190 lines
20 KiB
XML
190 lines
20 KiB
XML
<document id="8B1059411772D713FB1811DEF5AAE32B" ID-DOI="10.1206/0003-0090(2005)290<0001:SBOTGA>2.0.CO;2" ID-ISSN="0003-0090" ID-Zenodo-Dep="5361751" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630237087708" checkinUser="felipe" docAuthor="WHEELER, QUENTIN D. & MILLER, KELLY B." docDate="2005" docId="1B4B762AFFCDFFECFF54449EB7F3F938" docLanguage="en" docName="B290.pdf" docOrigin="Bulletin of the American Museum of Natural History 2005 (290)" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0090%282005%29290%3C0001%3ASBOTGA%3E2.0.CO%3B2" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Agathidium dioperculum Wheeler and Miller 2005, new species" docType="treatment" docVersion="3" lastPageNumber="56" masterDocId="E7720E52FFF8FFDBFFCF4073B500FFF5" masterDocTitle="Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part I. Coleoptera: Leiodidae" masterLastPageNumber="95" masterPageNumber="1" pageNumber="54" updateTime="1699194440882" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="546D4879311828BE69A66667AE52FD41">Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part I. Coleoptera: Leiodidae</mods:title>
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<mods:namePart id="D95C7EABD0D71D82D208251BD5378BF9">WHEELER, QUENTIN D.</mods:namePart>
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<mods:namePart id="8F6E58D665B6B08BFCF7EFA7A50FA6CE">MILLER, KELLY B.</mods:namePart>
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<mods:typeOfResource id="F999C35DC1C4728070883CF5CFCA3B88">text</mods:typeOfResource>
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<mods:title id="067DE7292201673712B5325A54B4A95F">Bulletin of the American Museum of Natural History</mods:title>
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<mods:part id="90006098A04AB3BD3E9DBC1ECCACBB7A">
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<mods:date id="82CF8A0197930859B80A8135A527A1EF">2005</mods:date>
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<mods:number id="B63C2F9B483DFD8187976B2F83315765">2005-03-24</mods:number>
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<mods:number id="AC7F4B0B31321EA7ED5F027175DB3117">2005</mods:number>
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<mods:number id="1F33D46075B01EC4129F855E60EE1F93">290</mods:number>
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<mods:url id="A2435A3A30B1F58D309B336DFC60BA62">http://www.bioone.org/doi/abs/10.1206/0003-0090%282005%29290%3C0001%3ASBOTGA%3E2.0.CO%3B2</mods:url>
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<mods:classification id="BA47E19C70B90B2430BAFB2EBC985D3B">journal article</mods:classification>
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<mods:identifier id="87C6587D2F83E51280CD38BED64A8A8B" type="DOI">10.1206/0003-0090(2005)290<0001:SBOTGA>2.0.CO;2</mods:identifier>
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<mods:identifier id="B3D8168FBB51F00E2F1A29424491D05F" type="ISSN">0003-0090</mods:identifier>
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<treatment id="1B4B762AFFCDFFECFF54449EB7F3F938" LSID="urn:lsid:plazi:treatment:1B4B762AFFCDFFECFF54449EB7F3F938" httpUri="http://treatment.plazi.org/id/1B4B762AFFCDFFECFF54449EB7F3F938" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
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<subSubSection id="DBF894B7FFCDFFEEFF54449EB4F0FACB" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="935DC73CFFCDFFEEFF54449EB4F0FACB" blockId="53.[155,618,1261,1342]" pageId="53" pageNumber="54">
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<heading id="C8157050FFCDFFEEFF54449EB4F0FACB" centered="true" fontSize="9" level="2" pageId="53" pageNumber="54" reason="2">
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<taxonomicName id="54E2BCBFFFCDFFEEFF54449EB454FAD4" ID-CoL="8XT5P" authority="Wheeler and Miller" authorityName="Wheeler and Miller" authorityYear="2005" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dioperculum" status="sp. nov.">
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<emphasis id="A1961B2EFFCDFFEEFF54449EB4CBFAF0" bold="true" box="[155,459,1261,1285]" italics="true" pageId="53" pageNumber="54">Agathidium dioperculum</emphasis>
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Wheeler and Miller
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</taxonomicName>
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,
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<taxonomicNameLabel id="BAA5A655FFCDFFEEFEAD4579B4F1FAD4" box="[354,497,1290,1313]" pageId="53" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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<figureCitation id="0BD9DBB9FFCDFFEEFEDA4554B496FACB" box="[277,406,1319,1342]" captionStart="Fig" captionStartId="50.[141,179,889,910]" captionTargetBox="[115,1188,187,872]" captionTargetPageId="50" captionText="Fig. 82. Agathidium brevisternum group geographic distribution: A. brevisternum Fall = •; A. dioperculum, n.sp. = ·; A. rhinocerellum, n.sp. = ^." figureDoi="http://doi.org/10.5281/zenodo.5361812" httpUri="https://zenodo.org/record/5361812/files/figure.png" pageId="53" pageNumber="54">Figures 82</figureCitation>
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,
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<figureCitation id="0BD9DBB9FFCDFFEEFE6A4554B4F0FACB" box="[421,496,1319,1342]" captionStart="Figs" captionStartId="53.[725,773,917,938]" captionTargetBox="[701,1226,188,895]" captionTargetId="figure-16@53.[699,1227,187,896]" captionTargetPageId="53" captionText="Figs. 94, 95. Agathidium dioperculum, n.sp., aedeagus: 94, lateral; 95, ventral." figureDoi="http://doi.org/10.5281/zenodo.5361819" httpUri="https://zenodo.org/record/5361819/files/figure.png" pageId="53" pageNumber="54">94, 95</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DBF894B7FFCDFFEEFF574525B78CF980" pageId="53" pageNumber="54" type="materials_examined">
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<paragraph id="935DC73CFFCDFFEEFF574525B720F9CE" blockId="53.[123,652,1366,1741]" pageId="53" pageNumber="54">
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TYPE
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<collectionCode id="F5F35FF9FFCDFFEEFF104525B454FA98" box="[223,340,1366,1389]" pageId="53" pageNumber="54">MATERIAL</collectionCode>
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:
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<typeStatus id="4C59799EFFCDFFEEFEA54525B4D9FA98" box="[362,473,1366,1389]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
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, Ƌ in
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<collectionCode id="F5F35FF9FFCDFFEEFDFD4525B78BFA98" box="[562,651,1366,1389]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="53" pageNumber="54">AMNH</collectionCode>
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(deposited from
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<collectingRegion id="512609DEFFCDFFEEFE9E4507B4C7FA7E" box="[337,455,1396,1419]" country="United States of America" name="California" pageId="53" pageNumber="54">California</collectingRegion>
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Department of Food and Agriculture) labeled ‘‘
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<collectionCode id="F5F35FF9FFCDFFEEFE3045E2B75BFA5D" box="[511,603,1425,1448]" pageId="53" pageNumber="54">CALIF</collectionCode>
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:Del Norte Co. Crescent City
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<date id="E75CE1FCFFCDFFEEFE3D45DDB78BFA30" box="[498,651,1454,1477]" pageId="53" pageNumber="54" value="1978-09-19">IX19 –1978</date>
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<collectionCode id="F5F35FF9FFCDFFEEFFB445BFB58EFA16" box="[123,142,1484,1507]" name="Tavera, Department of Geology and Geophysics" pageId="53" pageNumber="54">T</collectionCode>
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.
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<collectionCode id="F5F35FF9FFCDFFEEFF5E45BFB5A6FA16" box="[145,166,1484,1507]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="53" pageNumber="54">R</collectionCode>
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.Haig,Coll. [date handwritten]/ Berlesed from Redwood Duff/
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<typeStatus id="4C59799EFFCDFFEEFE43459AB725F9F5" box="[396,549,1513,1536]" pageId="53" pageNumber="54" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="54E2BCBFFFCDFFEEFDFB459AB78AF9E8" authority="Wheeler and Miller, 2002" authorityName="Wheeler and Miller" authorityYear="2002" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dioperculum">
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<emphasis id="A1961B2EFFCDFFEEFDFB459AB452F9EB" italics="true" pageId="53" pageNumber="54">Agathidium dioperculum</emphasis>
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Wheeler and Miller, 2002
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</taxonomicName>
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[red label with black line border]’’.
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</paragraph>
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<paragraph id="935DC73CFFCDFFEEFF574632B78CF980" blockId="53.[123,652,1366,1741]" pageId="53" pageNumber="54">
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<collectionCode id="F5F35FF9FFCDFFEEFF574632B44CF9A2" box="[152,332,1601,1624]" pageId="53" pageNumber="54">TYPE LOCALITY</collectionCode>
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:
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<collectingCountry id="EBF587ACFFCDFFEEFE914632B700F9AD" box="[350,512,1601,1624]" name="United States of America" pageId="53" pageNumber="54">United States</collectingCountry>
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,
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<collectingRegion id="512609DEFFCDFFEEFDC14632B787F9AD" box="[526,647,1601,1624]" country="United States of America" name="California" pageId="53" pageNumber="54">California</collectingRegion>
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, Del Norte Co., Crescent City, redwood duff.
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</paragraph>
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</subSubSection>
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<subSubSection id="DBF894B7FFCDFFEEFF57460FB104FBBD" pageId="53" pageNumber="54" type="diagnosis">
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<paragraph id="935DC73CFFCDFFEEFF57460FB104FBBD" blockId="53.[123,652,1366,1741]" lastBlockId="53.[699,1228,1014,1741]" pageId="53" pageNumber="54">
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DIAGNOSIS: This species is separable from other
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<taxonomicName id="54E2BCBFFFCDFFEEFF7046EAB47AF945" box="[191,378,1689,1712]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="brevisternum">
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<emphasis id="A1961B2EFFCDFFEEFF7046EAB47AF945" box="[191,378,1689,1712]" italics="true" pageId="53" pageNumber="54">A. brevisternum</emphasis>
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</taxonomicName>
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group members by the male median lobe with operculum deeply di vided into two elongate plates diverging apically, with halves connected by a broad, truncate, hyaline plate (fig. 95).
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</paragraph>
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</subSubSection>
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<subSubSection id="DBF894B7FFCDFFECFD17443DB618FBF9" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="description">
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<paragraph id="935DC73CFFCDFFEEFD17443DB6B0FB2E" blockId="53.[699,1228,1014,1741]" pageId="53" pageNumber="54">
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DESCRIPTION: Body very broadly rounded, strongly convex, and partially contractile; TBL = about
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<quantity id="541A6AD9FFCDFFEEFCAB44FAB6C2FB55" box="[868,962,1161,1184]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="53" pageNumber="54" unit="mm" value="2.2">2.2 mm</quantity>
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. Color reddishbrown; antenna pale, concolorous; venter and appendages testaceous.
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</paragraph>
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<paragraph id="935DC73CFFCDFFECFD174492B426FAE0" blockId="53.[699,1228,1014,1741]" lastBlockId="55.[123,652,780,1741]" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
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Head large, very wide, broadly rounded in front, more or less quadrate and dorsoventrally flattened; OHW/MDL = 2.0; eyes very large, elongate, situated on sides of head, with large, coarse facets numbering less than
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<quantity id="541A6AD9FFCDFFEEFD744507B603FA7E" box="[699,771,1396,1419]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="53" pageNumber="54" unit="in" value="36.0">36 in</quantity>
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dorsal view; postocular temporum short, inconspicuous; OHW/PHW = 1.1; anterior marginal carina very conspicuous from edge of labrum to point posterior to eye, narrowly bordering eye; antenna with abrupt 3 segmented club; antennomere III elongate, narrow, longer than II, subequal to length of I; II:III = 1:1.8; VII, VIII subequal, small, submoniliform; VII:VIII:IX = 0.7:1:1.4; frontoclypeal suture absent, extremely faint hint of suture under high magnification; surface of head shiny, nearly impunctate; under high magnification, sparse, microscopic punctules exist bearing very fine setae; labrum narrow, elongate, distally tapered with blunt apex slightly emarginate medially. Pronotum very broad, convex; anterior margin broadly rounded; posterior angles weakly defined; surface shiny, nearly impunctate, as on head, except punctures slightly less dense; PNW/PNL = 1.8; PNL/PNH = 1.6; PNW/ PNH = 2.8. Combined elytral width, in dorsal view, about equal to length (ELW/SEL = 1.3); in posterior view, extreme apex attenuate; sutural stria evident in apical half; surface shiny, nearly impunctate, as on pronotum; epipleuron very wide, bordered by sharp carinae, ending shortly before apex of elytron. Mesosternum large, simple; without lateral carinae; without median carina; surface microscopically alutaceous; MSL/MTL = 2.1. Metasternum very short medially, sides at widest point shorter than mesosternum medially, narrowed to bands between meso and metacoxae, narrower than onehalf maximum width of metacoxa; surface with microscopic alutaceous pattern; MTL/ MTW = 0.11.
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</paragraph>
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<caption id="C79D97B4FFCEFFEDFE48458CB6CAF9E1" ID-DOI="http://doi.org/10.5281/zenodo.5361821" ID-Zenodo-Dep="5361821" box="[391,970,1535,1556]" httpUri="https://zenodo.org/record/5361821/files/figure.png" pageId="54" pageNumber="55" startId="54.[391,429,1535,1556]" targetBox="[188,1146,193,1512]" targetPageId="54">
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<paragraph id="935DC73CFFCEFFEDFE48458CB6CAF9E1" blockId="54.[391,970,1535,1556]" box="[391,970,1535,1556]" pageId="54" pageNumber="55">
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Fig. 96.
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<taxonomicName id="54E2BCBFFFCEFFEDFE3B458CB61AF9E1" authority="Fall" authorityName="Fall" box="[500,794,1535,1556]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="depressum">
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<emphasis id="A1961B2EFFCEFFEDFE3B458CB7E7F9E1" box="[500,743,1535,1556]" italics="true" pageId="54" pageNumber="55">Agathidium depressum</emphasis>
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Fall
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</taxonomicName>
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, habitus, dorsal.
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</paragraph>
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</caption>
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<caption id="C79D97B4FFCFFFECFF7642C4B18CFD39" ID-DOI="http://doi.org/10.5281/zenodo.5361823" ID-Zenodo-Dep="5361823" box="[185,1164,695,717]" httpUri="https://zenodo.org/record/5361823/files/figure.png" pageId="55" pageNumber="56" startId="55.[185,233,695,716]" targetBox="[135,1218,188,673]" targetPageId="55">
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||
<paragraph id="935DC73CFFCFFFECFF7642C4B18CFD39" blockId="55.[185,1164,695,717]" box="[185,1164,695,717]" pageId="55" pageNumber="56">
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Figs. 97, 98.
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<taxonomicName id="54E2BCBFFFCFFFECFE9542CBB783FD39" authority="Fall" authorityName="Fall" box="[346,643,695,717]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="depressum">
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<emphasis id="A1961B2EFFCFFFECFE9542CBB74DFD38" box="[346,589,696,717]" italics="true" pageId="55" pageNumber="56">Agathidium depressum</emphasis>
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Fall
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</taxonomicName>
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:
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<emphasis id="A1961B2EFFCFFFECFD4142C4B7AFFD39" bold="true" box="[654,687,695,716]" pageId="55" pageNumber="56">97,</emphasis>
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habitus, lateral;
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<emphasis id="A1961B2EFFCFFFECFCAB42C4B685FD39" bold="true" box="[868,901,695,716]" pageId="55" pageNumber="56">98,</emphasis>
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male metasternal fovea.
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</paragraph>
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</caption>
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<paragraph id="935DC73CFFCFFFECFF57456FB64FFC27" blockId="55.[123,652,780,1741]" lastBlockId="55.[699,1228,780,1741]" pageId="55" pageNumber="56">Male tarsi 5–5–4; pro and mesobasotarsomeres slightly expanded, with spatulate ventral setae; metasternal fovea median, punctiform, with few long setae; metafemur narrow, without tooth; left mandible with dorsally directed, curved, apically blunt tusk arising dorsally from side near base mandible; with tuft of apical setae. Aedeagus with short, stout median lobe, narrowed at apex in ventral view, curved ventrad with apex recurved dorsad (fig. 95); operculum with sclerotized basal and lateral areas forming deeply emarginate, divided parts, the apices of which bear two pores and the medial surfaces of which are not excavated, the distal region between two sclerotized ‘‘arms’’ connected by truncate, transparent, extremely fine, membranelike structure (fig. 95); lateral lobes gradually narrowed apically, subapical setae similar in size (figs. 94, 95) (foreshortened due to angle of vision in ventral drawing, fig. 95).</paragraph>
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<paragraph id="935DC73CFFCFFFECFD1743ABB618FBF9" blockId="55.[699,1228,780,1741]" pageId="55" pageNumber="56">Female tarsi 5–4–4; spermatheca not examined.</paragraph>
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||
</subSubSection>
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<subSubSection id="DBF894B7FFCFFFECFD174461B64AFB6B" pageId="55" pageNumber="56" type="etymology">
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<paragraph id="935DC73CFFCFFFECFD174461B64AFB6B" blockId="55.[699,1228,780,1741]" pageId="55" pageNumber="56">
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ETYMOLOGY: This species is named from
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<emphasis id="A1961B2EFFCFFFECFD744443B7D2FBB2" box="[699,722,1072,1095]" italics="true" pageId="55" pageNumber="56">di</emphasis>
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, Latin for ‘‘two’’, and
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<emphasis id="A1961B2EFFCFFFECFC144443B157FBB2" box="[987,1111,1072,1095]" italics="true" pageId="55" pageNumber="56">operculum</emphasis>
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, Latin for ‘‘lid’’, in reference to the deeply divided operculum of the median lobe in males of this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="DBF894B7FFCFFFECFD1744D7B1A2FA92" pageId="55" pageNumber="56" type="distribution">
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<paragraph id="935DC73CFFCFFFECFD1744D7B6D1FB00" blockId="55.[699,1228,780,1741]" pageId="55" pageNumber="56">
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DISTRIBUTION: This species is only known from the
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<typeStatus id="4C59799EFFCFFFECFCE444B2B65DFB2D" box="[811,861,1217,1240]" pageId="55" pageNumber="56">type</typeStatus>
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locality in extreme northwestern
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<collectingRegion id="512609DEFFCFFFECFD2544ADB660FB00" box="[746,864,1246,1269]" country="United States of America" name="California" pageId="55" pageNumber="56">California</collectingRegion>
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(fig. 82).
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</paragraph>
|
||
<paragraph id="935DC73CFFCFFFECFD1A4570B1A2FA92" blockId="55.[699,1228,780,1741]" pageId="55" pageNumber="56">
|
||
<typeStatus id="4C59799EFFCFFFECFD1A4570B646FAE2" box="[725,838,1283,1304]" pageId="55" pageNumber="56" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<collectingCountry id="EBF587ACFFCFFFECFC9A4570B11FFAED" box="[853,1055,1283,1304]" name="United States of America" pageId="55" pageNumber="56">UNITED STATES</collectingCountry>
|
||
:
|
||
<emphasis id="A1961B2EFFCFFFECFBE64570B19FFAED" box="[1065,1183,1283,1304]" italics="true" pageId="55" pageNumber="56">
|
||
<collectingRegion id="512609DEFFCFFFECFBE64570B199FAED" box="[1065,1177,1283,1304]" country="United States of America" name="California" pageId="55" pageNumber="56">California</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
Del Norte Co.: Crescent City,
|
||
<date id="E75CE1FCFFCFFFECFC1F456EB159FAC7" box="[976,1113,1309,1330]" pageId="55" pageNumber="56" value="1991-04-24">24 Apr 1991</date>
|
||
, pine duff, berlese, TR Haig (4, CASC, deposited from
|
||
<collectingRegion id="512609DEFFCFFFECFB51454BB601FA92" country="United States of America" name="California" pageId="55" pageNumber="56">California</collectingRegion>
|
||
Department of Food and Agriculture).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DBF894B7FFCFFFECFD174504B7F3F938" pageId="55" pageNumber="56" type="discussion">
|
||
<paragraph id="935DC73CFFCFFFECFD174504B7F3F938" blockId="55.[699,1228,780,1741]" pageId="55" pageNumber="56">
|
||
DISCUSSION: This form is hypothesized to be a distinct species largely because the connective structure between the two sclerotized lateral arms of the operculum was not seen in individuals of related species. Because this form appears to by sympatric with related species and because variation in some structures have been observed, including the number of pores on the operculum, it is possible that this is simply a variant of
|
||
<taxonomicName id="54E2BCBFFFCFFFECFBF9460FB7EDF945" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="brevisternum">
|
||
<emphasis id="A1961B2EFFCFFFECFBF9460FB7EDF945" italics="true" pageId="55" pageNumber="56">A. brevisternum</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="4C59799EFFCFFFECFCFE46EAB663F945" box="[817,867,1689,1712]" pageId="55" pageNumber="56">type</typeStatus>
|
||
series was collected from pine duff.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |