535 lines
110 KiB
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535 lines
110 KiB
XML
<document id="569A731367AC43308AB3F03108D84582" ID-CLB-Dataset="274285" ID-DOI="10.5852/ejt.2023.893.2285" ID-GBIF-Dataset="e441bc9d-bb44-4ee6-b29f-a6724b96592b" ID-ISSN="2118-9773" ID-Zenodo-Dep="8385630" ID-ZooBank="D9DA4879-BC1D-4947-96E8-7D574DE334BA" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1695891225180" checkinUser="plazi" docAuthor="Leng, Zhaoqi, Tran, Anh Duc & Ye, Zhen" docDate="2023" docId="045E878DFFCD292E3A9AFEBA75710D26" docLanguage="en" docName="EJT.2023.893.1-73.pdf" docOrigin="European Journal of Taxonomy 893 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2285/9863" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhyacobates bui Leng & Tran 2023, sp. nov." docType="treatment" docUuid="8A1A2DC9-C314-4458-AF0C-752FEECA39C7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="19" masterDocId="F867FFF5FFDC293B381FFFB270530F1A" masterDocTitle="Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="18" updateTime="1699690649759" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5DA400070652CDF45AAFF278A9CD13A9">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:namePart id="5259C22648B64E715D640D8FAA79F49C">Leng, Zhaoqi</mods:namePart>
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<mods:affiliation id="F44142179D021C5742CB9764DA7E01D3">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:nameIdentifier id="0E34FC6F3888DF2FEF7278AD0C889C4B" type="email">zhaoqileng0712@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FF3B457147AE15F5EFCF015D9E06AE7D">Tran, Anh Duc</mods:namePart>
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<mods:affiliation id="674F291BEECFE384C4F8AB48369AD5B1">Faculty of Biology, VNU University of Science, Vietnam National University, Hanoi, Nguyen Trai, Thanh Xuan, Hanoi, Vietnam.</mods:affiliation>
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<mods:nameIdentifier id="0A23713A84CF0897BA0A1CAEDEE8CB69" type="email">tran.anhduc@hus.edu.vn</mods:nameIdentifier>
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<mods:namePart id="9FC187EC14BE80663E9558759F82240D">Ye, Zhen</mods:namePart>
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<mods:affiliation id="7752DD51D4FF65603890574057DB8D67">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:title id="BD51BCFF703DD3AE08E62120E85B81EF">European Journal of Taxonomy</mods:title>
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<mods:date id="025E16C7554497AA39F763A34D5B2C38">2023</mods:date>
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<treatment id="045E878DFFCD292E3A9AFEBA75710D26" ID-DOI="http://doi.org/10.5281/zenodo.10064059" ID-GBIF-Taxon="214246589" ID-Zenodo-Dep="10064059" LSID="urn:lsid:zoobank.org:act:8A1A2DC9-C314-4458-AF0C-752FEECA39C7" httpUri="http://treatment.plazi.org/id/045E878DFFCD292E3A9AFEBA75710D26" lastPageId="21" lastPageNumber="19" pageId="17" pageNumber="18">
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<heading id="D70081F7FFCD292A3A9AFEBA73FC0E39" box="[645,943,264,291]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
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<taxonomicName id="4BF74D18FFCD292A3A9AFEBA731E0E39" authority="Leng & Tran, 2023" authorityName="Leng & Tran" authorityYear="2023" box="[645,845,264,291]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bui" status="sp. nov.">
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<emphasis id="BE83EA89FFCD292A3A9AFEBA731E0E39" bold="true" box="[645,845,264,291]" italics="true" pageId="17" pageNumber="18">Rhyacobates bui</emphasis>
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<taxonomicNameLabel id="A5B057F2FFCD292A3B4BFEBB73FC0E39" box="[852,943,265,291]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C4ED6510FFCD292A3A45FEFD73890E70" box="[602,986,335,362]" pageId="17" pageNumber="18" type="description">
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<paragraph id="8C48369BFFCD292A3A45FEFD73890E70" blockId="17.[359,1228,264,362]" box="[602,986,335,362]" pageId="17" pageNumber="18">
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<figureCitation id="14CC2A1EFFCD292A3A45FEFD72EA0E73" box="[602,697,335,361]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="17" pageNumber="18">Figs 3A</figureCitation>
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,
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<figureCitation id="14CC2A1EFFCD292A3AD9FEFD72BE0E73" box="[710,749,335,361]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="17" pageNumber="18">4A</figureCitation>
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,
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<figureCitation id="14CC2A1EFFCD292A3AE6FEFD734C0E70" box="[761,799,335,362]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="17" pageNumber="18">5A</figureCitation>
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,
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<figureCitation id="14CC2A1EFFCD292A3B34FEFD73010E73" box="[811,850,335,361]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="17" pageNumber="18">6A</figureCitation>
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<figureCitation id="14CC2A1EFFCD292A3B41FEFD73D70E73" box="[862,900,335,361]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="17" pageNumber="18">8A</figureCitation>
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<figureCitation id="14CC2A1EFFCD292A3B8FFEFD73890E70" box="[912,986,335,362]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="19.[189,232,1963,1989]" captionStartId-1="20.[189,232,1787,1813]" captionTargetBox-0="[197,1362,400,1884]" captionTargetBox-1="[280,1309,276,1715]" captionTargetId-0="figure-11@19.[189,1398,372,1910]" captionTargetId-1="figure-14@20.[245,1342,265,1743]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="Fig. 15. Habitus of Rhyacobates bui sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." captionText-1="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385668" figureDoi-1="http://doi.org/10.5281/zenodo.8385670" httpUri-0="https://zenodo.org/record/8385668/files/figure.png" httpUri-1="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">15–16</figureCitation>
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</paragraph>
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<subSubSection id="C4ED6510FFCD292A38A2FE22726D0C0C" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="8C48369BFFCD292A38A2FE2271650EB0" blockId="17.[189,310,400,426]" box="[189,310,400,426]" pageId="17" pageNumber="18">
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<heading id="D70081F7FFCD292A38A2FE2271650EB0" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="BE83EA89FFCD292A38A2FE2271650EB0" bold="true" box="[189,310,400,426]" pageId="17" pageNumber="18">Diagnosis</emphasis>
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<paragraph id="8C48369BFFCD292A38A2FE0D726D0C0C" blockId="17.[188,1400,447,791]" pageId="17" pageNumber="18">
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Body length of apterous females 8.81–9.30, of apterous males 6.50–6.91. In both sexes, mesonotum and metanotum completely black, without median yellow stripe (
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<figureCitation id="14CC2A1EFFCD292A3B88FE5073A80EE7" box="[919,1019,482,509]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="17" pageNumber="18">Figs 3A</figureCitation>
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,
|
||
<figureCitation id="14CC2A1EFFCD292A3C15FE5174610EE7" box="[1034,1074,483,509]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="17" pageNumber="18">4A</figureCitation>
|
||
). Female: posterior margin of mediotergite VII with a distinctly long, slender median process (
|
||
<figureCitation id="14CC2A1EFFCD292A3BFBFDB4742A0D3A" box="[996,1145,518,544]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">Fig. 16A, D</figureCitation>
|
||
); posterior margin of abdominal segment VII with five processes, including a blunt process terminating each connexivum, a pair of blunt processes laterally, and a semicircular median process ventrally (
|
||
<figureCitation id="14CC2A1EFFCD292A3C99FDFE75480D7C" box="[1158,1307,588,614]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">Fig. 16D–G</figureCitation>
|
||
). Male: middle trochanter with 3–4 small spines; middle femur with scattered small spines arranged in a row, but with irregular distance between spines (
|
||
<figureCitation id="14CC2A1EFFCD292A3ACAFD2073680DB6" box="[725,827,658,684]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">Fig. 16J</figureCitation>
|
||
); length of middle tibia ca 1.6 times length of hind tibia; proctiger laterally with subtrapezoid lobes, dorsally with brownish setae (
|
||
<figureCitation id="14CC2A1EFFCD292A3CA1FD0475700DCA" box="[1214,1315,694,720]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="17" pageNumber="18">Figs 6A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCD292A3D2DFD04753B0DCA" box="[1330,1384,694,720]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">16K</figureCitation>
|
||
); paramere relatively slender, strongly curved at basal third, distal part tapering towards distinctly curved and narrow apex (
|
||
<figureCitation id="14CC2A1EFFCD292A398FFD4E71A20C0C" box="[400,497,764,790]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="17" pageNumber="18">Figs 8A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCD292A39E1FD4E72620C0C" box="[510,561,764,790]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">16L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFCD292A38A2FC8C75240CB0" pageId="17" pageNumber="18" type="etymology">
|
||
<paragraph id="8C48369BFFCD292A38A2FC8C71170C42" blockId="17.[189,324,830,856]" box="[189,324,830,856]" pageId="17" pageNumber="18">
|
||
<heading id="D70081F7FFCD292A38A2FC8C71170C42" bold="true" box="[189,324,830,856]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="BE83EA89FFCD292A38A2FC8C71170C42" bold="true" box="[189,324,830,856]" pageId="17" pageNumber="18">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38A2FCDE75240CB0" blockId="17.[189,1399,876,938]" pageId="17" pageNumber="18">
|
||
This species dedicated to Prof. Wenjun Bu (Institute of Entomology, Nankai University,
|
||
<collectingCountry id="F4E0760BFFCD292A3CE2FCDE751B0C9C" box="[1277,1352,876,902]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
|
||
), in recognition of his contributions to the study ofthe taxonomy, phylogeny and biogeography of
|
||
<taxonomicName id="4BF74D18FFCD292A3CFAFC2275200CB0" authorityName="Latreille" authorityYear="1810" box="[1253,1395,912,938]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="3" phylum="Arthropoda" rank="subOrder" subOrder="Heteroptera">Heteroptera</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFCD292A38A2FC6275520A75" pageId="17" pageNumber="18" type="materials_examined">
|
||
<paragraph id="8C48369BFFCD292A38A2FC6271F90CF0" blockId="17.[189,426,976,1002]" box="[189,426,976,1002]" pageId="17" pageNumber="18">
|
||
<heading id="D70081F7FFCD292A38A2FC6271F90CF0" bold="true" box="[189,426,976,1002]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="BE83EA89FFCD292A38A2FC6271F90CF0" bold="true" box="[189,426,976,1002]" pageId="17" pageNumber="18">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38F5FC4D71B30B00" blockId="17.[189,1398,1023,1120]" box="[234,480,1023,1050]" pageId="17" pageNumber="18">
|
||
<emphasis id="BE83EA89FFCD292A38F5FC4D710F0B03" bold="true" box="[234,348,1023,1049]" pageId="17" pageNumber="18">
|
||
<typeStatus id="534C8839FFCD292A38F5FC4D710F0B03" box="[234,348,1023,1049]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="14CC2A1EFFCD292A3973FC4D71850B00" box="[364,470,1023,1050]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[197,1362,400,1884]" captionTargetId="figure-11@19.[189,1398,372,1910]" captionTargetPageId="19" captionText="Fig. 15. Habitus of Rhyacobates bui sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385668" httpUri="https://zenodo.org/record/8385668/files/figure.png" pageId="17" pageNumber="18">Fig. 15A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38A2FB91753F0B7A" blockId="17.[189,1398,1023,1120]" pageId="17" pageNumber="18">
|
||
<materialsCitation id="3C9F3CC6FFCD292A38A2FB91753F0B7A" ID-GBIF-Occurrence="4429869386" collectingDate="2019-08-13" collectionCode="NKUM" collectorName="Zhen Ye & Si-qi Wang" country="China" county="Shang-si County" elevation="350" latitude="21.901138" location="City" longLatPrecision="1" longitude="107.90614" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="holotype">
|
||
<collectingCountry id="F4E0760BFFCD292A38A2FB91714B0B27" box="[189,280,1059,1085]" name="China" pageId="17" pageNumber="18">CHINA</collectingCountry>
|
||
•
|
||
<specimenCount id="9AF1FD12FFCD292A392CFB91711A0B27" box="[307,329,1059,1085]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
(apterous);
|
||
<collectingRegion id="4E33F879FFCD292A39C6FB9072E20B26" box="[473,689,1058,1084]" country="China" name="Guangxi" pageId="17" pageNumber="18">Guangxi Province</collectingRegion>
|
||
, Fang-cheng-gang
|
||
<location id="89286040FFCD292A3B8BFB90739B0B26" LSID="urn:lsid:plazi:treatment:045E878DFFCD292E3A9AFEBA75710D26:89286040FFCD292A3B8BFB90739B0B26" box="[916,968,1058,1084]" country="China" county="Shang-si County" latitude="21.901138" longLatPrecision="1" longitude="107.90614" name="City" pageId="17" pageNumber="18" stateProvince="Guangxi">City</location>
|
||
,
|
||
<collectingCounty id="65294E17FFCD292A3BCBFB9074C80B27" box="[980,1179,1058,1085]" pageId="17" pageNumber="18">Shang-si County</collectingCounty>
|
||
, Shi-wan-da-shan;
|
||
<geoCoordinate id="E9C3505CFFCD292A38A2FBF671090B7A" box="[189,346,1092,1120]" degrees="21" direction="north" minutes="54" orientation="latitude" pageId="17" pageNumber="18" precision="1" seconds="4.1" value="21.901138">21°54′4.1″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFCD292A3979FBF6724E0B7A" box="[358,541,1092,1120]" degrees="107" direction="east" minutes="54" orientation="longitude" pageId="17" pageNumber="18" precision="1" seconds="22.1" value="107.90614">107°54′22.1″ E</geoCoordinate>
|
||
;
|
||
<elevation id="07DAD1A8FFCD292A3A35FBF472BD0B7A" box="[554,750,1094,1120]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="17" pageNumber="18" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
|
||
<quantity id="4B0F9B7EFFCD292A3A35FBF472FC0B7A" box="[554,687,1094,1120]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="17" pageNumber="18" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">300–400 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="F849105BFFCD292A3AE3FBF473CD0B7A" box="[764,926,1094,1120]" pageId="17" pageNumber="18" value="2019-08-13">
|
||
<collectingDate id="E80DE9B3FFCD292A3AE3FBF473CD0B7A" box="[764,926,1094,1120]" pageId="17" pageNumber="18" value="2019-08-13">13 Aug. 2019</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFCD292A3BB5FBF4745F0B7A" box="[938,1036,1094,1120]" pageId="17" pageNumber="18">Zhen Ye</collectorName>
|
||
and
|
||
<collectorName id="2102534DFFCD292A3C5AFBF4749A0B7A" box="[1093,1225,1094,1120]" pageId="17" pageNumber="18">Si-qi Wang</collectorName>
|
||
leg.;
|
||
<collectionCode id="EAE6AE5EFFCD292A3D14FBF4753A0B7A" box="[1291,1385,1094,1120]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="17" pageNumber="18">NKUM</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38F5FB3571BA0BB8" blockId="17.[189,1049,1159,1221]" box="[234,489,1159,1186]" pageId="17" pageNumber="18">
|
||
<emphasis id="BE83EA89FFCD292A38F5FB3571340BBB" bold="true" box="[234,359,1159,1185]" pageId="17" pageNumber="18">
|
||
<typeStatus id="534C8839FFCD292A38F5FB3571340BBB" box="[234,359,1159,1185]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="14CC2A1EFFCD292A3967FB3571B30BB8" box="[376,480,1159,1186]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[197,1362,400,1884]" captionTargetId="figure-11@19.[189,1398,372,1910]" captionTargetPageId="19" captionText="Fig. 15. Habitus of Rhyacobates bui sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385668" httpUri="https://zenodo.org/record/8385668/files/figure.png" pageId="17" pageNumber="18">Fig. 15B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38A2FB18744A0BDF" blockId="17.[189,1049,1159,1221]" box="[189,1049,1192,1221]" pageId="17" pageNumber="18">
|
||
<materialsCitation id="3C9F3CC6FFCD292A38A2FB1874460BDF" ID-GBIF-Occurrence="4429869379" box="[189,1045,1192,1221]" collectingDate="2019-08-13" collectionCode="NKUM" collectorName="Zhen Ye & Si-qi Wang" country="China" county="Shang-si County" elevation="350" latitude="21.901138" location="City" longLatPrecision="1" longitude="107.90614" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
|
||
<collectingCountry id="F4E0760BFFCD292A38A2FB18714B0BDE" box="[189,280,1194,1220]" name="China" pageId="17" pageNumber="18">CHINA</collectingCountry>
|
||
• 3 ♁♁ (apterous); same collection data as for holotype;
|
||
<collectionCode id="EAE6AE5EFFCD292A3BA8FB1974460BDF" box="[951,1045,1195,1221]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="17" pageNumber="18">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38A2FB5E75520A75" blockId="17.[189,1399,1258,1392]" pageId="17" pageNumber="18">
|
||
<materialsCitation id="3C9F3CC6FFCD292A38A2FB5E74AF0A33" ID-GBIF-Occurrence="4429869301" collectingDate="2020-11-07" collectionCode="ZVNU" collectorName="A. D. Tran" country="Vietnam" latitude="21.816471" location="oc Binh" longLatPrecision="1" longitude="106.99897" pageId="17" pageNumber="18" specimenCount="3" specimenCount-female="3" stateProvince="ang SOn Province">
|
||
<collectingCountry id="F4E0760BFFCD292A38A2FB5E71170A1C" box="[189,324,1260,1286]" name="Vietnam" pageId="17" pageNumber="18">VIETNAM</collectingCountry>
|
||
• 10 ♁♁,
|
||
<specimenCount id="9AF1FD12FFCD292A39A4FB5E71AE0A1C" box="[443,509,1260,1286]" pageId="17" pageNumber="18" type="female">3 ♀♀</specimenCount>
|
||
(apterous); L
|
||
<collectingRegion id="4E33F879FFCD292A3A83FB5E733D0A1C" box="[668,878,1260,1288]" pageId="17" pageNumber="18">ạng SƠn Province</collectingRegion>
|
||
, L
|
||
<location id="89286040FFCD292A3B94FB5E73B90A1C" LSID="urn:lsid:plazi:treatment:045E878DFFCD292E3A9AFEBA75710D26:89286040FFCD292A3B94FB5E73B90A1C" box="[907,1002,1260,1286]" country="Vietnam" latitude="21.816471" longLatPrecision="1" longitude="106.99897" name="oc Binh" pageId="17" pageNumber="18" stateProvince="ang SOn Province">ộc Bình</location>
|
||
, M
|
||
<location id="89286040FFCD292A3C10FB5E74370A12" LSID="urn:lsid:plazi:treatment:045E878DFFCD292E3A9AFEBA75710D26:89286040FFCD292A3C10FB5E74370A12" box="[1039,1124,1260,1288]" country="Vietnam" latitude="21.816471" longLatPrecision="1" longitude="106.99897" name="au SOn" pageId="17" pageNumber="18" stateProvince="ang SOn Province">ẫu SƠn</location>
|
||
, upstream of
|
||
<location id="89286040FFCD292A3D1EFB5E715C0A33" LSID="urn:lsid:plazi:treatment:045E878DFFCD292E3A9AFEBA75710D26:89286040FFCD292A3D1EFB5E715C0A33" country="Vietnam" latitude="21.816471" longLatPrecision="1" longitude="106.99897" name="Long Dau stream" pageId="17" pageNumber="18" stateProvince="ang SOn Province">Long Đǻu stream</location>
|
||
;
|
||
<geoCoordinate id="E9C3505CFFCD292A3906FABF71970A33" box="[281,452,1293,1321]" degrees="21" direction="north" minutes="48" orientation="latitude" pageId="17" pageNumber="18" precision="1" seconds="59.3" value="21.816471">21°48′59.3″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFCD292A39D0FABF72D60A33" box="[463,645,1293,1321]" degrees="106" direction="east" minutes="59" orientation="longitude" pageId="17" pageNumber="18" precision="1" seconds="56.3" value="106.99897">106°59′56.3″ E</geoCoordinate>
|
||
;
|
||
<date id="F849105BFFCD292A3A8FFABD73720A33" box="[656,801,1295,1321]" pageId="17" pageNumber="18" value="2020-11-07">
|
||
<collectingDate id="E80DE9B3FFCD292A3A8FFABD73720A33" box="[656,801,1295,1321]" pageId="17" pageNumber="18" value="2020-11-07">7 Nov. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFCD292A3B34FABD73CC0A33" box="[811,927,1295,1321]" pageId="17" pageNumber="18">A.D. Tran</collectorName>
|
||
<emphasis id="BE83EA89FFCD292A3BB9FAA2738C0A33" box="[934,991,1295,1321]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
|
||
leg.; TAD20-37;
|
||
<collectionCode id="EAE6AE5EFFCD292A3CB4FABD74AF0A33" box="[1195,1276,1295,1321]" pageId="17" pageNumber="18">ZVNU</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3C9F3CC6FFCD292A3D0CFABD73310A57" ID-GBIF-Occurrence="4429869359" collectingDate="2020-11-07" collectionCode="NHMW" collectorName="A. D. Tran" country="Vietnam" latitude="21.816471" location="oc Binh" longLatPrecision="1" longitude="106.99897" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="ang SOn Province">
|
||
1 ♁,
|
||
<specimenCount id="9AF1FD12FFCD292A3D53FABD75250A33" box="[1356,1398,1295,1321]" count="1" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
|
||
(apterous); same collection data as for preceding;
|
||
<collectionCode id="EAE6AE5EFFCD292A3B1DFA8173310A57" box="[770,866,1331,1357]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3C9F3CC6FFCD292A3B66FA8071970A6A" ID-GBIF-Occurrence="4429869382" collectingDate="2020-11-07" collectionCode="NKUM" collectorName="A. D. Tran" country="Vietnam" latitude="21.816471" location="oc Binh" longLatPrecision="1" longitude="106.99897" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="ang SOn Province">
|
||
1 ♁,
|
||
<specimenCount id="9AF1FD12FFCD292A3BADFA80738E0A57" box="[946,989,1330,1357]" count="1" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
|
||
(apterous); same collection data as for preceding;
|
||
<collectionCode id="EAE6AE5EFFCD292A3975FAE471970A6A" box="[362,452,1366,1392]" collectionName="China, Tianjin, Tianjin, Nankai University" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="17" pageNumber="18">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3C9F3CC6FFCD292A39C3FAE774AE0A75" ID-GBIF-Occurrence="4429869373" box="[476,1277,1363,1392]" collectingDate="2020-11-07" collectionCode="ZRC" collectorName="A. D. Tran" country="Vietnam" latitude="21.816471" location="oc Binh" longLatPrecision="1" longitude="106.99897" pageId="17" pageNumber="18" specimenCount="2" specimenCount-female="2" stateProvince="ang SOn Province">
|
||
2 ♁♁,
|
||
<specimenCount id="9AF1FD12FFCD292A3A32FAE7723C0A6A" box="[557,623,1365,1392]" count="2" pageId="17" pageNumber="18" type="female">2 ♀♀</specimenCount>
|
||
(apterous); same collection data as for preceding;
|
||
<collectionCode id="EAE6AE5EFFCD292A3CDFFAE774AE0A75" box="[1216,1277,1365,1391]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="17" pageNumber="18">ZRC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFCD292938A2FA2471250AAD" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="8C48369BFFCD292A38A2FA24711D0AAA" blockId="17.[189,334,1430,1456]" box="[189,334,1430,1456]" pageId="17" pageNumber="18">
|
||
<heading id="D70081F7FFCD292A38A2FA24711D0AAA" bold="true" box="[189,334,1430,1456]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="BE83EA89FFCD292A38A2FA24711D0AAA" bold="true" box="[189,334,1430,1456]" pageId="17" pageNumber="18">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38F5FA7771EB0AC5" blockId="17.[185,1399,1477,1751]" box="[234,440,1477,1503]" pageId="17" pageNumber="18">
|
||
<emphasis id="BE83EA89FFCD292A38F5FA7771EB0AC5" bold="true" box="[234,440,1477,1503]" pageId="17" pageNumber="18">Apterous female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38A2FA5B731A09CC" blockId="17.[185,1399,1477,1751]" pageId="17" pageNumber="18">
|
||
<smallCapsWord id="8AAEA047FFCD292A38A2FA5B7138091B" baselines="1532,1532" box="[189,363,1513,1539]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="17" pageNumber="18">MEASUREMENTS</smallCapsWord>
|
||
. Body length 8.81–9.30 (
|
||
<typeStatus id="534C8839FFCD292A3ABFFA5A73540918" box="[672,775,1512,1538]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
8.81), width 2.34–2.52 (
|
||
<typeStatus id="534C8839FFCD292A3C2CFA5A74C90918" box="[1075,1178,1512,1538]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
2.52), head width 1.41, interocular width 0.75, eye length (dorsal view) 0.62; relative lengths of antennal segments I–IV: 3.34: 1.02: 1.12:0.71; pronotum: length 0.72, width 1.51; mesonotum: length 2.34, width 2.52 (
|
||
<typeStatus id="534C8839FFCD292A3D10F99D75240953" box="[1295,1399,1583,1609]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
2.52); metanotum: length 0.84, width 2.21; abdomen length (ventral view) 4.16; abdominal sternum VII: length 1.16, width 1.15; abdominal mediotergite I: length 0.35, width 0.92; relative lengths of leg segments (femur:tibia: tarsal segment I: tarsal segment II): fore leg: 3.56: 2.84:1.46:0.78, middle leg: 10.01:5.94:2.63:0.44, hind leg: 10.16: 3.88: 0.11:0.17.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCD292A38A2F94F727F08BE" blockId="17.[189,1399,1789,1956]" pageId="17" pageNumber="18">
|
||
<smallCapsWord id="8AAEA047FFCD292A38A2F94F711B080C" baselines="1808,1809" box="[189,328,1789,1815]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="17" pageNumber="18">COLORATION</smallCapsWord>
|
||
. Median black spot of head reduced, indistinctly bifid or dissolved into smaller spots posteriorly (
|
||
<figureCitation id="14CC2A1EFFCD292A3950F89271E30820" box="[335,432,1824,1850]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="17" pageNumber="18">Fig. 3A</figureCitation>
|
||
). Pronotum mainly black with a median brownish-yellow spot. Mesonotum and metanotum completely black, without yellow markings (
|
||
<figureCitation id="14CC2A1EFFCD292A3B4DF8F173E00844" box="[850,947,1859,1886]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="17" pageNumber="18">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCD292A3BA0F8F173A60847" box="[959,1013,1859,1885]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">16A</figureCitation>
|
||
). Connexivum dorsally blackish. Venter of female mainly light yellow, except mesosternum and mesopleuron black laterally (
|
||
<figureCitation id="14CC2A1EFFCD292A3CE2F8D5753B089B" box="[1277,1384,1895,1921]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="17" pageNumber="18">Fig. 16B</figureCitation>
|
||
). Abdominal venter light yellow.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938A2FEBB74070D26" blockId="18.[188,1401,265,573]" pageId="18" pageNumber="19">
|
||
<smallCapsWord id="8AAEA047FFCE292938A2FEBB717F0E3B" baselines="284,284" box="[189,300,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="18" pageNumber="19">ABDOMEN</smallCapsWord>
|
||
. Abdomen relatively long, nearly straight, gradually tapering towards apex (
|
||
<figureCitation id="14CC2A1EFFCE29293CDDFEBB75750E3E" box="[1218,1318,265,292]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="18" pageNumber="19">Figs 5A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCE29293D2BFEBB753A0E39" box="[1332,1385,265,291]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">16C</figureCitation>
|
||
). Connexiva erect on abdominal segments I–VI, parallel to each other above mediotergites, reflexed over terminal mediotergites (
|
||
<figureCitation id="14CC2A1EFFCE29293A08FEFD72FE0E73" box="[535,685,335,361]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Fig. 16A, D</figureCitation>
|
||
). Abdominal mediotergite I not swollen, fully covered with silvery pubescence (
|
||
<figureCitation id="14CC2A1EFFCE292939B4FEC0725B0E97" box="[427,520,370,397]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="18" pageNumber="19">Fig. 3A</figureCitation>
|
||
). Posterior margin of mediotergite VII with a distinctly long median process (
|
||
<figureCitation id="14CC2A1EFFCE292938DBFE2471010EAA" box="[196,338,406,432]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Fig. 16A, D</figureCitation>
|
||
), about half the length of mediotergite VII (excluding the process). Abdominal segment VII elongate, nearly as long as three preceding abdominal segments, completely enclosing genital segments. Abdominal dorsal margin not curved in lateral view (
|
||
<figureCitation id="14CC2A1EFFCE29293B5AFE6E73FA0EED" box="[837,937,476,503]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="18" pageNumber="19">Figs 5A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCE29293BA6FE6E73BD0EEC" box="[953,1006,476,502]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">16C</figureCitation>
|
||
). Posterior margin of abdominal segment VII with five processes, including a blunt process terminating each connexivum, a pair of blunt processes laterally, and a semicircular median process ventrally (
|
||
<figureCitation id="14CC2A1EFFCE29293BAAFD9074140D26" box="[949,1095,546,572]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Fig. 16D–G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938F5FDD671F20D64" blockId="18.[185,1399,612,885]" box="[234,417,612,638]" pageId="18" pageNumber="19">
|
||
<emphasis id="BE83EA89FFCE292938F5FDD671F20D64" bold="true" box="[234,417,612,638]" pageId="18" pageNumber="19">Apterous male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938A2FD3A71BC0C6F" blockId="18.[185,1399,612,885]" pageId="18" pageNumber="19">
|
||
<smallCapsWord id="8AAEA047FFCE292938A2FD3A713D0DBA" baselines="667,667" box="[189,366,648,674]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="18" pageNumber="19">MEASUREMENTS</smallCapsWord>
|
||
. Body length 6.50–6.91, width 1.62–1.97. head width 1.21, interocular width 0.52, eye length (dorsal view) 0.54; relative lengths of antennal segments I–IV: 3.11: 0.88: 0.98: 1.08; pronotum: length 0.70, width 1.35; mesonotum: length 1.87, width 1.62; metanotum: length 0.65, width 1.60; abdomen length (ventral view) 2.72; abdominal sternum VII: length 0.38, width 0.52; abdominal mediotergite I: length 0.22, width 0.57; relative lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II): fore leg: 3.01: 2.34: 0.78: 0.52, middle leg: 8.88: 4.65: 2.10: 0.38, hind leg: 8.56:2.97: 0.11: 0.15.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938A2FC2E727F0B59" blockId="18.[187,1399,924,1091]" pageId="18" pageNumber="19">
|
||
<smallCapsWord id="8AAEA047FFCE292938A2FC2E711B0CAE" baselines="943,943" box="[189,328,924,950]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="18" pageNumber="19">COLORATION</smallCapsWord>
|
||
. Median black spot of head reduced, indistinctly bifid or dissolved in smaller spots posteriorly (
|
||
<figureCitation id="14CC2A1EFFCE29293956FC0D71FA0CC3" box="[329,425,959,985]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="18" pageNumber="19">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCE292939ACFC0D71BA0CC3" box="[435,489,959,985]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">16H</figureCitation>
|
||
). Pronotum and mesonotum mainly black with a median brownish-yellow spot. Mesonotum and metanotum completely black, without yellow markings (
|
||
<figureCitation id="14CC2A1EFFCE29293C3FFC5074D00CE6" box="[1056,1155,994,1020]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="18" pageNumber="19">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCE29293C8EFC5074940CE6" box="[1169,1223,994,1020]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">16H</figureCitation>
|
||
). Connexivum dorsally blackish. Mesosternum chiefly blackish with a median subtriangular yellowish spot (
|
||
<figureCitation id="14CC2A1EFFCE29293D17FBB7753A0B05" box="[1288,1385,1029,1055]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Fig. 16I</figureCitation>
|
||
). Abdominal venter light yellow.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938A2FBD871610BBD" blockId="18.[189,1398,1130,1191]" pageId="18" pageNumber="19">
|
||
<smallCapsWord id="8AAEA047FFCE292938A2FBD8717F0B99" baselines="1149,1150" box="[189,300,1130,1156]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="18" pageNumber="19">ABDOMEN</smallCapsWord>
|
||
. Abdominal mediotergite I not swollen, nearly completely covered by silvery pubescence (
|
||
<figureCitation id="14CC2A1EFFCE292938D9FB3F71760BBD" box="[198,293,1165,1191]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="18" pageNumber="19">Fig. 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938A2FB7D71D40A16" blockId="18.[189,1399,1231,1292]" pageId="18" pageNumber="19">
|
||
<smallCapsWord id="8AAEA047FFCE292938A2FB7D70B80BFD" baselines="1250,1250" box="[189,235,1231,1257]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="18" pageNumber="19">LEG</smallCapsWord>
|
||
. Middle trochanter with 3–4 small spines; middle femur with a row of irregularly-spaced small spines (
|
||
<figureCitation id="14CC2A1EFFCE29293909FB4071290A16" box="[278,378,1266,1292]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Fig. 16J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938A2FA8171250AAD" blockId="18.[189,1399,1331,1463]" pageId="18" pageNumber="19">
|
||
<smallCapsWord id="8AAEA047FFCE292938A2FA8171620A56" baselines="1350,1351" box="[189,305,1331,1357]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="18" pageNumber="19">GENITALIA</smallCapsWord>
|
||
. Abdominal segment VIII ventro-laterally depressed. Pygophore large, ovate. Proctiger laterally with subtrapezoid lobes, dorsally with brownish setae (
|
||
<figureCitation id="14CC2A1EFFCE29293BD6FAE4747C0A6A" box="[969,1071,1366,1392]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="18" pageNumber="19">Figs 6A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCE29293C20FAE474260A6A" box="[1087,1141,1366,1392]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">16K</figureCitation>
|
||
). Paramere relatively slender, strongly curved basal third, distal part tapering towards distinctly curved and narrowed apex (
|
||
<figureCitation id="14CC2A1EFFCE292938D8FA2F717A0AAD" box="[199,297,1437,1463]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="18" pageNumber="19">Figs 8A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCE2929392AFA2F713A0AAD" box="[309,361,1437,1463]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">16L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFCE292938A2FA6C72200933" box="[189,627,1502,1577]" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="8C48369BFFCE292938A2FA6C71050AE2" blockId="18.[189,342,1502,1528]" box="[189,342,1502,1528]" pageId="18" pageNumber="19">
|
||
<heading id="D70081F7FFCE292938A2FA6C71050AE2" bold="true" box="[189,342,1502,1528]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
||
<emphasis id="BE83EA89FFCE292938A2FA6C71050AE2" bold="true" box="[189,342,1502,1528]" pageId="18" pageNumber="19">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292938A2F9BF72200933" blockId="18.[189,627,1549,1577]" box="[189,627,1549,1577]" pageId="18" pageNumber="19">
|
||
<collectingCountry id="F4E0760BFFCE292938A2F9BF715B093D" box="[189,264,1549,1575]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="4E33F879FFCE2929390CF9BF712F093D" box="[275,380,1549,1575]" country="China" name="Guangxi" pageId="18" pageNumber="19">Guangxi</collectingRegion>
|
||
.
|
||
<collectingCountry id="F4E0760BFFCE29293999F9BF71BD093D" box="[390,494,1549,1575]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
|
||
: Lạng SƠn.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFCE292E38A2F9FC75710D26" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="8C48369BFFCE292938A2F9FC71F90972" blockId="18.[189,426,1614,1640]" box="[189,426,1614,1640]" pageId="18" pageNumber="19">
|
||
<heading id="D70081F7FFCE292938A2F9FC71F90972" bold="true" box="[189,426,1614,1640]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
||
<emphasis id="BE83EA89FFCE292938A2F9FC71F90972" bold="true" box="[189,426,1614,1640]" pageId="18" pageNumber="19">Comparative notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFCE292E38A2F9CF75710D26" blockId="18.[187,1400,1661,2004]" lastBlockId="21.[188,1399,265,573]" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19">
|
||
This new species is closely related to
|
||
<taxonomicName id="4BF74D18FFCE29293A95F9CC734C098D" authority="Leng & Tran, 2023" authorityName="Leng & Tran" authorityYear="2023" box="[650,799,1661,1687]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
|
||
<emphasis id="BE83EA89FFCE29293A95F9CC734C098D" box="[650,799,1661,1687]" italics="true" pageId="18" pageNumber="19">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B057F2FFCE29293B36F9CF73D5098D" box="[809,902,1661,1687]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in having an elongate medial process on the posterior margin of mediotergite VII and five processes on abdominal segment VII of the female, including two connexival processes, two lateral blunt processes and a semicircular median process on the ventral side (
|
||
<figureCitation id="14CC2A1EFFCE2929399EF954725D081A" box="[385,526,1766,1792]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Figs 16D–F</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFCE29293A05F9557223081B" box="[538,624,1767,1793]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[335,1283,278,1755]" captionTargetId="figure-14@24.[238,1348,265,1779]" captionTargetPageId="24" captionText="Fig. 18. Morphological features of Rhyacobates elongatus sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385674" httpUri="https://zenodo.org/record/8385674/files/figure.png" pageId="18" pageNumber="19">18D–F</figureCitation>
|
||
). However, the abdominal mediotergite I of
|
||
<taxonomicName id="4BF74D18FFCE29293C6AF95574E8081A" authority="Leng & Tran, 2023" authorityName="Leng & Tran" authorityYear="2023" box="[1141,1211,1766,1792]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bui" status="sp. nov.">
|
||
<emphasis id="BE83EA89FFCE29293C6AF95574E8081A" box="[1141,1211,1766,1792]" italics="true" pageId="18" pageNumber="19">R. bui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B057F2FFCE29293CDCF955754E081B" box="[1219,1309,1767,1793]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is fully covered with silvery pubescence, which is much denser than in
|
||
<taxonomicName id="4BF74D18FFCE29293B8EF8B87471083E" authorityName="Leng & Tran" authorityYear="2023" box="[913,1058,1802,1828]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFCE29293B8EF8B87471083E" box="[913,1058,1802,1828]" italics="true" pageId="18" pageNumber="19">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="14CC2A1EFFCE29293C2DF8B874E2083E" box="[1074,1201,1802,1828]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="18" pageNumber="19">Fig. 3A–B</figureCitation>
|
||
). The connexival processes of abdominal segment VII of
|
||
<taxonomicName id="4BF74D18FFCE29293ABFF89C72B4085D" authorityName="Leng & Tran" authorityYear="2023" box="[672,743,1837,1863]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFCE29293ABFF89C72B4085D" box="[672,743,1837,1863]" italics="true" pageId="18" pageNumber="19">R. bui</emphasis>
|
||
</taxonomicName>
|
||
are much shorter and blunter (
|
||
<figureCitation id="14CC2A1EFFCE29293C41F89F7498085D" box="[1118,1227,1837,1863]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Fig. 16D</figureCitation>
|
||
) than those of
|
||
<taxonomicName id="4BF74D18FFCE292938A0F8E371010870" authorityName="Leng & Tran" authorityYear="2023" box="[191,338,1872,1898]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFCE292938A0F8E371010870" box="[191,338,1872,1898]" italics="true" pageId="18" pageNumber="19">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="14CC2A1EFFCE2929397BF8E271800870" box="[356,467,1872,1898]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[335,1283,278,1755]" captionTargetId="figure-14@24.[238,1348,265,1779]" captionTargetPageId="24" captionText="Fig. 18. Morphological features of Rhyacobates elongatus sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385674" httpUri="https://zenodo.org/record/8385674/files/figure.png" pageId="18" pageNumber="19">Fig. 18D</figureCitation>
|
||
). The abdomen of the female of
|
||
<taxonomicName id="4BF74D18FFCE29293B7EF8E373FA0870" authorityName="Leng & Tran" authorityYear="2023" box="[865,937,1872,1898]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFCE29293B7EF8E373FA0870" box="[865,937,1872,1898]" italics="true" pageId="18" pageNumber="19">R. bui</emphasis>
|
||
</taxonomicName>
|
||
is almost straight (
|
||
<figureCitation id="14CC2A1EFFCE29293C8BF8E274AC0870" box="[1172,1279,1872,1898]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="18" pageNumber="19">Fig. 16C</figureCitation>
|
||
), whereas that of
|
||
<taxonomicName id="4BF74D18FFCE2929390AF8C671FA0897" authorityName="Leng & Tran" authorityYear="2023" box="[277,425,1907,1933]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFCE2929390AF8C671FA0897" box="[277,425,1907,1933]" italics="true" pageId="18" pageNumber="19">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
is curved dorsad in the caudal part (
|
||
<figureCitation id="14CC2A1EFFCE29293B41F8C173980897" box="[862,971,1907,1933]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[335,1283,278,1755]" captionTargetId="figure-14@24.[238,1348,265,1779]" captionTargetPageId="24" captionText="Fig. 18. Morphological features of Rhyacobates elongatus sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385674" httpUri="https://zenodo.org/record/8385674/files/figure.png" pageId="18" pageNumber="19">Fig. 18C</figureCitation>
|
||
). These two species are also related to five species (i.e.,
|
||
<taxonomicName id="4BF74D18FFCE292939B8F825727C08AA" authorityName="Tran & Nguyen" authorityYear="2016" box="[423,559,1943,1968]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="angustus">
|
||
<emphasis id="BE83EA89FFCE292939B8F825727C08AA" box="[423,559,1943,1968]" italics="true" pageId="18" pageNumber="19">R. angustus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4BF74D18FFCE29293A21F825729C08AA" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[574,719,1942,1968]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lundbladi">
|
||
<emphasis id="BE83EA89FFCE29293A21F825729C08AA" box="[574,719,1942,1968]" italics="true" pageId="18" pageNumber="19">R. lundbladi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4BF74D18FFCE29293AC1F825732808AA" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[734,891,1942,1968]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="svenhedini">
|
||
<emphasis id="BE83EA89FFCE29293AC1F825732808AA" box="[734,891,1942,1968]" italics="true" pageId="18" pageNumber="19">R. svenhedini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4BF74D18FFCE29293B94F825747608AA" authorityName="Esaki" authorityYear="1923" box="[907,1061,1942,1968]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="takahashii">
|
||
<emphasis id="BE83EA89FFCE29293B94F825747608AA" box="[907,1061,1942,1968]" italics="true" pageId="18" pageNumber="19">R. takahashii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4BF74D18FFCE29293C79F825749F08AA" authorityName="Tran & Nguyen" authorityYear="2016" box="[1126,1228,1942,1968]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="zetteli">
|
||
<emphasis id="BE83EA89FFCE29293C79F825749F08AA" box="[1126,1228,1942,1968]" italics="true" pageId="18" pageNumber="19">R. zetteli</emphasis>
|
||
</taxonomicName>
|
||
) in having the median process on the posterior margin of mediotergite VII in the female, but this structure in
|
||
<taxonomicName id="4BF74D18FFCE29293D1FF808751708CE" authorityName="Leng & Tran" authorityYear="2023" box="[1280,1348,1978,2004]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFCE29293D1FF808751708CE" box="[1280,1348,1978,2004]" italics="true" pageId="18" pageNumber="19">R. bui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFC9292E38A2FEB8711D0E39" authorityName="Leng & Tran" authorityYear="2023" box="[189,334,265,291]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFC9292E38A2FEB8711D0E39" box="[189,334,265,291]" italics="true" pageId="21" pageNumber="22">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
is much longer than that of the five latter species. Furthermore, both
|
||
<taxonomicName id="4BF74D18FFC9292E3C75FEB874FD0E39" authorityName="Leng & Tran" authorityYear="2023" box="[1130,1198,265,291]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFC9292E3C75FEB874FD0E39" box="[1130,1198,265,291]" italics="true" pageId="21" pageNumber="22">R. bui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFC9292E3CFBFEB875260E39" authorityName="Leng & Tran" authorityYear="2023" box="[1252,1397,265,291]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFC9292E3CFBFEB875260E39" box="[1252,1397,265,291]" italics="true" pageId="21" pageNumber="22">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
have entirely black mesonota and metanota (
|
||
<figureCitation id="14CC2A1EFFC9292E3AD7FE9E731F0E5C" box="[712,844,300,326]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="21" pageNumber="22">Figs 3A–B</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC9292E3B47FE9E73F10E5C" box="[856,930,300,326]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="21" pageNumber="22">4A–B</figureCitation>
|
||
), without yellow stripes (in both sexes), which is a character state shared with
|
||
<taxonomicName id="4BF74D18FFC9292E3A9CFEE273770E73" authorityName="Tran & Nguyen" authorityYear="2016" box="[643,804,336,361]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC9292E3A9CFEE273770E73" box="[643,804,336,361]" italics="true" pageId="21" pageNumber="22">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
. Females of
|
||
<taxonomicName id="4BF74D18FFC9292E3BDFFEE274550E73" authorityName="Leng & Tran" authorityYear="2023" box="[960,1030,335,361]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFC9292E3BDFFEE274550E73" box="[960,1030,335,361]" italics="true" pageId="21" pageNumber="22">R. bui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFC9292E3C5DFEE274860E73" authorityName="Leng & Tran" authorityYear="2023" box="[1090,1237,335,361]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFC9292E3C5DFEE274860E73" box="[1090,1237,335,361]" italics="true" pageId="21" pageNumber="22">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
can be easily separated from
|
||
<taxonomicName id="4BF74D18FFC9292E396BFEC172450E96" authorityName="Tran & Nguyen" authorityYear="2016" box="[372,534,371,396]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC9292E396BFEC172450E96" box="[372,534,371,396]" italics="true" pageId="21" pageNumber="22">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
by the following characteristics: (1)
|
||
<taxonomicName id="4BF74D18FFC9292E3BD6FEC1745C0E96" authorityName="Leng & Tran" authorityYear="2023" box="[969,1039,370,396]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFC9292E3BD6FEC1745C0E96" box="[969,1039,370,396]" italics="true" pageId="21" pageNumber="22">R. bui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFC9292E3C56FEC174880E96" authorityName="Leng & Tran" authorityYear="2023" box="[1097,1243,370,396]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFC9292E3C56FEC174880E96" box="[1097,1243,370,396]" italics="true" pageId="21" pageNumber="22">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
have slender bodies (
|
||
<figureCitation id="14CC2A1EFFC9292E3905FE2471C80EAA" box="[282,411,406,432]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="21" pageNumber="22">Fig. 3A–B</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="4BF74D18FFC9292E3A08FE2572EA0EAA" authorityName="Tran & Nguyen" authorityYear="2016" box="[535,697,407,432]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC9292E3A08FE2572EA0EAA" box="[535,697,407,432]" italics="true" pageId="21" pageNumber="22">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
has a wider, stouter body (
|
||
<figureCitation id="14CC2A1EFFC9292E3BE9FE24741B0EAA" box="[1014,1096,406,432]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="21" pageNumber="22">Fig. 3I</figureCitation>
|
||
); (2) the posterior margin of mediotergite VII of
|
||
<taxonomicName id="4BF74D18FFC9292E39DDFE0872550EC9" authorityName="Leng & Tran" authorityYear="2023" box="[450,518,441,467]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFC9292E39DDFE0872550EC9" box="[450,518,441,467]" italics="true" pageId="21" pageNumber="22">R. bui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFC9292E3A21FE08729C0EC9" authorityName="Leng & Tran" authorityYear="2023" box="[574,719,441,467]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFC9292E3A21FE08729C0EC9" box="[574,719,441,467]" italics="true" pageId="21" pageNumber="22">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
has a median process (
|
||
<figureCitation id="14CC2A1EFFC9292E3BC5FE0B743E0ECE" box="[986,1133,441,468]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="21" pageNumber="22">Figs 16A, D</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC9292E3C68FE0B74830EC9" box="[1143,1232,441,467]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[335,1283,278,1755]" captionTargetId="figure-14@24.[238,1348,265,1779]" captionTargetPageId="24" captionText="Fig. 18. Morphological features of Rhyacobates elongatus sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385674" httpUri="https://zenodo.org/record/8385674/files/figure.png" pageId="21" pageNumber="22">18A, D</figureCitation>
|
||
), whereas that of
|
||
<taxonomicName id="4BF74D18FFC9292E38C2FE6F712C0EEC" authorityName="Tran & Nguyen" authorityYear="2016" box="[221,383,477,502]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC9292E38C2FE6F712C0EEC" box="[221,383,477,502]" italics="true" pageId="21" pageNumber="22">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
lacks a median process (
|
||
<figureCitation id="14CC2A1EFFC9292E3ABBFE6E72A90EEC" box="[676,762,476,502]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="21" pageNumber="22">Figs 3I</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC9292E3B18FE6E736F0EEC" box="[775,828,476,502]" captionStart="Fig" captionStartId="42.[189,232,1928,1954]" captionTargetBox="[217,1371,266,1884]" captionTargetId="figure-15@42.[217,1371,266,1884]" captionTargetPageId="42" captionText="Fig. 26.Photographs of Rhyacobates constrictus Tran & Nguyen, 2016. A–B. Holotype, apterous female (ZVNU). A. Body, lateral view. B. Habitus, dorsal view. C. Allotype, apterous male (ZVNU), habitus, dorsal view. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385692" httpUri="https://zenodo.org/record/8385692/files/figure.png" pageId="21" pageNumber="22">26B</figureCitation>
|
||
); (3) the posterior margin of abdominal segment VII of
|
||
<taxonomicName id="4BF74D18FFC9292E390FFDB271050D03" authorityName="Leng & Tran" authorityYear="2023" box="[272,342,511,537]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bui">
|
||
<emphasis id="BE83EA89FFC9292E390FFDB271050D03" box="[272,342,511,537]" italics="true" pageId="21" pageNumber="22">R. bui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFC9292E398EFDB272760D03" authorityName="Leng & Tran" authorityYear="2023" box="[401,549,511,537]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="BE83EA89FFC9292E398EFDB272760D03" box="[401,549,511,537]" italics="true" pageId="21" pageNumber="22">R. elongatus</emphasis>
|
||
</taxonomicName>
|
||
has five processes (
|
||
<figureCitation id="14CC2A1EFFC9292E3B0EFE4D73CC0D03" box="[785,927,511,537]" captionStart="Fig" captionStartId="20.[189,232,1787,1813]" captionTargetBox="[280,1309,276,1715]" captionTargetId="figure-14@20.[245,1342,265,1743]" captionTargetPageId="20" captionText="Fig. 16. Morphological features of Rhyacobates bui sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, distal view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385670" httpUri="https://zenodo.org/record/8385670/files/figure.png" pageId="21" pageNumber="22">Figs 16D–F</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC9292E3BB3FDB274520D00" box="[940,1025,512,538]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[335,1283,278,1755]" captionTargetId="figure-14@24.[238,1348,265,1779]" captionTargetPageId="24" captionText="Fig. 18. Morphological features of Rhyacobates elongatus sp. nov. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385674" httpUri="https://zenodo.org/record/8385674/files/figure.png" pageId="21" pageNumber="22">18D–F</figureCitation>
|
||
), whereas that of
|
||
<taxonomicName id="4BF74D18FFC9292E3CCBFDB275240D03" authorityName="Tran & Nguyen" authorityYear="2016" box="[1236,1399,512,537]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC9292E3CCBFDB275240D03" box="[1236,1399,512,537]" italics="true" pageId="21" pageNumber="22">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
has only one median process ventrally (
|
||
<figureCitation id="14CC2A1EFFC9292E3A91FD90734E0D26" box="[654,797,546,572]" captionStart="Fig" captionStartId="42.[189,232,1928,1954]" captionTargetBox="[217,1371,266,1884]" captionTargetId="figure-15@42.[217,1371,266,1884]" captionTargetPageId="42" captionText="Fig. 26.Photographs of Rhyacobates constrictus Tran & Nguyen, 2016. A–B. Holotype, apterous female (ZVNU). A. Body, lateral view. B. Habitus, dorsal view. C. Allotype, apterous male (ZVNU), habitus, dorsal view. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385692" httpUri="https://zenodo.org/record/8385692/files/figure.png" pageId="21" pageNumber="22">Fig. 26A–B</figureCitation>
|
||
; also see
|
||
<bibRefCitation id="E8664B6AFFC9292E3B93FD9174D80D26" author="Tran & Nguyen" box="[908,1163,546,573]" firstAuthor="Tran" pageId="21" pageNumber="22" pagination="501 - 516" refId="ref33477" refString="Tran A. D. & Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran & Nguyen 2016</bibRefCitation>
|
||
: figs 36–39).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |