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<document ID-DOI="http://dx.doi.org/10.3897/alpento.7.106089" ID-Pensoft-Pub="2535-0889-7-83" ID-Pensoft-UUID="1413FD1E0A7D5CB9A8D543F2604617E5" ID-ZooBank="EC4C4A0EA1F04555B5D4A4F18AFED1AD" ModsDocID="2535-0889-7-83" checkinTime="1688785175868" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Vuataz, Laurent &amp; Gattolliat, Jean-Luc" docDate="2023" docId="17E28F33352254ED9F66685E12980967" docLanguage="en" docName="AlpEntomol 7: 83-134" docOrigin="Alpine Entomology 7" docPubDate="2023-07-07" docSource="http://dx.doi.org/10.3897/alpento.7.106089" docTitle="Labiobaetis kokoda Kaltenbach &amp; Vuataz &amp; Gattolliat 2023, sp. nov." docType="treatment" docUuid="91E672B7-A49D-4490-9D3F-6A2B92FD41EA" docUuidSource="ZooBank" docVersion="1" id="1413FD1E0A7D5CB9A8D543F2604617E5" lastPageNumber="83" masterDocId="1413FD1E0A7D5CB9A8D543F2604617E5" masterDocTitle="New species of Labiobaetis Novikova &amp; Kluge from New Guinea (Ephemeroptera, Baetidae): a never-ending story of diversity" masterLastPageNumber="134" masterPageNumber="83" pageNumber="83" updateTime="1688785175868" updateUser="pensoft">
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<mods:title id="363D321C9905596E7F7E64504B79D8BA">New species of Labiobaetis Novikova &amp; Kluge from New Guinea (Ephemeroptera, Baetidae): a never-ending story of diversity</mods:title>
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<mods:namePart id="05EC45011795CFA5324AFF3507590754">Kaltenbach, Thomas</mods:namePart>
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<mods:affiliation id="E6D0588096E8E1432906F63DE5364A0B">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier id="CDCD215CCB938248E2B160ED95691AD3" type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
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<mods:namePart id="1F8BE1617FD7DADCCCD2BF346FE054A5">Vuataz, Laurent</mods:namePart>
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<mods:affiliation id="64603DFB025462FE1E7DF0B7942E387B">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:namePart id="0155E42F32F141EC067D3C92DABF7331">Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier id="D13628956014088B699F037B31FE476E" type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation id="208BAEC7CB4A81819473CC419A3C8D16">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:title id="C2B557742DDD64DE7ED19483C27A2A37">Alpine Entomology</mods:title>
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<mods:date id="BC37D7BB4CDB39B82EDCC8D56AE22F3D">2023</mods:date>
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<subSubSection id="72B207688ED0B3F67F7DEEA95B11CF23" pageId="0" pageNumber="83" type="nomenclature">
<paragraph id="0CC51E5B2B22C0364E85CE8370BFEAC8" pageId="0" pageNumber="83">
10.
<taxonomicName id="FD3A829914A3780E89D8EC47B7037A34" LSID="https://zoobank.org/91E672B7-A49D-4490-9D3F-6A2B92FD41EA" authority="Kaltenbach &amp; Vuataz &amp; Gattolliat, 2023" authorityName="Kaltenbach &amp; Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kokoda" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kokoda" status="sp. nov.">Labiobaetis kokoda</taxonomicName>
<taxonomicNameLabel id="3D6EF94215783F499F3E3DFF2110130A" pageId="0" pageNumber="83">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CD18E1207598850807B9ED72AE81808F" pageId="0" pageNumber="83" type="description">
<paragraph id="A118688ECC2F039077E64B65BD16868E" pageId="0" pageNumber="83">
<figureCitation id="CB25B2BC1330DCFCCABEADFEA1B651F7" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis kokoda sp. nov., larva habitus: a. Dorsal view; b. Ventral view. Scale bars 1 mm." figureDoi="10.3897/alpento.7.106089.figure32" httpUri="https://binary.pensoft.net/fig/874320" pageId="0" pageNumber="83">Figs 32</figureCitation>
<figureCitation id="6E1DBE6B3962C4DD3F2B00DA07B50A38" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Labiobaetis kokoda sp. nov., larva morphology: a. Labrum; b, c. Sections of labrum, dorsal focus; d, e. Right mandible; f, g. Left mandible. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83">, 33</figureCitation>
<figureCitation id="4F0E2446973434D8E3E40745551F0111" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Labiobaetis kokoda sp. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Maxillary palp; d. Section of maxilla. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure34" httpUri="https://binary.pensoft.net/fig/874322" pageId="0" pageNumber="83">, 34</figureCitation>
<figureCitation id="B09207E17785DE62DB13EFCF6AE1E9FC" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Labiobaetis kokoda sp. nov., larva morphology: a. Labium; b. Glossa, paraglossa and mentum, ventral focus; c. Section of paraglossa, dorsal focus; d. Labial palp, ventral focus; e. Section of labial palp, dorsal focus. Scale bar: 50 µm." figureDoi="10.3897/alpento.7.106089.figure35" httpUri="https://binary.pensoft.net/fig/874323" pageId="0" pageNumber="83">, 35</figureCitation>
<figureCitation id="605A8E0A4348CA6E2FC04FA7ABEB8D3C" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis kokoda sp. nov., larva morphology: a. Fore leg; b. Fore femur, dorsal margin; c. Fore femur, ventral margin; d. Fore femur, setae on distomedial surface; e. Fore tibia, dorsal margin; f. Fore tibia, ventral margin; g. Fore tarsus, dorsal margin; h. Fore tarsus, ventral margin; i. Fore claw; j. Fore femur, apex, posterior side. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure36" httpUri="https://binary.pensoft.net/fig/874324" pageId="0" pageNumber="83">, 36</figureCitation>
<figureCitation id="1EE3EB65EE2843DB9A891165821E5391" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis kokoda sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Costal margin of tergalius IV; f. Anal margin of tergalius IV; g. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83">, 37</figureCitation>
<figureCitation id="7D5519E82BB4FC36828BA5E792C51A46" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution of species treated in this study: a. Species of group Labiobaetis petersorum and L. kokoda sp. nov.; b. MOTUs of group Labiobaetis petersorum, not described in this study." figureDoi="10.3897/alpento.7.106089.figure38" httpUri="https://binary.pensoft.net/fig/874326" pageId="0" pageNumber="83">, 38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6DABC27B11B04230C7D04F3BCD48028F" pageId="0" pageNumber="83" type="diagnosis">
<paragraph id="55AD87561A40F25E7CD9B0BF70EF947E" pageId="0" pageNumber="83">Diagnosis.</paragraph>
<paragraph id="515C22D94EEF841315094AD720DF9AF3" pageId="0" pageNumber="83">
<emphasis id="75C2BDE9C3D63581B6112BEA1FB1935D" bold="true" pageId="0" pageNumber="83">Larva.</emphasis>
Following combination of characters differentiates
<taxonomicName id="9BEE5B92E1BB8A20AD1EF810DBAA7A48" lsidName="L. kokoda" pageId="0" pageNumber="83" rank="species" species="kokoda">
<emphasis id="516E782B706DE3704AA7EB8DD643692F" italics="true" pageId="0" pageNumber="83">L. kokoda</emphasis>
</taxonomicName>
sp. nov. from other species of
<taxonomicName id="1BE7F33C0E80D43A6E481B1623AAA84C" authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="4D6221547CDD839FDBA6ACBEA7A6E8D2" italics="true" pageId="0" pageNumber="83">Labiobaetis</emphasis>
</taxonomicName>
: A) labrum length 0.7
<normalizedToken id="C91120668A37178E7E6392934958C251" originalValue="×">x</normalizedToken>
maximal width; dorsal submarginal arc of setae consisting of one plus 3-5 simple setae, 1st and 2nd setae after submedian seta closely together (Fig.
<figureCitation id="AC65E119901CFD65BE1F48F8140F8178" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Labiobaetis kokoda sp. nov., larva morphology: a. Labrum; b, c. Sections of labrum, dorsal focus; d, e. Right mandible; f, g. Left mandible. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83">33a-c</figureCitation>
); B) incisor and kinetodontium of right mandible with four and three denticles; margin between prostheca and mola slightly convex, smooth (Fig.
<figureCitation id="B5F9CA0B6DAF6158E1C864131F6D76B8" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Labiobaetis kokoda sp. nov., larva morphology: a. Labrum; b, c. Sections of labrum, dorsal focus; d, e. Right mandible; f, g. Left mandible. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83">33d, e</figureCitation>
); C) incisor and kinetodontium of left mandible with three and three denticles; margin between prostheca and mola almost straight, smooth (Fig.
<figureCitation id="1117EC326781883B348CFA6F2249AADB" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Labiobaetis kokoda sp. nov., larva morphology: a. Labrum; b, c. Sections of labrum, dorsal focus; d, e. Right mandible; f, g. Left mandible. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83">33f, g</figureCitation>
); D) hypopharynx with well-developed medial tuft of stout setae (Fig.
<figureCitation id="B5328623A4C16C64A279A36FB87B0F71" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Labiobaetis kokoda sp. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Maxillary palp; d. Section of maxilla. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure34" httpUri="https://binary.pensoft.net/fig/874322" pageId="0" pageNumber="83">34a</figureCitation>
); E) maxillary palp longer than galea-lacinia; segment II without distolateral excavation, apically pointed, constricted (Fig.
<figureCitation id="B6833FB464231854E243BD978FBD393C" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Labiobaetis kokoda sp. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Maxillary palp; d. Section of maxilla. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure34" httpUri="https://binary.pensoft.net/fig/874322" pageId="0" pageNumber="83">34b, c</figureCitation>
); F) labial palp segment II with thumb-like, distomedial protuberance; segment III oblong (Fig.
<figureCitation id="2205219EE0E67BEED78A8A468BA4D5A4" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Labiobaetis kokoda sp. nov., larva morphology: a. Labium; b. Glossa, paraglossa and mentum, ventral focus; c. Section of paraglossa, dorsal focus; d. Labial palp, ventral focus; e. Section of labial palp, dorsal focus. Scale bar: 50 µm." figureDoi="10.3897/alpento.7.106089.figure35" httpUri="https://binary.pensoft.net/fig/874323" pageId="0" pageNumber="83">35a, d</figureCitation>
); G) femur dorsally with row of ca. 12 long, spine-like setae on margin; several such setae additionally in partial 2nd row near margin (Fig.
<figureCitation id="A55ED074D5D82F999F67FB40E0CE4161" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis kokoda sp. nov., larva morphology: a. Fore leg; b. Fore femur, dorsal margin; c. Fore femur, ventral margin; d. Fore femur, setae on distomedial surface; e. Fore tibia, dorsal margin; f. Fore tibia, ventral margin; g. Fore tarsus, dorsal margin; h. Fore tarsus, ventral margin; i. Fore claw; j. Fore femur, apex, posterior side. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure36" httpUri="https://binary.pensoft.net/fig/874324" pageId="0" pageNumber="83">36a, b</figureCitation>
); H) claw with ca. 14 denticles (Fig.
<figureCitation id="B35C32B70B585B2510FECC6F2268B9C0" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis kokoda sp. nov., larva morphology: a. Fore leg; b. Fore femur, dorsal margin; c. Fore femur, ventral margin; d. Fore femur, setae on distomedial surface; e. Fore tibia, dorsal margin; f. Fore tibia, ventral margin; g. Fore tarsus, dorsal margin; h. Fore tarsus, ventral margin; i. Fore claw; j. Fore femur, apex, posterior side. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure36" httpUri="https://binary.pensoft.net/fig/874324" pageId="0" pageNumber="83">36i</figureCitation>
); I) posterior margin of abdominal tergum IV with triangular or rounded spines, wider than long (Fig.
<figureCitation id="79F919183B9517912422A792BF6D975A" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis kokoda sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Costal margin of tergalius IV; f. Anal margin of tergalius IV; g. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83">37c</figureCitation>
); J) antennal scape without distolateral process (Fig.
<figureCitation id="EB259723FE99AC0020745AB4F7425E18" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis kokoda sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Costal margin of tergalius IV; f. Anal margin of tergalius IV; g. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83">37a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4DB079F55DCEF786AF779DA0DF8D33DC" pageId="0" pageNumber="83" type="description">
<paragraph id="4822B1AA250F32A0EC7B7DBFAF6E7C2E" pageId="0" pageNumber="83">Description.</paragraph>
<paragraph id="1B5C91A80580F47155654AF32678ECB8" pageId="0" pageNumber="83">
<emphasis id="CF3E82FB24B411190FB4CB8190106B8B" bold="true" pageId="0" pageNumber="83">Larva</emphasis>
(Figs
<figureCitation id="67CA235DC0D977CE4D459006960C6D5A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis kokoda sp. nov., larva habitus: a. Dorsal view; b. Ventral view. Scale bars 1 mm." figureDoi="10.3897/alpento.7.106089.figure32" httpUri="https://binary.pensoft.net/fig/874320" pageId="0" pageNumber="83">32</figureCitation>
-
<figureCitation id="D565D2A928F7B07EBCB57C07CFC35733" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis kokoda sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Costal margin of tergalius IV; f. Anal margin of tergalius IV; g. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83">37</figureCitation>
). Body length ca. 3 mm (immature). Cerci: broken. Paracercus ca. 0.4
<normalizedToken id="2775C42ACA202FA3EB6EBBE52CE1E9B9" originalValue="×">x</normalizedToken>
body length. Antenna ca. 2.5
<normalizedToken id="31A8A302DD77034044461EB40C8C9B5F" originalValue="×">x</normalizedToken>
head length.
</paragraph>
<caption id="32E6DD649BBB544A9FADE2C7A588EEA6" doi="10.3897/alpento.7.106089.figure32" httpUri="https://binary.pensoft.net/fig/874320" pageId="0" pageNumber="83" start="Figure 32" startId="F32">
<paragraph id="373B5FC96683FA2DD96EFAB5BCDA9C69" pageId="0" pageNumber="83">
<emphasis id="98DFA5DC489EEEAFA11EBFAAAC25DB6F" bold="true" pageId="0" pageNumber="83">Figure 32.</emphasis>
<taxonomicName id="515B1026484DAC3F95A717AEB4CFCDA2" authorityName="Kaltenbach &amp; Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kokoda" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kokoda">
<emphasis id="49513D625E79951BB6D0115D361F9F78" italics="true" pageId="0" pageNumber="83">Labiobaetis kokoda</emphasis>
</taxonomicName>
sp. nov., larva habitus:
<emphasis id="42522B874F98559BEE5E1DE6AC1F778D" bold="true" pageId="0" pageNumber="83">a.</emphasis>
Dorsal view;
<emphasis id="C4E336FC5220E1B972D7CC7AB97DA2B8" bold="true" pageId="0" pageNumber="83">b.</emphasis>
Ventral view. Scale bars 1 mm.
</paragraph>
</caption>
<caption id="B956612276F0938F10CE12FEEDE9B834" doi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83" start="Figure 33" startId="F33">
<paragraph id="42770050AF202B5EF3C1F6E8552872CE" pageId="0" pageNumber="83">
<emphasis id="1403EFE04435E30B7773C587BFA8AEA4" bold="true" pageId="0" pageNumber="83">Figure 33.</emphasis>
<taxonomicName id="1D15F9624A50BE5D12C964358AB2692D" authorityName="Kaltenbach &amp; Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kokoda" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kokoda">
<emphasis id="B5900E66CA5595A2EF7406652690751A" italics="true" pageId="0" pageNumber="83">Labiobaetis kokoda</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis id="81E801A560DAEC4E42F22B55B8DB9649" bold="true" pageId="0" pageNumber="83">a.</emphasis>
Labrum;
<emphasis id="C0940A37DC9485E1CE868E34A06DE74F" bold="true" pageId="0" pageNumber="83">b, c.</emphasis>
Sections of labrum, dorsal focus;
<emphasis id="FB931677130C8669B4279BB7AC1E1089" bold="true" pageId="0" pageNumber="83">d, e.</emphasis>
Right mandible;
<emphasis id="5E9288DAEF47D1FB3473254ECB98D129" bold="true" pageId="0" pageNumber="83">f, g.</emphasis>
Left mandible. Scale bars: 50
<normalizedToken id="C3C47DC10C6F02AF7107F7ECBA1550A3" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption id="BABE435D15C4E09359ED7F3A688D683A" doi="10.3897/alpento.7.106089.figure34" httpUri="https://binary.pensoft.net/fig/874322" pageId="0" pageNumber="83" start="Figure 34" startId="F34">
<paragraph id="3668A56A62BAEDB992190CB2380215D2" pageId="0" pageNumber="83">
<emphasis id="F5D4474BF26129358B747C3BBE77DB35" bold="true" pageId="0" pageNumber="83">Figure 34.</emphasis>
<taxonomicName id="81BB9D20D69974DFB453975499110C49" authorityName="Kaltenbach &amp; Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kokoda" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kokoda">
<emphasis id="832175BC3E20A6F5F3B028E7AB9D14B1" italics="true" pageId="0" pageNumber="83">Labiobaetis kokoda</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis id="08A91267B6C25105F63A04D4422EE7E4" bold="true" pageId="0" pageNumber="83">a.</emphasis>
Hypopharynx and superlinguae;
<emphasis id="A9E9ED380237D41A0A02C0691A28EB44" bold="true" pageId="0" pageNumber="83">b.</emphasis>
Maxilla;
<emphasis id="A7945ACBA4C7BA629A4C2F89CEA22D7D" bold="true" pageId="0" pageNumber="83">c.</emphasis>
Maxillary palp;
<emphasis id="CB792F393F586D75335BE1E649A200E7" bold="true" pageId="0" pageNumber="83">d.</emphasis>
Section of maxilla. Scale bars: 50
<normalizedToken id="560D6F3DBB189449D0C2BDDC1007EC03" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption id="028A3A70D8F7AE78E3CF2F62D7165253" doi="10.3897/alpento.7.106089.figure35" httpUri="https://binary.pensoft.net/fig/874323" pageId="0" pageNumber="83" start="Figure 35" startId="F35">
<paragraph id="D9ED1F8F8A3CB63744BCD3A1C64F5505" pageId="0" pageNumber="83">
<emphasis id="107E85C1A708FCDB009B50D4A97B1B8D" bold="true" pageId="0" pageNumber="83">Figure 35.</emphasis>
<taxonomicName id="298B723A88079C1579CBE692109B9457" authorityName="Kaltenbach &amp; Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kokoda" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kokoda">
<emphasis id="99DB30E3A79C7E9C915DA5A89B7C378C" italics="true" pageId="0" pageNumber="83">Labiobaetis kokoda</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis id="9604E18E5B4147327E5A4E00BBFBE385" bold="true" pageId="0" pageNumber="83">a.</emphasis>
Labium;
<emphasis id="9DF015C2984401C77AFEBE5137A0ADD5" bold="true" pageId="0" pageNumber="83">b.</emphasis>
Glossa, paraglossa and mentum, ventral focus;
<emphasis id="23807780AF867F7D7ABC1E8592FBBDB0" bold="true" pageId="0" pageNumber="83">c.</emphasis>
Section of paraglossa, dorsal focus;
<emphasis id="D79D1952B6963165037F98333705C1B5" bold="true" pageId="0" pageNumber="83">d.</emphasis>
Labial palp, ventral focus;
<emphasis id="E41FBDF04F3F6DC79A48AEB1F87FDAE9" bold="true" pageId="0" pageNumber="83">e.</emphasis>
Section of labial palp, dorsal focus. Scale bar: 50
<normalizedToken id="FB966D6E05113724EC73418E4CA017A1" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption id="0C9501F79AC8D9F71A79B812F5BCA9CE" doi="10.3897/alpento.7.106089.figure36" httpUri="https://binary.pensoft.net/fig/874324" pageId="0" pageNumber="83" start="Figure 36" startId="F36">
<paragraph id="5E4C3AE77F058F2822FDA09205447FEC" pageId="0" pageNumber="83">
<emphasis id="4DAC886ACD123A35EA292C5D2F1AD796" bold="true" pageId="0" pageNumber="83">Figure 36.</emphasis>
<taxonomicName id="808173D1688698E45C954AC2C526AB34" authorityName="Kaltenbach &amp; Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kokoda" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kokoda">
<emphasis id="C37422F24BD3FBB86CE1C8D5F5325A99" italics="true" pageId="0" pageNumber="83">Labiobaetis kokoda</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis id="F91EA776A385150E8978B744922821B2" bold="true" pageId="0" pageNumber="83">a.</emphasis>
Fore leg;
<emphasis id="5817BD21BE78398D2EC541B0EBFFE32E" bold="true" pageId="0" pageNumber="83">b.</emphasis>
Fore femur, dorsal margin;
<emphasis id="B61E7D5B577DFA4069C0C015DF139F21" bold="true" pageId="0" pageNumber="83">c.</emphasis>
Fore femur, ventral margin;
<emphasis id="AD1B630602F287614F230989C1357767" bold="true" pageId="0" pageNumber="83">d.</emphasis>
Fore femur, setae on distomedial surface;
<emphasis id="7BB923279327A55803CE17FD0981D20E" bold="true" pageId="0" pageNumber="83">e.</emphasis>
Fore tibia, dorsal margin;
<emphasis id="262CD3F6C63EFFBB39D63E35180723F9" bold="true" pageId="0" pageNumber="83">f.</emphasis>
Fore tibia, ventral margin;
<emphasis id="2CF6C08DE2D4E5A1203E3A270DB21B66" bold="true" pageId="0" pageNumber="83">g.</emphasis>
Fore tarsus, dorsal margin;
<emphasis id="2AF5678B84EB1FB1B789FCA74C024645" bold="true" pageId="0" pageNumber="83">h.</emphasis>
Fore tarsus, ventral margin;
<emphasis id="E016E23BCAD78182B5DD7294C68C15B8" bold="true" pageId="0" pageNumber="83">i.</emphasis>
Fore claw;
<emphasis id="AA6858FB882EB123256B36DFEA554D10" bold="true" pageId="0" pageNumber="83">j.</emphasis>
Fore femur, apex, posterior side. Scale bars: 50
<normalizedToken id="112EBDEA4DB6020EE05B6C68D564E4D1" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption id="249196694611E0F61CD047C9A37841E6" doi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83" start="Figure 37" startId="F37">
<paragraph id="338A38F1230B60EEB042D826816E8026" pageId="0" pageNumber="83">
<emphasis id="06EC8646A0007EB0D6B11B9A75EB096A" bold="true" pageId="0" pageNumber="83">Figure 37.</emphasis>
<taxonomicName id="A0F88C233AC5FD896C3E507AE26B27F1" authorityName="Kaltenbach &amp; Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis kokoda" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kokoda">
<emphasis id="14F0406C7FD6A2AE7F91869B393FA102" italics="true" pageId="0" pageNumber="83">Labiobaetis kokoda</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis id="D9530F25953F259A8CD3B0611BC0AA2B" bold="true" pageId="0" pageNumber="83">a.</emphasis>
Base of antenna;
<emphasis id="AEB062A551DA12B535CDBE02B532F91A" bold="true" pageId="0" pageNumber="83">b.</emphasis>
Antenna;
<emphasis id="860073A1F6A5ED9C8BD38189C2C531B6" bold="true" pageId="0" pageNumber="83">c.</emphasis>
Abdominal tergum IV;
<emphasis id="7394ADFE7E046DE6D20C6B99947E1312" bold="true" pageId="0" pageNumber="83">d.</emphasis>
Tergalius IV;
<emphasis id="E28264AADF29898E894C3FDB4DD80470" bold="true" pageId="0" pageNumber="83">e.</emphasis>
Costal margin of tergalius IV;
<emphasis id="F8A5692D9DD028EE0F062C082952ECB5" bold="true" pageId="0" pageNumber="83">f.</emphasis>
Anal margin of tergalius IV;
<emphasis id="F6EF8CC474686412E8489CE6CFF44F5D" bold="true" pageId="0" pageNumber="83">g.</emphasis>
Paraproct. Scale bars: 50
<normalizedToken id="3AFD404535A4CD934F70DB1EDE300ADB" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id="9F987CA86C523EFCA755915D52B9242A" pageId="0" pageNumber="83">
<emphasis id="A7BE5231AE5333B6DAB8470F6DEE8BF9" bold="true" italics="true" pageId="0" pageNumber="83">Cuticular colouration</emphasis>
(Fig.
<figureCitation id="3FBDF3E7848527DA6B360517CF40EA2E" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis kokoda sp. nov., larva habitus: a. Dorsal view; b. Ventral view. Scale bars 1 mm." figureDoi="10.3897/alpento.7.106089.figure32" httpUri="https://binary.pensoft.net/fig/874320" pageId="0" pageNumber="83">32a, b</figureCitation>
). Antenna light brown, darker at distal margins of segments. Head, thorax and abdomen dorsally brown, with pattern as in Fig.
<figureCitation id="7CE242D6C603CFE8458D34F550949386" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis kokoda sp. nov., larva habitus: a. Dorsal view; b. Ventral view. Scale bars 1 mm." figureDoi="10.3897/alpento.7.106089.figure32" httpUri="https://binary.pensoft.net/fig/874320" pageId="0" pageNumber="83">32a</figureCitation>
; abdominal terga I, V and IX-X brighter. Head, thorax and abdomen ventrally light brown; abdominal sterna VI-VIII darker, IX-X brighter. Legs light brown, darker along dorsal margins of tibia and tarsus, femur medially and apically darker. Caudalii light brown.
</paragraph>
<paragraph id="A28BFEAB61BC4ADEB386F90D933270EA" pageId="0" pageNumber="83">
<emphasis id="25D350D7811667B5FA2D5B1F31BC4A1C" bold="true" italics="true" pageId="0" pageNumber="83">Antenna</emphasis>
(Fig.
<figureCitation id="78191FAE2428CCB19E5916173E41676A" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis kokoda sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Costal margin of tergalius IV; f. Anal margin of tergalius IV; g. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83">37a</figureCitation>
) with scape and pedicel sub-cylindrical, without distolateral process at scape. Pedicel distally with triangular scales.
</paragraph>
<paragraph id="A5A317CF83E27889A6EBE193BCFEB01E" pageId="0" pageNumber="83">
<emphasis id="3D49A1FAD28A4384BC9725F75DF20891" bold="true" italics="true" pageId="0" pageNumber="83">Labrum</emphasis>
(Fig.
<figureCitation id="B0F6FD6DCE2F83B02E5756F01B0E40A6" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Labiobaetis kokoda sp. nov., larva morphology: a. Labrum; b, c. Sections of labrum, dorsal focus; d, e. Right mandible; f, g. Left mandible. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83">33a-c</figureCitation>
). Sub-rectangular, length 0.7
<normalizedToken id="EB8A781AFC8ED0CED49FD27B28F5F083" originalValue="×">x</normalizedToken>
maximum width. Distal margin with deep medial emargination and small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 3-5 long, simple setae, 1st and 2nd seta after submedian seta closely together. Ventrally with marginal row of setae composed of anterolateral long, feathered setae and medial long, bifid setae.
</paragraph>
<paragraph id="94E79024B1063F11ADF539BD9B3C70DF" pageId="0" pageNumber="83">
<emphasis id="80D880B83D2ED099094847A9479943BD" bold="true" italics="true" pageId="0" pageNumber="83">Right mandible</emphasis>
(Fig.
<figureCitation id="6EFA2D2E8AE1AF6ECAF3CA93D488A80C" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Labiobaetis kokoda sp. nov., larva morphology: a. Labrum; b, c. Sections of labrum, dorsal focus; d, e. Right mandible; f, g. Left mandible. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83">33d, e</figureCitation>
). Incisor and kinetodontium fused. Incisor with four denticles; kinetodontium with three denticles, inner margin of innermost denticle with row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex, smooth. Tuft of setae at apex of mola present.
</paragraph>
<paragraph id="FE522A0DC66F64B91D9AD9AA2FC2C7A1" pageId="0" pageNumber="83">
<emphasis id="1EF4291795AB16D6DB9387311BDB6A13" bold="true" italics="true" pageId="0" pageNumber="83">Left mandible</emphasis>
(Fig.
<figureCitation id="D71BAFB71A64357EC139C05B1E9BA0ED" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Labiobaetis kokoda sp. nov., larva morphology: a. Labrum; b, c. Sections of labrum, dorsal focus; d, e. Right mandible; f, g. Left mandible. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure33" httpUri="https://binary.pensoft.net/fig/874321" pageId="0" pageNumber="83">33f, g</figureCitation>
). Incisor and kinetodontium fused. Incisor with three denticles; kinetodontium with three denticles. Prostheca robust, apicolaterally with small denticles and comb-shaped structure. Margin between prostheca and mola almost straight, smooth. Subtriangular process above level of area between prostheca and mola. Tuft of setae at apex of mola present.
</paragraph>
<paragraph id="13E84FCEDAAAF3CD0DCA472BC8B95DBF" pageId="0" pageNumber="83">Both mandibles with lateral margins almost straight.</paragraph>
<paragraph id="3EC2540E0E672C729C30646A77B20F9F" pageId="0" pageNumber="83">
<emphasis id="D5B55557894C8024B10FB407BDACB3F1" bold="true" italics="true" pageId="0" pageNumber="83">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation id="86ADB89B7ECD4FBAB1866167D3FD7E2F" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Labiobaetis kokoda sp. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Maxillary palp; d. Section of maxilla. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure34" httpUri="https://binary.pensoft.net/fig/874322" pageId="0" pageNumber="83">34a</figureCitation>
). Lingua slightly longer than superlinguae. Lingua longer than broad; medial tuft of stout setae well developed, distolaterally with two additional tufts of setae; distal half laterally slightly expanded. Superlinguae distolaterally slightly protruding; lateral margins rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph id="8747B3CA9F0C1454A993CEFB858C1805" pageId="0" pageNumber="83">
<emphasis id="B33E185BF3C57FF1C168822FB74473DF" bold="true" italics="true" pageId="0" pageNumber="83">Maxilla</emphasis>
(Fig.
<figureCitation id="C2925A9EC8BADF281FEC29890B7F0741" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Labiobaetis kokoda sp. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Maxillary palp; d. Section of maxilla. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure34" httpUri="https://binary.pensoft.net/fig/874322" pageId="0" pageNumber="83">34b-d</figureCitation>
). Galea-lacinia ventrally with two simple, apical setae below canines. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one feathered spine-like seta and ca. six long, simple setae. Maxillary palp ca. 1.1
<normalizedToken id="9324EFDB7496DA24CDF76D9AD583E0FA" originalValue="×">x</normalizedToken>
length of galea-lacinia; 2-segmented; palp segment II subequal in length to segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment without distolateral excavation, apically slightly pointed, constricted.
</paragraph>
<paragraph id="279E5CFC5505D16E713A9A4193EB6574" pageId="0" pageNumber="83">
<emphasis id="4ABC3379D470F32FD8C395B8164FAA80" bold="true" italics="true" pageId="0" pageNumber="83">Labium</emphasis>
(Fig.
<figureCitation id="0B6C2A06A65ECB140E4D286C7AA6C159" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Labiobaetis kokoda sp. nov., larva morphology: a. Labium; b. Glossa, paraglossa and mentum, ventral focus; c. Section of paraglossa, dorsal focus; d. Labial palp, ventral focus; e. Section of labial palp, dorsal focus. Scale bar: 50 µm." figureDoi="10.3897/alpento.7.106089.figure35" httpUri="https://binary.pensoft.net/fig/874323" pageId="0" pageNumber="83">35a-e</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca. six spine-like seta; apex with two long and one medium robust, apically pectinate setae; outer margin with ca. six spine-like setae; ventral surface with fine, simple, scattered setae. Paraglossa sub-rectangular, slightly curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area; ventrally ca. four medium, fine, simple setae in anteromedial area; dorsally with five long, spine-like setae near inner margin. Labial palp with segment I approx. as long as length of segments II and III combined. Segment II with thumb-like, distomedial protuberance; distomedial protuberance 0.4
<normalizedToken id="C2C6E48F52C6964F0E14708302F36A3E" originalValue="×">x</normalizedToken>
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with row of ca. four spine-like setae near outer margin. Segment III oblong; length ca. 1.1
<normalizedToken id="020F2A35D6568945785DDD4DD6AEE81E" originalValue="×">x</normalizedToken>
width; ventrally covered with short to medium, spine-like, simple setae and short, fine, simple setae. Mentum ventrally with medium fine, simple setae scattered on distal part of surface.
</paragraph>
<paragraph id="E0269D220D042F129A052E3AA50B8CA7" pageId="0" pageNumber="83">
<emphasis id="381F10940E481940879E518267C58C56" bold="true" italics="true" pageId="0" pageNumber="83">Hind protoptera</emphasis>
absent.
</paragraph>
<paragraph id="FAE0250AF9F652EBFB3CF13EDCB49AA3" pageId="0" pageNumber="83">
<emphasis id="8CB6BDCF1582BE55BF80F56A77810DDA" bold="true" italics="true" pageId="0" pageNumber="83">Legs</emphasis>
(Fig.
<figureCitation id="3322F6DB66702F16ACE3AC720487A803" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis kokoda sp. nov., larva morphology: a. Fore leg; b. Fore femur, dorsal margin; c. Fore femur, ventral margin; d. Fore femur, setae on distomedial surface; e. Fore tibia, dorsal margin; f. Fore tibia, ventral margin; g. Fore tarsus, dorsal margin; h. Fore tarsus, ventral margin; i. Fore claw; j. Fore femur, apex, posterior side. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure36" httpUri="https://binary.pensoft.net/fig/874324" pageId="0" pageNumber="83">36a-j</figureCitation>
). Ratio of foreleg segments 1.5:1.0:0.8:0.3.
<emphasis id="DF5A2CFAFF5135A04AA24132D42447D6" bold="true" pageId="0" pageNumber="83">
<emphasis id="AFCE34F83A41885E3E3A07CD87DE92C8" italics="true" pageId="0" pageNumber="83">Femur</emphasis>
.
</emphasis>
Femur very wide, length ca. 2.3
<normalizedToken id="D3D3DF6676CA75D8CCDA45395715AC66" originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with row of ca. 12 long, curved, spine-like setae, and some additional spine-like setae in partial 2nd row near margin; length of setae 0.2
<normalizedToken id="6961D66B872CF233F0C6D2686A4609DA" originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded, with a pair of long, spine-like setae and several medium, spine-like setae. Apex on posterior side with few short, stout, setae on fore and middle leg, absent on hind leg. Stout, lanceolate, pointed setae scattered along ventral margin; several of such setae on distomedial surface. Femoral patch rudimentary on fore and middle legs, reduced on hind leg.
<emphasis id="C999F084204780C56846043933C8AE8E" bold="true" pageId="0" pageNumber="83">
<emphasis id="7BB406C29FBC2D2921318CBF56841EF8" italics="true" pageId="0" pageNumber="83">Tibia</emphasis>
.
</emphasis>
Dorsal margin with row of short, spine-like, pointed setae. Ventral margin with row of short, curved, spine-like setae, on apex a tuft of fine, simple setae. Anterior surface with short, stout, lanceolate, pointed setae along patellatibial suture. Posterior surface without stout setae. Patellatibial suture present on basal 2/3 area.
<emphasis id="7BFE2C467A0EAFCDD2D03C237B11C9E6" bold="true" pageId="0" pageNumber="83">
<emphasis id="7ECCC23AB8CEF5C9911892BA8B4EB1A0" italics="true" pageId="0" pageNumber="83">Tarsus</emphasis>
.
</emphasis>
Dorsal margin with few short, stout setae. Ventral margin with row of curved, spine-like setae increasing in length distally. Claw with one row of ca. 14 denticles; distally pointed; without subapical seta on.
</paragraph>
<paragraph id="8A331AFA7B81C630B439EF4BFADFD189" pageId="0" pageNumber="83">
<emphasis id="9A8171A7DDE91B01AEC2FBED84514E68" bold="true" italics="true" pageId="0" pageNumber="83">Abdominal terga</emphasis>
(Fig.
<figureCitation id="EF1BF92A9E066AFDBFA27DD1E340B503" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis kokoda sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Costal margin of tergalius IV; f. Anal margin of tergalius IV; g. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83">37c</figureCitation>
). Surface with irregular rows of U-shaped scale bases. Posterior margin of terga: I smooth, without spines; II with rounded spines, much wider than long; III-VI with triangular or rounded spines, wider than long; VII-IX with triangular spines, longer than wide.
</paragraph>
<paragraph id="3B9BDCC244908C8FE81DBE38403CDA5E" pageId="0" pageNumber="83">
<emphasis id="A53E0A040C58E51AAD245EA591A6FEBA" bold="true" pageId="0" pageNumber="83">
<emphasis id="5890E78619DCD8BA883AFAC467243310" italics="true" pageId="0" pageNumber="83">Abdominal sterna</emphasis>
.
</emphasis>
Posterior margin of sterna: I-VI smooth, without spines; VII-IX with small, triangular spines.
</paragraph>
<paragraph id="4464B8970C9219855D5ACCE28A03E8FE" pageId="0" pageNumber="83">
<emphasis id="841085D64714EB15E204FC409BBED782" bold="true" italics="true" pageId="0" pageNumber="83">Tergalii</emphasis>
(Fig.
<figureCitation id="73DA32520BA089C4C1BFBF7E27FFD2E1" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis kokoda sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Costal margin of tergalius IV; f. Anal margin of tergalius IV; g. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure37" httpUri="https://binary.pensoft.net/fig/874325" pageId="0" pageNumber="83">37d-f</figureCitation>
). Present on segments II-VII. Margin with small denticles intercalating fine, simple setae. Anal margin with both short and long, fine setae. Costal margin with alternating larger and smaller denticles. Tracheae not reaching inner and outer margins. Tergalius IV as long as length of segments V and VI combined.
</paragraph>
<paragraph id="9FC30EDDEC6F866F9A0B3EDCF86AED02" pageId="0" pageNumber="83">
<emphasis id="92380AD3BF4A5D5032078C016B17DB13" bold="true" italics="true" pageId="0" pageNumber="83">Paraproct</emphasis>
(Fig.
<figureCitation id="24EAF93DE7ADE60D7DC275A01F89076F" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis simbuensis sp. nov., larva morphology: a. Base of antenna; b. Antenna; c. Abdominal tergum IV; d. Tergalius IV; e. Anal margin of tergalius IV; f. Paraproct. Scale bars: 50 µm." figureDoi="10.3897/alpento.7.106089.figure31" httpUri="https://binary.pensoft.net/fig/874319" pageId="0" pageNumber="83">31f</figureCitation>
). Distally slightly expanded, with ca. 22 stout, marginal spines. Surface scattered with U-shaped scale bases. Cercotractor with numerous small, marginal spines.
</paragraph>
</subSubSection>
<subSubSection id="0FF90B28836A459EE31EA9B0846D8ADD" pageId="0" pageNumber="83" type="etymology">
<paragraph id="4193F585C0C6CFCAD3D8D6A4A2B1F716" pageId="0" pageNumber="83">Etymology.</paragraph>
<paragraph id="1321DECE1AB4AD77948EDCA5A685239B" pageId="0" pageNumber="83">Referring to the Kokoda Trek in Central Prov. of Papua New Guinea, along which the specimens were collected.</paragraph>
</subSubSection>
<subSubSection id="F8A715EC49A2E0E385A9C7545EA873D7" pageId="0" pageNumber="83" type="distribution">
<paragraph id="A29BFBC076AA6A2C95E89341B8CD92B9" pageId="0" pageNumber="83">Distribution.</paragraph>
<paragraph id="5FE8626C263D019B1951D7FFB78DDEAF" pageId="0" pageNumber="83">
Papua New Guinea, Central Prov. (Fig.
<figureCitation id="7A8730DCC6EF8242B3FF43F798B77043" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution of species treated in this study: a. Species of group Labiobaetis petersorum and L. kokoda sp. nov.; b. MOTUs of group Labiobaetis petersorum, not described in this study." figureDoi="10.3897/alpento.7.106089.figure38" httpUri="https://binary.pensoft.net/fig/874326" pageId="0" pageNumber="83">38a</figureCitation>
).
</paragraph>
<caption id="F6469955B52A2BC042FE874B25B43C11" doi="10.3897/alpento.7.106089.figure38" httpUri="https://binary.pensoft.net/fig/874326" pageId="0" pageNumber="83" start="Figure 38" startId="F38">
<paragraph id="6559A05EE77A18D2A8AA926C1D30F2D3" pageId="0" pageNumber="83">
<emphasis id="93BE8B4C8A44BB0A460D87C82227AC5A" bold="true" pageId="0" pageNumber="83">Figure 38.</emphasis>
Distribution of species treated in this study:
<emphasis id="311D9BFA0218A7453F6F99D92355E711" bold="true" pageId="0" pageNumber="83">a.</emphasis>
Species of group
<taxonomicName id="CDD29F46CB4040E70288AA4E2A568BAD" baseAuthorityName="Lugo-Ortiz &amp; McCafferty" baseAuthorityYear="1999" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis petersorum" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="petersorum">
<emphasis id="319E2FFC3ED2E8C15FE3F009B93F5F72" italics="true" pageId="0" pageNumber="83">Labiobaetis petersorum</emphasis>
</taxonomicName>
and
<taxonomicName id="248150EE1F1C6E725991097637983E9B" lsidName="L. kokoda" pageId="0" pageNumber="83" rank="species" species="kokoda">
<emphasis id="3C5A699A0B2A0E08E36A594C0F9D42D3" italics="true" pageId="0" pageNumber="83">L. kokoda</emphasis>
</taxonomicName>
sp. nov.;
<emphasis id="25AE5A8B0526638D305C68F6AD26E6E2" bold="true" pageId="0" pageNumber="83">b.</emphasis>
MOTUs of group
<taxonomicName id="67D42F73598637FC24C4E926C5EEDFAF" baseAuthorityName="Lugo-Ortiz &amp; McCafferty" baseAuthorityYear="1999" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis petersorum" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="petersorum">
<emphasis id="43336BEB964CCBCC305C278A0BACBA2D" italics="true" pageId="0" pageNumber="83">Labiobaetis petersorum</emphasis>
</taxonomicName>
, not described in this study.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="569A076A243E3C5559B146D066B580EC" pageId="0" pageNumber="83" type="biological aspects">
<paragraph id="1B299BBCB998A91D722B65ADDEC39371" pageId="0" pageNumber="83">Biological aspects.</paragraph>
<paragraph id="8F34BB10640F6BC09D6365DD887A1AE0" pageId="0" pageNumber="83">
The specimens were collected at an altitude of 1390 m, together with
<taxonomicName id="0668E072C1C29DBE2B35EFFBB86DAC98" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2018" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis lobatus" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="lobatus">
<emphasis id="F75D5F43D0E39448AF9963C578B58FFE" italics="true" pageId="0" pageNumber="83">Labiobaetis lobatus</emphasis>
</taxonomicName>
Kaltenbach &amp; Gattolliat, 2018 and other species of
<taxonomicName id="8D8293294D892B60A6B7361246C1ED1F" authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis" order="Ephemeroptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="24960167BCF872F419F191A4CB2C1B5D" italics="true" pageId="0" pageNumber="83">Labiobaetis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0F122BD64E99E568018E69E6A1F04F90" pageId="0" pageNumber="83" type="type material">
<paragraph id="3304C8AB88DC920160F7B9B4FF745C82" pageId="0" pageNumber="83">Type material.</paragraph>
<paragraph id="D3344124D5AB969BB485375F4D778C98" pageId="0" pageNumber="83">
<emphasis id="C4701CACA686355EE280C22E3FB93719" bold="true" pageId="0" pageNumber="83">
<emphasis id="29AFAA6995D2D9665EA75A4FF5E1FD29" italics="true" pageId="0" pageNumber="83">Holotype</emphasis>
.
</emphasis>
PAPUA NEW GUINEA • larva; Central Prov., Kokoda Trek;
<geoCoordinate id="364BFE71274F5DC20E4481E9AC314E7F" degrees="09" direction="south" minutes="00" orientation="latitude" precision="15" seconds="20" value="-9.005555">09°00'20&quot;S</geoCoordinate>
,
<geoCoordinate id="B4513B42A561DD8584A2D92AE4E3586C" degrees="147" direction="east" minutes="44" orientation="longitude" precision="15" seconds="15" value="147.7375">147°44'15&quot;E</geoCoordinate>
; 1390 m; i.2008; leg. Posman; (PNG 173); on slide; GBIFCH00975629; ZSM.
<emphasis id="056224AFCA275428EEBD5B5196CCF83E" bold="true" pageId="0" pageNumber="83">
<emphasis id="CE32C8FB2EE5FCC03D49123BFA365F95" italics="true" pageId="0" pageNumber="83">Paratypes</emphasis>
.
</emphasis>
PAPUA NEW GUINEA • 2 larvae; same data as holotype; on slides; GBIFCH00592683, GBIFCH00592684; MZL.
</paragraph>
</subSubSection>
</treatment>
</document>