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<document id="FA06C83FC7CACC260E48E9F20CA48D55" ID-CLB-Dataset="21278" ID-DOI="10.11646/zootaxa.4786.4.5" ID-GBIF-Dataset="7a4a33fc-a6ef-46c0-92a9-9f5f113da7e6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3990820" ID-ZooBank="33A7EFE7-356D-4D55-907C-D6C62D33C5A2" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597795496756" checkinUser="felipe" docAuthor="Vicentini, Heinrich, Wyler, Sofia &amp; Waringer, Johann" docDate="2020" docId="125D9655FC5BB341FF08E7E086AAFC6E" docLanguage="en" docName="zootaxa.4786.4.5.pdf.imf" docOrigin="Zootaxa 4786 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydropsyche doehleri Tobias 1972" docType="treatment" docVersion="6" lastPageNumber="540" masterDocId="EE64EE2DFC5AB344FF9FE10D8458FFA2" masterDocTitle="The larva of Hydropsyche doehleri Tobias 1972, based on Swiss material (Hydropsychidae, Trichoptera)" masterLastPageNumber="545" masterPageNumber="535" pageNumber="536" updateTime="1698844788515" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A01EC788B24B5D2A5A882C9AAEAF7BEC">The larva of Hydropsyche doehleri Tobias 1972, based on Swiss material (Hydropsychidae, Trichoptera)</mods:title>
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<mods:namePart id="F0943D9ACA62D16EFD6FCE67BEAA1E14">Vicentini, Heinrich</mods:namePart>
<mods:affiliation id="D998077C7BDD84F52241074BD3A768F7">Wasserschöpfi 48, CH- 8055 Zürich, Switzerland &amp; h. vic @ bluewin. ch; https: // orcid. org / 0000 - 0002 - 6854 - 1415 &amp; h. vic @ bluewin. ch; https: // orcid. org / 0000 - 0002 - 6854 - 1415</mods:affiliation>
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<mods:namePart id="997C7AC88826003FA28EEA994B703512">Wyler, Sofia</mods:namePart>
<mods:affiliation id="FB9E3BEEC4C3DBF89549374C27863AE0">SwissBOL, Université de Neuchâtel, Bellevaux 51, CH- 2000 Neuchâtel, Switzerland &amp; sofia. wyler @ unine. ch; https: // orcid. org / 0000 - 0002 - 3882 - 5679 &amp; sofia. wyler @ unine. ch; https: // orcid. org / 0000 - 0002 - 3882 - 5679</mods:affiliation>
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<mods:namePart id="33935EAA83E3AE5605AF1BC33154D753">Waringer, Johann</mods:namePart>
<mods:affiliation id="FA10CDEC73C0F6C10C0F257424D4B7FF">Department of Functional and Evolutionary Ecology, Division Limnology, University of Vienna, Althanstrasse 14, A- 1090 Vienna, Austria &amp; * johann. waringer @ univie. ac. at; https: // orcid. org / 0000 - 0002 - 0114 - 1636 &amp; * johann. waringer @ univie. ac. at; https: // orcid. org / 0000 - 0002 - 0114 - 1636</mods:affiliation>
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<subSection id="F37B3EA8FC5BB341FF08E7E086AAFC6E" lastPageId="5" lastPageNumber="540" pageId="1" pageNumber="536" type="multiple">
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<subSubSection id="D2EE74C8FC5BB345FF08E7E08043F8A5" box="[151,1051,1773,1800]" pageId="1" pageNumber="536" type="nomenclature">
<paragraph id="9A4B2743FC5BB345FF08E7E08043F8A5" blockId="1.[151,1051,1773,1800]" box="[151,1051,1773,1800]" pageId="1" pageNumber="536">
<heading id="C103902FFC5BB345FF08E7E08043F8A5" bold="true" box="[151,1051,1773,1800]" fontSize="11" level="2" pageId="1" pageNumber="536">
<emphasis id="A880FB51FC5BB345FF08E7E08043F8A5" bold="true" box="[151,1051,1773,1800]" pageId="1" pageNumber="536">
Description of the final instar larva of
<taxonomicName id="5DF45CC0FC5BB345FDE8E7E08043F8A5" ID-CoL="7VGMN" authority="Tobias 1972" authorityName="Tobias" authorityYear="1972" box="[631,1051,1773,1800]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="536" phylum="Arthropoda" rank="species" species="doehleri">
<emphasis id="A880FB51FC5BB345FDE8E7E08727F8AA" bold="true" box="[631,895,1773,1800]" italics="true" pageId="1" pageNumber="536">Hydropsyche doehleri</emphasis>
<bibRefCitation id="FE655AB2FC5BB345FC18E7E08043F8A5" author="Tobias, W." box="[903,1051,1773,1799]" pageId="1" pageNumber="536" pagination="245 - 268" refId="ref5552" refString="Tobias, W. (1972) Zur Kenntnis europaischer Hydropsychidae (Insecta: Trichoptera), II. Senckenbergiana Biologica, 53, 245 - 268." type="journal article" year="1972">Tobias 1972</bibRefCitation>
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<subSubSection id="D2EE74C8FC5BB341FF08E63886AAFC6E" lastPageId="5" lastPageNumber="540" pageId="1" pageNumber="536" type="description">
<paragraph id="9A4B2743FC5BB345FF08E638878CF8F2" blockId="1.[151,1437,1845,2016]" box="[151,980,1845,1872]" pageId="1" pageNumber="536">
<emphasis id="A880FB51FC5BB345FF08E6388555F8F2" bold="true" box="[151,269,1845,1872]" pageId="1" pageNumber="536">Biometry.</emphasis>
Body length 15.0
<quantity id="5D0C8AA6FC5BB345FE7BE6388611F8F2" box="[484,585,1845,1872]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="1" pageNumber="536" unit="mm" value="17.8">17.8 mm</quantity>
, head width
<quantity id="5D0C8AA6FC5BB345FD48E6388722F8F2" box="[727,890,1845,1872]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.72" metricValueMin="1.56" pageId="1" pageNumber="536" unit="mm" value="1.64" valueMax="1.72" valueMin="1.56">1.561.72 mm</quantity>
(n = 7).
</paragraph>
<paragraph id="9A4B2743FC5BB346FF58E65781DEFDC1" blockId="1.[151,1437,1845,2016]" lastBlockId="2.[151,1437,153,899]" lastPageId="2" lastPageNumber="537" pageId="1" pageNumber="536">
<emphasis id="A880FB51FC5BB345FF58E6578556F8D6" bold="true" box="[199,270,1882,1908]" pageId="1" pageNumber="536">Head.</emphasis>
Head capsule prognathous, surface smooth, roundish, maximum width at mid-length, tapering anteriorly and posteriorly (
<figureCitation id="02CF3BC6FC5BB345FED1E67085E3F83A" box="[334,443,1917,1944]" captionStart="FIGURES 25" captionStartId="3.[151,264,1914,1939]" captionTargetBox="[161,1427,203,1841]" captionTargetId="figure@3.[159,1428,201,1888]" captionTargetPageId="3" captionText="FIGURES 25. Hydropsyche doehleri Tobias 1972, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="1" pageNumber="536">Figs. 25</figureCitation>
). Coloration chestnut brown; on parietalia, with large pale rings around eyes (
<figureCitation id="02CF3BC6FC5BB345FA84E6708107F83A" box="[1307,1375,1917,1944]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="1" pageNumber="536">Fig. 6</figureCitation>
), and with yellow ventrolateral area extending from near anterior to posterior borders; posterior section of yellow area only half as wide as anterior area (
<figureCitation id="02CF3BC6FC5BB345FDBAE6C88633F842" box="[549,619,1989,2016]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="1" pageNumber="536">Fig. 6</figureCitation>
). Frontoclypeal apotome ogival, its maximum width at straight anterior border (
<figureCitation id="02CF3BC6FC58B346FF70E194856BFF11" box="[239,307,153,179]" captionStart="FIGURES 25" captionStartId="3.[151,264,1914,1939]" captionTargetBox="[161,1427,203,1841]" captionTargetId="figure@3.[159,1428,201,1888]" captionTargetPageId="3" captionText="FIGURES 25. Hydropsyche doehleri Tobias 1972, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="2" pageNumber="537">Fig. 2</figureCitation>
); anterior 2/3 of frontoclypeal suture slightly tapering posteriorly and almost parallel-sided; posterior branches of frontoclypeal suture meeting at near right angle (
<figureCitation id="02CF3BC6FC58B346FCCCE1B087C2FF75" box="[851,922,189,215]" captionStart="FIGURES 25" captionStartId="3.[151,264,1914,1939]" captionTargetBox="[161,1427,203,1841]" captionTargetId="figure@3.[159,1428,201,1888]" captionTargetPageId="3" captionText="FIGURES 25. Hydropsyche doehleri Tobias 1972, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="2" pageNumber="537">Fig. 2</figureCitation>
). Frontoclypeal step continuous, present also at center of sclerite (
<figureCitation id="02CF3BC6FC58B346FE1FE1EC859DFF59" box="[384,453,225,251]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 6</figureCitation>
, arrow). Typically, yellow patches on frontoclypeal apotome consisting of two pairs of yellow lateral patches, mesal anterior and posterior patches indistinct (
<figureCitation id="02CF3BC6FC58B346FC34E0088041FEBD" box="[939,1049,261,287]" captionStart="FIGURES 25" captionStartId="3.[151,264,1914,1939]" captionTargetBox="[161,1427,203,1841]" captionTargetId="figure@3.[159,1428,201,1888]" captionTargetPageId="3" captionText="FIGURES 25. Hydropsyche doehleri Tobias 1972, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="2" pageNumber="537">Figs. 24</figureCitation>
); in dark individuals, only anterior pair of lateral patches distinctly yellow (
<figureCitation id="02CF3BC6FC58B346FDC4E02486F8FEE1" box="[603,672,296,323]" captionStart="FIGURES 25" captionStartId="3.[151,264,1914,1939]" captionTargetBox="[161,1427,203,1841]" captionTargetId="figure@3.[159,1428,201,1888]" captionTargetPageId="3" captionText="FIGURES 25. Hydropsyche doehleri Tobias 1972, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="2" pageNumber="537">Fig. 5</figureCitation>
). Borders of frontoclypeal apotome and lateral sections of parietalia densely covered by secondary acuminate peg setae (
<figureCitation id="02CF3BC6FC58B346FD44E040871FFEC5" box="[731,839,333,359]" captionStart="FIGURES 25" captionStartId="3.[151,264,1914,1939]" captionTargetBox="[161,1427,203,1841]" captionTargetId="figure@3.[159,1428,201,1888]" captionTargetPageId="3" captionText="FIGURES 25. Hydropsyche doehleri Tobias 1972, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="2" pageNumber="537">Figs. 24</figureCitation>
,
<figureCitation id="02CF3BC6FC58B346FCCEE0408739FECA" box="[849,865,333,360]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">6</figureCitation>
). Ventrally, parietalia dark brown, with narrow yellow anterior border and narrow yellow band along anterior half of hypocranial suture (
<figureCitation id="02CF3BC6FC58B346FBC3E07C80F8FE29" box="[1116,1184,369,395]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 7</figureCitation>
). Striae of stridulatory file within dark brown coloration (
<figureCitation id="02CF3BC6FC58B346FD87E0988632FE0D" box="[536,618,405,431]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 10</figureCitation>
, dotted oval). Parietalia touching ventrally, thereby creating anterior ventral apotome (= submentum; sm in
<figureCitation id="02CF3BC6FC58B346FDBEE0B4863CFE71" box="[545,612,441,467]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 7</figureCitation>
); posterior ventral apotome reduced. Anterior section of submentum with pronounced median division (arrow in
<figureCitation id="02CF3BC6FC58B346FDD8E0D086D1FE55" box="[583,649,477,503]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 7</figureCitation>
), posterior section broadly triangular with short, wide lateral lobes (sm in
<figureCitation id="02CF3BC6FC58B346FF29E30C84A3FDB9" box="[182,251,513,539]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 7</figureCitation>
). Yellowish-brown mandibles slightly asymmetrical, with black cutting edges and teeth. Each mandible consisting of dorsal and ventral blades along mesal margin, each blade with one apical tooth; each ventral blade with 3 subapical teeth along its edge, each dorsal blade with small subapical tooth and curved cutting edge (
<figureCitation id="02CF3BC6FC58B346FA88E3448122FDC1" box="[1303,1402,585,611]" captionStart="FIGURES 25" captionStartId="3.[151,264,1914,1939]" captionTargetBox="[161,1427,203,1841]" captionTargetId="figure@3.[159,1428,201,1888]" captionTargetPageId="3" captionText="FIGURES 25. Hydropsyche doehleri Tobias 1972, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="2" pageNumber="537">
Fig.
<quantity id="5D0C8AA6FC58B346FAD4E3448122FDC1" box="[1355,1402,585,611]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="537" unit="m" value="2.0">2 m</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="9A4B2743FC58B346FF58E3608047FC21" blockId="2.[151,1437,153,899]" pageId="2" pageNumber="537">
<emphasis id="A880FB51FC58B346FF58E360857FFD25" bold="true" box="[199,295,621,647]" pageId="2" pageNumber="537">Thorax.</emphasis>
Pro-, meso- and metanota fully sclerotized; pronotal sclerites separated by median dorsal ecdysial line, the latter lacking on meso- and metanotal sclerites (
<figureCitation id="02CF3BC6FC58B346FD49E39C8742FD09" box="[726,794,657,683]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 8</figureCitation>
). Transverse ecdysial suture lacking on all notal sclerites. Tergites densely covered by small bristles. Pronotal sclerite (pn in
<figureCitation id="02CF3BC6FC58B346FC10E3B8878AFD6D" box="[911,978,693,719]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 8</figureCitation>
) dark brown, with black, straight posterior border. Mesonotal sclerites (msn in
<figureCitation id="02CF3BC6FC58B346FDC3E3D486C6FD51" box="[604,670,729,755]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 8</figureCitation>
) with black borders and sinuate, M-shaped posterior border heavily sclerotized in median section; mesonotal surface with dark median longitudinal band and dark bands extending from anterolateral corners to center of posterior border. Metanotal sclerites (mtn in
<figureCitation id="02CF3BC6FC58B346FC6EE22C8069FC99" box="[1009,1073,801,827]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="2" pageNumber="537">Fig. 8</figureCitation>
) yellowish, with black trimming along lateral border and anterolateral corners; posterior border slightly concave, with black central bar; black bands extending from anterolatereal corners to brownish area at rear center of sclerites.
</paragraph>
<caption id="CE8B77CBFC58B346FF08E601802AF87B" ID-DOI="http://doi.org/10.5281/zenodo.3990822" ID-Zenodo-Dep="3990822" httpUri="https://zenodo.org/record/3990822/files/figure.png" pageId="2" pageNumber="537" startId="2.[151,250,1804,1829]" targetBox="[161,1429,925,1770]" targetPageId="2">
<paragraph id="9A4B2743FC58B346FF08E601802AF87B" blockId="2.[151,1436,1804,2009]" pageId="2" pageNumber="537">
<emphasis id="A880FB51FC58B346FF08E601854CF887" bold="true" box="[151,276,1804,1829]" pageId="2" pageNumber="537">FIGURE 1.</emphasis>
MCL tree of four
<taxonomicName id="5DF45CC0FC58B346FE4FE601860DF886" authorityName="Curtis" authorityYear="1835" box="[464,597,1804,1828]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="A880FB51FC58B346FE4FE601860DF886" box="[464,597,1804,1828]" italics="true" pageId="2" pageNumber="537">Hydropsyche</emphasis>
</taxonomicName>
species with
<taxonomicName id="5DF45CC0FC58B346FD7FE6018014F887" authority="Pictet 1834" authorityName="Pictet" authorityYear="1834" box="[736,1100,1804,1829]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="torrentium">
<emphasis id="A880FB51FC58B346FD7FE6018789F886" box="[736,977,1804,1828]" italics="true" pageId="2" pageNumber="537">Rhyacophila torrentium</emphasis>
<bibRefCitation id="FE655AB2FC58B346FC47E6018014F887" author="Pictet, F. J." box="[984,1100,1804,1829]" pageId="2" pageNumber="537" refId="ref5338" refString="Pictet, F. J. (1834) Recherches pour Servir a l'Histoire et a l'Anatomie des Phryganides. A. Cherbuliez, rue de la Cit, Geneva, 235 pp. https: // doi. org / 10.5962 / bhl. title. 8547" type="book" year="1834">Pictet 1834</bibRefCitation>
</taxonomicName>
as outgroup taxon, based on 658 base pairs from mtCOI and including the herein associated life stages of
<taxonomicName id="5DF45CC0FC58B346FCE6E63C87B4F8EA" authorityName="Tobias" authorityYear="1972" box="[889,1004,1840,1864]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="doehleri">
<emphasis id="A880FB51FC58B346FCE6E63C87B4F8EA" box="[889,1004,1840,1864]" italics="true" pageId="2" pageNumber="537">H. doehleri</emphasis>
</taxonomicName>
. Branch lengths = number of substitutions per site; topology with highest log likelihood value (-2391.83). L = larvae, M = males, F = females. HV = material collected by H. Vicentini in stream Gaggiolo near Arzo, Southern Tessin, Switzerland; GBIFCH00660138 = male collected by H. Vicentini in stream Breggia at Caslaccio, Morbio Inferiore, Valle di Muggio, Southern Tessin, Switzerland on 7 August 2003; HMKKT743-10 = male collected by H. Malicky at Verzi, Savona Region, Italy, on 16 May 2006.
</paragraph>
</caption>
<caption id="CE8B77CBFC59B347FF08E6778000F814" ID-DOI="http://doi.org/10.5281/zenodo.3990824" ID-Zenodo-Dep="3990824" httpUri="https://zenodo.org/record/3990824/files/figure.png" pageId="3" pageNumber="538" startId="3.[151,264,1914,1939]" targetBox="[161,1427,203,1841]" targetPageId="3">
<paragraph id="9A4B2743FC59B347FF08E6778000F814" blockId="3.[151,1436,1914,1975]" pageId="3" pageNumber="538">
<emphasis id="A880FB51FC59B347FF08E677856CF831" bold="true" box="[151,308,1914,1939]" pageId="3" pageNumber="538">FIGURES 25</emphasis>
.
<taxonomicName id="5DF45CC0FC59B347FEDEE67786FCF830" authority="Tobias 1972" authorityName="Tobias" authorityYear="1972" box="[321,676,1914,1939]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="doehleri">
<emphasis id="A880FB51FC59B347FEDEE6778678F830" box="[321,544,1914,1938]" italics="true" pageId="3" pageNumber="538">Hydropsyche doehleri</emphasis>
<bibRefCitation id="FE655AB2FC59B347FDB9E67786FCF830" author="Tobias, W." box="[550,676,1914,1939]" pageId="3" pageNumber="538" pagination="245 - 268" refId="ref5552" refString="Tobias, W. (1972) Zur Kenntnis europaischer Hydropsychidae (Insecta: Trichoptera), II. Senckenbergiana Biologica, 53, 245 - 268." type="journal article" year="1972">Tobias 1972</bibRefCitation>
</taxonomicName>
, 5th instar larvae, heads, dorsal (m = mandible; dashed white lines indicate maximum width of frontoclypeal apotome and its near-right-angled apex). Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="CE8B77CBFC5EB340FF08E61280A9F849" ID-DOI="http://doi.org/10.5281/zenodo.3990826" ID-Zenodo-Dep="3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="4" pageNumber="539" startId="4.[151,264,1823,1848]" targetBox="[231,1362,183,1796]" targetPageId="4">
<paragraph id="9A4B2743FC5EB340FF08E61280A9F849" blockId="4.[151,1437,1823,2028]" pageId="4" pageNumber="539">
<emphasis id="A880FB51FC5EB340FF08E6128518F89A" bold="true" box="[151,320,1823,1848]" pageId="4" pageNumber="539">FIGURES 611</emphasis>
.
<taxonomicName id="5DF45CC0FC5EB340FED3E62D86F7F89A" authority="Tobias 1972" authorityName="Tobias" authorityYear="1972" box="[332,687,1823,1848]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="539" phylum="Arthropoda" rank="species" species="doehleri">
<emphasis id="A880FB51FC5EB340FED3E62D8672F89A" box="[332,554,1824,1848]" italics="true" pageId="4" pageNumber="539">Hydropsyche doehleri</emphasis>
<bibRefCitation id="FE655AB2FC5EB340FDAEE61286F7F89A" author="Tobias, W." box="[561,687,1823,1848]" pageId="4" pageNumber="539" pagination="245 - 268" refId="ref5552" refString="Tobias, W. (1972) Zur Kenntnis europaischer Hydropsychidae (Insecta: Trichoptera), II. Senckenbergiana Biologica, 53, 245 - 268." type="journal article" year="1972">Tobias 1972</bibRefCitation>
</taxonomicName>
, 5th instar larvae.
<emphasis id="A880FB51FC5EB340FCF7E612872DF89A" bold="true" box="[872,885,1823,1848]" pageId="4" pageNumber="539">6</emphasis>
, head, right lateral (arrow = frontoclypeal step present also at center of sclerite);
<emphasis id="A880FB51FC5EB340FE05E64985FFF8FE" bold="true" box="[410,423,1860,1884]" pageId="4" pageNumber="539">7</emphasis>
, head, ventral (sm = submentum; arrow = submental notch);
<emphasis id="A880FB51FC5EB340FB93E64E8078F8FE" bold="true" box="[1036,1056,1859,1884]" pageId="4" pageNumber="539">8,</emphasis>
thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum);
<emphasis id="A880FB51FC5EB340FDB5E66A866FF822" bold="true" box="[554,567,1895,1920]" pageId="4" pageNumber="539">9</emphasis>
, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite);
<emphasis id="A880FB51FC5EB340FE21E686858FF806" bold="true" box="[446,471,1931,1956]" pageId="4" pageNumber="539">10</emphasis>
, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation);
<emphasis id="A880FB51FC5EB340FC02E6A287EDF86A" bold="true" box="[925,949,1967,1992]" pageId="4" pageNumber="539">11</emphasis>
, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="9A4B2743FC5FB341FF58E1948642FEE1" blockId="5.[151,1437,153,972]" pageId="5" pageNumber="540">
Prosternal plate (a in
<figureCitation id="02CF3BC6FC5FB341FE24E19485A7FF11" box="[443,511,153,179]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Fig. 9</figureCitation>
) broadly trapezoidal, with elongate and enlarged anterolateral corners and pair of symmetrical, shallow concavities along black anterior border; with dark bar along posterior border (posterior part of a in
<figureCitation id="02CF3BC6FC5FB341FF75E1EC8577FF59" box="[234,303,225,251]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Fig. 9</figureCitation>
). Intersegmental fold posterior to prosternal plate with pair of sclerites, each one consisting of dark brown rhomboidal median section (pm in
<figureCitation id="02CF3BC6FC5FB341FDF8E00886F0FEBD" box="[615,680,261,287]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Fig. 9</figureCitation>
) and yellowish, funnel-shaped lateral section (pl in
<figureCitation id="02CF3BC6FC5FB341FB7BE008817FFEBD" box="[1252,1319,261,287]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Fig. 9</figureCitation>
), the latter with concave, black lateral border.
</paragraph>
<paragraph id="9A4B2743FC5FB341FF58E0408537FD9D" blockId="5.[151,1437,153,972]" pageId="5" pageNumber="540">
Foretrochantin (ft in
<figureCitation id="02CF3BC6FC5FB341FE2FE0408658FEC5" box="[432,512,333,359]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Fig. 10</figureCitation>
) yellow, with black ventral border, forked (arrows in
<figureCitation id="02CF3BC6FC5FB341FBC9E04080F0FEC5" box="[1110,1192,333,359]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Fig. 10</figureCitation>
). Mid- and hind pleural sclerites smaller, elongate triangular, with blackish ventral margins (mps and hps in
<figureCitation id="02CF3BC6FC5FB341FBF5E07C80E3FE29" box="[1130,1211,369,395]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Fig. 11</figureCitation>
, respectively). Legs yellow to yellowish brown, with numerous secondary setae on coxae, trochanters, and femora; tibiae and tarsi with smaller numbers of setae (
<figureCitation id="02CF3BC6FC5FB341FE21E0B4861DFE71" box="[446,581,441,467]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Figs. 10, 11</figureCitation>
). Each forefemur club-shaped, with broad, concave anterodorsal border apically. All femora with long ventral setal fringes. Tarsal claws long and slender, slightly curved, pointed, each with basal spur (
<figureCitation id="02CF3BC6FC5FB341FE84E30C85FDFDB9" box="[283,421,513,539]" captionStart="FIGURES 611" captionStartId="4.[151,264,1823,1848]" captionTargetBox="[231,1362,183,1796]" captionTargetId="figure@4.[224,1363,181,1798]" captionTargetPageId="4" captionText="FIGURES 611. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 6, head, right lateral (arrow = frontoclypeal step present also at center of sclerite); 7, head, ventral (sm = submentum; arrow = submental notch); 8, thoracic nota, dorsal (pn = pronotum; msn = mesonotum; mtn = metanotum); 9, prosternum, ventral (a = anterior prosternite; pm = median posterior prosternite; pl = lateral posterior prosternite); 10, head, right ventrolateral, and forelegs, with posterior face of right foreleg (ft = fore trochantin; arrows = forks of foretrochantin; dotted oval = striae of file for stridulation); 11, right midleg and hind leg, posterior faces (ml = midleg; hl = hind leg; mps = pleural sclerite of midleg; hps = pleural sclerite of hind leg). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990826" httpUri="https://zenodo.org/record/3990826/files/figure.png" pageId="5" pageNumber="540">Figs. 10, 11</figureCitation>
). Prosternum without gills, gills of meso- and metasterna each with branches originating from truncate base.
</paragraph>
<paragraph id="9A4B2743FC5FB341FF58E34586AAFC6E" blockId="5.[151,1437,153,972]" pageId="5" pageNumber="540">
<emphasis id="A880FB51FC5FB341FF58E345851BFDC1" bold="true" box="[199,323,584,611]" pageId="5" pageNumber="540">Abdomen.</emphasis>
Abdomen near white to pale purplishblue, with coloured areas subdivided by pale bands (
<figureCitation id="02CF3BC6FC5FB341FADBE34481C0FDC1" box="[1348,1432,585,611]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 12</figureCitation>
; coloration may be modified by preservation), and densely covered with hairlike setae (hs in
<figureCitation id="02CF3BC6FC5FB341FB27E3608152FD25" box="[1208,1290,621,647]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 13</figureCitation>
) and clavate scale hairs (sh in
<figureCitation id="02CF3BC6FC5FB341FECBE39C85FDFD09" box="[340,421,657,683]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 13</figureCitation>
). Each segment with 2 pairs of groups of dorsal primary setae, including posterior subdorsal pair (
<emphasis id="A880FB51FC5FB341FF4DE3BB84B3FD6D" box="[210,235,694,719]" italics="true" pageId="5" pageNumber="540">sa</emphasis>
<quantity id="5D0C8AA6FC5FB341FF74E3B8854DFD6D" box="[235,277,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="540" unit="in" value="2.0">2 in</quantity>
<figureCitation id="02CF3BC6FC5FB341FE84E3B88533FD6D" box="[283,363,693,719]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 12</figureCitation>
) and dorsolateral pair (
<emphasis id="A880FB51FC5FB341FDF2E3BB86DEFD6D" box="[621,646,694,719]" italics="true" pageId="5" pageNumber="540">sa</emphasis>
<quantity id="5D0C8AA6FC5FB341FD19E3B886E8FD6D" box="[646,688,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="540" unit="in" value="3.0">3 in</quantity>
<figureCitation id="02CF3BC6FC5FB341FD29E3B88750FD6D" box="[694,776,693,719]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 12</figureCitation>
). In addition, with pupal gill buds [not appearing before the final (fifth) instar in
<taxonomicName id="5DF45CC0FC5FB341FEE4E3D5866EFD51" authorityName="J Curtis" authorityYear="1835" box="[379,566,728,755]" class="Insecta" family="Hydropsychidae" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="540" phylum="Arthropoda" rank="family">Hydropsychidae</taxonomicName>
] (dotted oval in
<figureCitation id="02CF3BC6FC5FB341FD74E3D48765FD51" box="[747,829,729,755]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 12</figureCitation>
). Abdominal segments 16 with pair of multibranched gills matching morphology of gills on thoracic segments (g in
<figureCitation id="02CF3BC6FC5FB341FCCEE3F087FBFCB5" box="[849,931,765,791]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 12</figureCitation>
). Lateral fringe lacking. Pair of ventral sclerites on each of abdominal segments VIII and IX yellowish brown, conical, pointing in anterior direction, with scler- ites of segment IX much larger (vs in
<figureCitation id="02CF3BC6FC5FB341FDA5E24886D4FCFD" box="[570,652,837,863]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 14</figureCitation>
). Segment IX with dark dorsal sclerite and pair of dark lateral sclerites (ds and ls in
<figureCitation id="02CF3BC6FC5FB341FEBBE264852DFC21" box="[292,373,872,899]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 15</figureCitation>
, respectively). Segment X with anal papillae (ag in
<figureCitation id="02CF3BC6FC5FB341FC2AE264805EFC21" box="[949,1030,872,899]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 15</figureCitation>
). Anal prolegs with sclerites orange, each consisting of lateral area sclerite, ventral sole plate, large, sickle-shaped anal claw, and apicodorsal brush of long setae (la, vsp, ac, and st in
<figureCitation id="02CF3BC6FC5FB341FE63E2BC8615FC69" box="[508,589,944,971]" captionStart="FIGURES 1215" captionStartId="7.[151,264,1672,1697]" captionTargetBox="[218,1372,183,1645]" captionTargetId="figure@7.[213,1373,181,1646]" captionTargetPageId="7" captionText="FIGURES 1215. Hydropsyche doehleri Tobias 1972, 5th instar larvae. 12, abdominal segment VI, left lateral (g = multibranched gill; sa2, sa3 = setae of setal areas 2 and 3; dotted oval = pupal gill bud); 13, abdominal tergum V, detail of right dorsolateral section (hs = hairlike setae; sh = clavate scale hairs); 14, abdominal segments VII to X and anal prolegs, ventral (ac = anal claw; vs = paired ventral sclerites of abdominal sterna VIII and IX); 15, abdominal segments V to IX and anal prolegs, right lateral (ac = anal claw; ag = anal papillae; ds = dorsal sclerite; la = lateral area sclerite; ls = lateral sclerite; st = setal tuft; vs = ventral sclerites; vsp = ventral sole plate; arrow = gills lacking on abdominal segment VII). Scale bars: 0.5 mm (except Fig. 13: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3990828" httpUri="https://zenodo.org/record/3990828/files/figure.png" pageId="5" pageNumber="540">Fig. 15</figureCitation>
, respectively).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>