxa. 4052.3.2" type="journal article" year="2015">
Álvarez-Campos
<emphasis id="9CE9B66BE672F65BFD72F63A912EFCFE" box="[753,801,799,825]" italics="true" pageId="15" pageNumber="246">et al</emphasis>
. 2015a
</bibRefCitation>
;
<bibRefCitation id="CA0C1788E672F65BFC03F63A967BFCFE" author="Ba-Akdah, M. A. &amp; Satheesh, S. &amp; Mohsin &amp; Al-Sofyani, A. M. A. &amp; Lucas, Y. &amp; Alvarez-Campos, P. &amp; San Martin, G." box="[896,1140,799,825]" pageId="15" pageNumber="246" pagination="790 - 805" refId="ref16611" refString="Ba-Akdah, M. A., Satheesh, S., Mohsin, Al-Sofyani, A. M. A., Lucas, Y., Alvarez-Campos, P. &amp; San Martin, G. (2018) Taxonomy of some species of the genus Syllis (Annelida: Syllidae: Syllinae) from the Red Sea found among the first colonizers of an artificial substrate. Marine Biology Research, 14 (8), 790 - 805. https: // doi. org / 10.1080 / 17451000.2018.1531133" type="journal article" year="2018">
Ba-Akdah
<emphasis id="9CE9B66BE672F65BFC7AF63A9625FCFE" box="[1017,1066,799,825]" italics="true" pageId="15" pageNumber="246">et al</emphasis>
. 2018
</bibRefCitation>
).
<taxonomicName id="699D11FAE672F65BFB06F63A93E5FC9A" authority="Aguado, San Martin &amp; Ten Hove, 2008" authorityName="Aguado, San Martin &amp; Ten Hove" authorityYear="2008" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="246" phylum="Annelida" rank="species" species="ypsiloides">
<emphasis id="9CE9B66BE672F65BFB06F63A9737FCFE" box="[1157,1336,799,825]" italics="true" pageId="15" pageNumber="246">Syllis ypsiloides</emphasis>
<bibRefCitation id="CA0C1788E672F65BFABEF63A93E5FC9A" author="Aguado, M. T. &amp; San Martin, G. &amp; Ten Hove, H." pageId="15" pageNumber="246" pagination="1 - 48" refId="ref15558" refString="Aguado, M. T., San Martin, G. &amp; Ten Hove, H. (2008) Syllidae (Annelida: Polychaeta) from Indonesia collected by the Siboga (1899 - 1900) and Snellius II (1984) expeditions. Zootaxa, 1673 (1), 1 - 48. https: // doi. org / 10.11646 / zootaxa. 1673.1.1" type="journal article" year="2008">Aguado, San Martín &amp; Ten Hove, 2008</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="D68A2AE9E672F65BFDB2F6669091FC9A" box="[561,670,835,861]" name="Indonesia" pageId="15" pageNumber="246">Indonesia</collectingCountry>
and the
<collectingCountry id="D68A2AE9E672F65BFD78F6669173FC9A" box="[763,892,835,861]" name="Philippines" pageId="15" pageNumber="246">Philippines</collectingCountry>
, also has fused shaft and blade in the most dorsal midbody chaetae and the ends of blades are more pointed than the other species (
<bibRefCitation id="CA0C1788E672F65BFBAAF6429707FC46" author="Aguado, M. T. &amp; San Martin, G. &amp; Ten Hove, H." box="[1065,1288,871,897]" pageId="15" pageNumber="246" pagination="1 - 48" refId="ref15558" refString="Aguado, M. T., San Martin, G. &amp; Ten Hove, H. (2008) Syllidae (Annelida: Polychaeta) from Indonesia collected by the Siboga (1899 - 1900) and Snellius II (1984) expeditions. Zootaxa, 1673 (1), 1 - 48. https: // doi. org / 10.11646 / zootaxa. 1673.1.1" type="journal article" year="2008">
Aguado
<emphasis id="9CE9B66BE672F65BFB08F64296B3FC46" box="[1163,1212,871,897]" italics="true" pageId="15" pageNumber="246">et al</emphasis>
. 2008
</bibRefCitation>
). There seem to be numerous species with these characteristics (i.e., midbody chaetae with enlarged shaft and different degrees of blade fusion) all over the world, which have been misdentified as
<taxonomicName id="699D11FAE672F65BFCF9F68A961BFC0E" authorityName="Grube" authorityYear="1840" box="[890,1044,943,969]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="246" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="9CE9B66BE672F65BFCF9F68A961BFC0E" box="[890,1044,943,969]" italics="true" pageId="15" pageNumber="246">Syllis gracilis</emphasis>
</taxonomicName>
(
<bibRefCitation id="CA0C1788E672F65BFBA2F695977EFC0E" author="Alvarez-Campos, P. &amp; Giribet, G. &amp; Riesgo, A." box="[1057,1393,943,971]" pageId="15" pageNumber="246" pagination="138 - 150" refId="ref16168" refString="Alvarez-Campos, P., Giribet, G. &amp; Riesgo, A. (2017 a) The Syllis gracilis species complex: A molecular approach to a difficult taxonomic problem (Annelida, Syllidae). Molecular Phylogenetics and Evolution, 109, 138 - 150. https: // doi. org / 10.1016 / j. ympev. 2016.12.036" type="journal article" year="2017">
Álvarez-Campos
<emphasis id="9CE9B66BE672F65BFB6BF68A9716FC0E" box="[1256,1305,943,969]" italics="true" pageId="15" pageNumber="246">et al</emphasis>
. 2017a
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E68739F2E672F65BFF44F6F690EDFBBA" pageId="15" pageNumber="246" type="etymology">
<paragraph id="AE226A79E672F65BFF44F6F696E6FB9E" blockId="15.[151,1437,151,1185]" pageId="15" pageNumber="246">
<emphasis id="9CE9B66BE672F65BFF44F6F69347FC2A" bold="true" box="[199,328,979,1005]" pageId="15" pageNumber="246">Etymology</emphasis>
. The specific name comes from the Indonesian
<emphasis id="9CE9B66BE672F65BFCE7F6F691B4FC2B" box="[868,955,979,1004]" italics="true" pageId="15" pageNumber="246">Hampir</emphasis>
, meaning “almost” and
<emphasis id="9CE9B66BE672F65BFB4FF6F69721FC2B" box="[1228,1326,979,1004]" italics="true" pageId="15" pageNumber="246">Menyatu</emphasis>
, meaning “fused”; it refers to the degree of fusion of the dorsal-most midbody chaetae.
<collectingCountry id="D68A2AE9E672F65BFC7BF6D2967CFBD6" box="[1016,1139,1015,1041]" name="Timor-Leste" pageId="15" pageNumber="246">East Timor</collectingCountry>
is one of the two independent states that constitute the island of Timor, where Indonesian, among other languages, is spoken. Therefore, it seemed appropriate that the etymology of this new species described had a meaning in Indonesian.
</paragraph>
<paragraph id="AE226A79E672F65BFF44F14790EDFBBA" blockId="15.[151,1437,151,1185]" box="[199,738,1122,1149]" pageId="15" pageNumber="246">
<emphasis id="9CE9B66BE672F65BFF44F147932CFBBB" bold="true" box="[199,291,1122,1148]" pageId="15" pageNumber="246">Habitat</emphasis>
. Dead coral rubble, around
<quantity id="6965C79CE672F65BFDDFF146909AFBBA" box="[604,661,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="246" unit="m" value="16.0">16 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="E68739F2E672F65BFF44F1A3919FFB66" box="[199,912,1158,1185]" pageId="15" pageNumber="246" type="distribution">
<paragraph id="AE226A79E672F65BFF44F1A3919FFB66" blockId="15.[151,1437,151,1185]" box="[199,912,1158,1185]" pageId="15" pageNumber="246">
<emphasis id="9CE9B66BE672F65BFF44F1A39356FB67" bold="true" box="[199,345,1158,1184]" pageId="15" pageNumber="246">Distribution</emphasis>
. Only known from the
<typeStatus id="7126D4DBE672F65BFDE3F1A2909FFB66" box="[608,656,1159,1185]" pageId="15" pageNumber="246">type</typeStatus>
locality at
<collectingCountry id="D68A2AE9E672F65BFC8DF1A29184FB66" box="[782,907,1159,1185]" name="Timor-Leste" pageId="15" pageNumber="246">East Timor</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>