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<document id="EF84137E7A98200A7C0E686ABB50D0D0" ID-DOI="10.15298/rusentj.31.1.17" ID-ISSN="0132-8069" ID-Zenodo-Dep="13177702" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722592858916" checkinUser="felipe" docAuthor="MacGowan, I. &amp; Ruchin, A. B." docDate="2022" docId="03E887E9615EFFB3596AFB160944FCC4" docLanguage="en" docName="RussEntomolJ.31.1.83-86.pdf" docOrigin="Russian Entomological Journal 31 (1)" docSource="http://dx.doi.org/10.15298/rusentj.31.1.17" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Lonchaea cryptica MacGowan et Ruchin 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="86" masterDocId="FFD1FF91615FFFB05A38FFEC0851FFA3" masterDocTitle="Two new species of Lonchaeidae (Diptera: Schizophora) from the Republic of Mordovia, Russia" masterLastPageNumber="86" masterPageNumber="83" pageNumber="84" updateTime="1722632040337" updateUser="ExternalLinkService">
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<mods:title id="0EDD7FD300828BED9C2480B08F85077C">Two new species of Lonchaeidae (Diptera: Schizophora) from the Republic of Mordovia, Russia</mods:title>
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MacGowan et Ruchin
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<figureCitation id="137A2A7A615EFFB15E27FAFB0CD1FA8C" box="[1055,1152,1303,1327]" captionStart="Figs 59" captionStartId="2.[226,263,1969,1988]" captionTargetBox="[197,1362,1192,1942]" captionTargetId="figure-672@2.[191,1391,1192,1939]" captionTargetPageId="2" captionText="Figs 59. Lonchaea cryptica sp.n.: 5 — epandrium and associated structures; 6 — inner surface of the surstylus; 7 — phallus; 89 — detail of apex of phallus; 5, 78 — lateral view; 9 — ventral view. Рис. 59. Lonchaea cryptica sp.n.: 5 — Эпандрий и прилежаЩие структуры; 6 — внутреннЯЯ поверхность сурстилей; 7 — фаллус; 89 — детали строениЯ верШины фаллуса; 5, 78 — сбоку; 9 — сниЗу." figureDoi="http://doi.org/10.5281/zenodo.13177708" httpUri="https://zenodo.org/record/13177708/files/figure.png" pageId="1" pageNumber="84">Figs 59</figureCitation>
.
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DESCRIPTION.
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,
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. Head: Eyes bare. Frons at narrowest point above lunule approximately 0.6x width of an eye, matt black, frontal and interfrontal setulae long, 0.6x length of orbital seta. Orbital plate black, dulled by microsculpture, bare apart from the orbital seta. Lunule ground colour black; face sub-shining black, parafacials slightly grey dusted. Anterior genal setulae forming a single regularly spaced row of 56 along mouth edge, these slightly stronger than other setulae on the genae. Palpi black, with numerous setae. Antennae entirely black, antennal postpedicel long, reaching mouth margin, length to depth ratio 3.0: 1. Arista entirely black.
</paragraph>
<caption id="DF3E6677615EFFB15ADAF86D0ADBF7B2" ID-DOI="http://doi.org/10.5281/zenodo.13177704" ID-Zenodo-Dep="13177704" httpUri="https://zenodo.org/record/13177704/files/figure.png" pageId="1" pageNumber="84" startId="1.[226,264,1921,1940]" targetBox="[194,769,997,1885]" targetPageId="1" targetType="figure">
<paragraph id="8BFE36FF615EFFB15ADAF86D0A66F867" blockId="1.[189,775,1921,2065]" pageId="1" pageNumber="84">
Figs 14.
<taxonomicName id="4C414D7C615EFFB15B79F86E09A4F837" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" box="[321,501,1921,1940]" class="Insecta" family="Lonchaeidae" genus="Earomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="mordovia" status="sp. nov.">
<emphasis id="B935EAED615EFFB15B79F86E09A4F837" box="[321,501,1921,1940]" italics="true" pageId="1" pageNumber="84">Earomyia mordovia</emphasis>
</taxonomicName>
<emphasis id="B935EAED615EFFB15BC6F86E0A7BF836" bold="true" box="[510,554,1922,1941]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="A2065796615EFFB15BC6F86E0A7BF836" box="[510,554,1922,1941]" pageId="1" pageNumber="84" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 12 — epandrium and associated structures; 3 — phallus; 4 — inner surface of the surstylus; 1, 3 — lateral view; 2 — ventral view.
</paragraph>
<paragraph id="8BFE36FF615EFFB2596AF87009CFFCC4" blockId="1.[812,1400,1237,2077]" lastBlockId="2.[189,776,315,1137]" lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84">
Thorax: Scutum sub shining blue-black, light grey dusted with a covering of rather dense long black setulae approximately 0.75x length of orbital seta. Humeri densely covered in setulae. Anepisternum grey dusted with 4 setae in a row along posterior margin, anterior setae difficult to distinguish from other numerous long setulae on the sclerite. Proepisternum with one seta on left, two on right, proepimeron with two setae, one weaker than the other on left, one on right. Katepisternum, sub-shining black, a single seta located near dorsal margin, remainder of sclerite, apart from posterior portion, covered in long setulae the majority of which are more than 0.5x the length of the seta, several setulae present posterior to the seta. Anepimeron bare. Scutellum sub-shining black, on margin with a pair of lateral and apical setae, six to seven setulae situated between lateral and apical setae, six between apical setae, these at least 0.5x as long as marginal setae, no setulae anterior to the lateral setae. Calypteres dark grey with a black margin and a dense, long black fringe of uniform length.Wing: length
<quantity id="4CB99B1A615DFFB25BEDFD7A0A4EFD08" box="[469,543,662,683]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="85" unit="mm" value="3.9">3.9 mm</quantity>
, veins yellowish-brown, intercostal section of wing, between insertion of Sc and R1, approximately 4x length of cross vein rm. Legs, black, basal tarsomere of fore legs and basal and second tarsomeres of hind legs yellow-brown ventrally with a ventral fringe of short, stiff golden setulae. Basal tarsomere of mid leg only very obscurely brownish at extreme ventral base, ventral fringe of setulae black.
</paragraph>
<paragraph id="8BFE36FF615EFFB15ADAF8220ADBF7B2" blockId="1.[189,775,1921,2065]" pageId="1" pageNumber="84">
Рис. 14.
<taxonomicName id="4C414D7C615EFFB15B70F82209AEF843" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" box="[328,511,1997,2016]" class="Insecta" family="Lonchaeidae" genus="Earomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="mordovia" status="sp. nov.">
<emphasis id="B935EAED615EFFB15B70F82209AEF843" box="[328,511,1997,2016]" italics="true" pageId="1" pageNumber="84">Earomyia mordovia</emphasis>
</taxonomicName>
<emphasis id="B935EAED615EFFB15833F8210A66F843" bold="true" box="[523,567,1997,2016]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="A2065796615EFFB15833F8210A66F843" box="[523,567,1997,2016]" pageId="1" pageNumber="84" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 12 — Эпандрий и прилежаЩие структуры; 3 — фаллус; 4 — внутреннЯЯ поверхность сурстилей; 1, 3 — сбоку; 2 — сниЗу.
</paragraph>
</caption>
<paragraph id="8BFE36FF615DFFB25ADAFC870B0CFE02" blockId="2.[189,776,315,1137]" lastBlockId="2.[812,1399,315,1137]" pageId="2" pageNumber="85">
Abdomen: 1
<superScript id="7C349BB7615DFFB25B66FC870936FCD4" attach="left" box="[350,359,875,887]" fontSize="5" pageId="2" pageNumber="85">st</superScript>
sternite without setulae. Male terminalia: (
<figureCitation id="137A2A7A615DFFB25AFBFC6A0970FC3F" box="[195,289,902,924]" captionStart="Figs 59" captionStartId="2.[226,263,1969,1988]" captionTargetBox="[197,1362,1192,1942]" captionTargetId="figure-672@2.[191,1391,1192,1939]" captionTargetPageId="2" captionText="Figs 59. Lonchaea cryptica sp.n.: 5 — epandrium and associated structures; 6 — inner surface of the surstylus; 7 — phallus; 89 — detail of apex of phallus; 5, 78 — lateral view; 9 — ventral view. Рис. 59. Lonchaea cryptica sp.n.: 5 — Эпандрий и прилежаЩие структуры; 6 — внутреннЯЯ поверхность сурстилей; 7 — фаллус; 89 — детали строениЯ верШины фаллуса; 5, 78 — сбоку; 9 — сниЗу." figureDoi="http://doi.org/10.5281/zenodo.13177708" httpUri="https://zenodo.org/record/13177708/files/figure.png" pageId="2" pageNumber="85">Figs 59</figureCitation>
): In lateral view epandrium slightly wider than high, with several strong setae on posterior and ventral margins. Cerci large and apically rounded, more than half height of epandrium; bearing numerous strong setae apically. Surstylus extending ventrally beyond shell of epandrium for all its length culminating in a large, rounded, slightly hooked, posterior process. Inner surface of surstylus with numerous strong black setae on ventral half, posterior process bare apart from a few setulae. Phallus un-segmented, a simple-J shape, base and stem darkened with sclerotization, apically divided into two equal processes, these lying at an angle to the main stem and serrated along their outer margins.
</paragraph>
<paragraph id="8BFE36FF615DFFB2596AFE4A0D27FE52" blockId="2.[812,1399,315,1137]" pageId="2" pageNumber="85">
Female: Associated females are similar to the males apart from a wider frons and generally shorter pilosity. However, at present they are not distinguishable from females of
<taxonomicName id="4C414D7C615DFFB25F24FE300D21FE53" authorityName="Malloch" authorityYear="1920" box="[1308,1392,475,497]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615DFFB25F24FE300D21FE53" box="[1308,1392,475,497]" italics="true" pageId="2" pageNumber="85">L. affinis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B6574615DFFB2596AFE1A0CC0FDE2" pageId="2" pageNumber="85" type="etymology">
<paragraph id="8BFE36FF615DFFB2596AFE1A0CC0FDE2" blockId="2.[812,1399,315,1137]" pageId="2" pageNumber="85">
ETYMOLOGY. The specific epithet refers to the fact that this species has until now remained undetected within series of the very similar
<taxonomicName id="4C414D7C615DFFB259D0FDC70CDBFDE3" authorityName="Malloch" authorityYear="1920" box="[1000,1162,555,576]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615DFFB259D0FDC70CDBFDE3" box="[1000,1162,555,576]" italics="true" pageId="2" pageNumber="85">Lonchaea affinis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B6574615DFFB3596AFDAA09D0FCB4" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85" type="discussion">
<paragraph id="8BFE36FF615DFFB2596AFDAA0BD7FCEF" blockId="2.[812,1399,315,1137]" pageId="2" pageNumber="85">
REMARKS. With bare eyes, anterior genal setae in a single row, tarsomeres partly pale, calypteres dark fringed, scutellar disc bare and proepimeron, in most cases with more than one seta, this species belongs to the
<taxonomicName id="4C414D7C615DFFB25EFAFD7B0B0DFD65" authority="Malloch, 1920" authorityName="Malloch" authorityYear="1920" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615DFFB25EFAFD7B0C9EFD0F" box="[1218,1231,663,684]" italics="true" pageId="2" pageNumber="85">L</emphasis>
.
<emphasis id="B935EAED615DFFB25EE4FD7A0D49FD08" box="[1244,1304,662,683]" italics="true" pageId="2" pageNumber="85">affinis</emphasis>
Malloch, 1920
</taxonomicName>
species-group within the Palearctic
<taxonomicName id="4C414D7C615DFFB25E82FD5D0D48FD65" authorityName="Fallen" authorityYear="1820" box="[1210,1305,689,710]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAED615DFFB25E82FD5D0D48FD65" box="[1210,1305,689,710]" italics="true" pageId="2" pageNumber="85">Lonchaea</emphasis>
</taxonomicName>
. This is a small group consisting of the widespread Holarctic
<taxonomicName id="4C414D7C615DFFB25F18FD200D27FD43" authorityName="Malloch" authorityYear="1920" box="[1312,1398,715,737]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615DFFB25F18FD200D27FD43" box="[1312,1398,715,737]" italics="true" pageId="2" pageNumber="85">L. affinis</emphasis>
</taxonomicName>
and the little-known
<taxonomicName id="4C414D7C615DFFB259CCFD0B0D55FD5F" authority="Hackman, 1956" authorityName="Hackman" authorityYear="1956" box="[1012,1284,742,764]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="sorocula">
<emphasis id="B935EAED615DFFB259CCFD0B0C32FD58" box="[1012,1123,742,764]" italics="true" pageId="2" pageNumber="85">L. sorocula</emphasis>
Hackman, 1956
</taxonomicName>
. Two of the
<typeStatus id="54FA885D615DFFB25914FCED0BD8FCB5" box="[812,905,769,790]" pageId="2" pageNumber="85" type="paratype">paratypes</typeStatus>
also have two setae on the proepisternum, the number of setulae on the proepimeron varies between one and four.
</paragraph>
<paragraph id="8BFE36FF615DFFB2596AFCBD0C99FBA4" blockId="2.[812,1399,315,1137]" pageId="2" pageNumber="85">
In external characters
<taxonomicName id="4C414D7C615DFFB25E12FCBD0CC2FCC5" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" box="[1066,1171,849,870]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="cryptica" status="sp.n.">
<emphasis id="B935EAED615DFFB25E12FCBD0C66FCC5" box="[1066,1079,849,870]" italics="true" pageId="2" pageNumber="85">L</emphasis>
.
<emphasis id="B935EAED615DFFB25E7CFCBD0CC2FCC5" box="[1092,1171,849,870]" italics="true" pageId="2" pageNumber="85">cryptica</emphasis>
</taxonomicName>
<emphasis id="B935EAED615DFFB25EA2FCBD0C9AFCC5" bold="true" box="[1178,1227,849,870]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A2065796615DFFB25EA2FCBD0C9AFCC5" box="[1178,1227,849,870]" pageId="2" pageNumber="85" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="4C414D7C615DFFB25914FC800BD3FC23" authorityName="Malloch" authorityYear="1920" box="[812,898,875,897]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615DFFB25914FC800BD3FC23" box="[812,898,875,897]" italics="true" pageId="2" pageNumber="85">L. affinis</emphasis>
</taxonomicName>
, the general shape and structures of the male terminalia are similar apart from the phallus which is clearly distinct. In terms of chaetotaxy
<taxonomicName id="4C414D7C615DFFB25E60FC4D0CEEFC15" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" box="[1112,1215,929,950]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="cryptica" status="sp.n.">
<emphasis id="B935EAED615DFFB25E60FC4D0CEEFC15" box="[1112,1215,929,950]" italics="true" pageId="2" pageNumber="85">L. cryptica</emphasis>
</taxonomicName>
<emphasis id="B935EAED615DFFB25EFCFC4D0CA4FC15" bold="true" box="[1220,1269,929,950]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A2065796615DFFB25EFCFC4D0CA4FC15" box="[1220,1269,929,950]" pageId="2" pageNumber="85" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
tends to have fewer setae on the proepimeron with numbers in the
<typeStatus id="54FA885D615DFFB25F7DFC500B0EFC48" pageId="2" pageNumber="85" type="paratype">paratypes</typeStatus>
ranging from one to four whereas in
<taxonomicName id="4C414D7C615DFFB25EFDFC3B0D4BFC48" authorityName="Malloch" authorityYear="1920" box="[1221,1306,982,1004]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615DFFB25EFDFC3B0D4BFC48" box="[1221,1306,982,1004]" italics="true" pageId="2" pageNumber="85">L. affinis</emphasis>
</taxonomicName>
there is a tendency for there to be a greater number.
</paragraph>
<paragraph id="8BFE36FF615DFFB3596AFBE00A54FD0F" blockId="2.[812,1399,315,1137]" lastBlockId="3.[189,775,315,684]" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85">
In the past there was confusion between
<taxonomicName id="4C414D7C615DFFB25EE5FBE00D62FB83" authorityName="Malloch" authorityYear="1920" box="[1245,1331,1035,1057]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615DFFB25EE5FBE00D62FB83" box="[1245,1331,1035,1057]" italics="true" pageId="2" pageNumber="85">L. affinis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C414D7C615DFFB25F5BFBE00B88FB9F" authority="Collin, 1953" authorityName="Collin" authorityYear="1953" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="laxa">
<emphasis id="B935EAED615DFFB25F5BFBE00D21FB82" box="[1379,1392,1036,1057]" italics="true" pageId="2" pageNumber="85">L</emphasis>
.
<emphasis id="B935EAED615DFFB25914FBCA0B04FB98" box="[812,853,1062,1083]" italics="true" pageId="2" pageNumber="85">laxa</emphasis>
Collin, 1953
</taxonomicName>
. The type series of
<taxonomicName id="4C414D7C615DFFB25EA1FBCB0C8CFB98" authorityName="Collin" authorityYear="1953" box="[1177,1245,1062,1084]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="laxa">
<emphasis id="B935EAED615DFFB25EA1FBCB0C8CFB98" box="[1177,1245,1062,1084]" italics="true" pageId="2" pageNumber="85">L. laxa</emphasis>
</taxonomicName>
consists of
<specimenCount id="9D47FD76615DFFB25F6AFBCA0B0AFBF4" count="2" pageId="2" pageNumber="85" type="male">two male</specimenCount>
and
<specimenCount id="9D47FD76615DFFB259B3FBAD0C5BFBF4" box="[907,1034,1089,1111]" count="7" pageId="2" pageNumber="85" type="female">seven female</specimenCount>
<typeStatus id="54FA885D615DFFB25E28FBAD0C34FBF5" box="[1040,1125,1089,1110]" pageId="2" pageNumber="85" type="syntype">syntypes</typeStatus>
[
<bibRefCitation id="EFD04B0E615DFFB25E48FBAD0C8EFBF5" author="Pont A. C." box="[1136,1247,1088,1110]" pageId="2" pageNumber="85" refId="ref3117" refString="Pont A. C. 1995. The Type-Material of Diptera (Insecta) Described by G. H. Verrall and J. E. Collin. Oxford, United Kingdom: Oxford Scientific Publications. 223 pp." type="book" year="1995">Pont, 1995</bibRefCitation>
], all except
<specimenCount id="9D47FD76615DFFB25F6BFBAD0B0AFBD2" count="1" pageId="2" pageNumber="85" type="male">one male</specimenCount>
were from the Scottish Highlands. Examination of the male from
<collectingCountry id="F356766F615CFFB35B1FFED7092AFEF2" box="[295,379,315,337]" name="United Kingdom" pageId="3" pageNumber="86">Scotland</collectingCountry>
confirmed that it was in fact a specimen of
<taxonomicName id="4C414D7C615CFFB35AE5FEBB096BFEC8" authorityName="Malloch" authorityYear="1920" box="[221,314,342,364]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615CFFB35AE5FEBB096BFEC8" box="[221,314,342,364]" italics="true" pageId="3" pageNumber="86">L. affinis</emphasis>
</taxonomicName>
and it was considered most likely that the associated females were also of this species. As a result,
<taxonomicName id="4C414D7C615CFFB358CBFE9D08B7FE03" authorityName="Collin" authorityYear="1953" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="laxa">
<emphasis id="B935EAED615CFFB358CBFE9D08B7FE03" italics="true" pageId="3" pageNumber="86">L. laxa</emphasis>
</taxonomicName>
was proposed as a junior synonym of
<taxonomicName id="4C414D7C615CFFB3587EFE600AC8FE03" authorityName="Malloch" authorityYear="1920" box="[582,665,395,417]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B935EAED615CFFB3587EFE600AC8FE03" box="[582,665,395,417]" italics="true" pageId="3" pageNumber="86">L. affinis</emphasis>
</taxonomicName>
[MacGow- an, 2020]. However, at the time of publication of the
<taxonomicName id="4C414D7C615CFFB358E0FE4A0940FE74" class="Insecta" family="Lonchaeidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="family">Lonchaeidae</taxonomicName>
volume of the “
<emphasis id="B935EAED615CFFB35B88FE2D0918FE52" italics="true" pageId="3" pageNumber="86">Handbooks for the Identification of British Insects</emphasis>
<taxonomicName id="4C414D7C615CFFB35B64FE3009F3FE53" authorityName="Collin" authorityYear="1953" box="[348,418,475,497]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="laxa">
<emphasis id="B935EAED615CFFB35B64FE3009F3FE53" box="[348,418,475,497]" italics="true" pageId="3" pageNumber="86">L. laxa</emphasis>
</taxonomicName>
was still considered a separate species, and the figure of the male terminalia was based on the second
<typeStatus id="54FA885D615CFFB35B30FDFD0902FD85" box="[264,339,529,550]" pageId="3" pageNumber="86" type="syntype">syntype</typeStatus>
male which was captured in southern
<collectingCountry id="F356766F615CFFB358E0FDFD08B6FDE2" name="United Kingdom" pageId="3" pageNumber="86">England</collectingCountry>
[
<bibRefCitation id="EFD04B0E615CFFB35ACCFDC70A40FDE3" author="MacGowan I. &amp; Rotheray G. E." box="[244,529,555,577]" pageId="3" pageNumber="86" refId="ref2933" refString="MacGowan I., Rotheray G. E. 2008. British Lonchaeidae. Diptera, Cyclorrhapha, Acalyptratae // Handbooks for the Identification of British Insects. Vol. 10. No. 15. London: Royal Entomological Society. 159 pp." type="book" year="2008">MacGowan, Rotheray, 2008</bibRefCitation>
: Fig. 242 on p.63]. That figure shows that the specimen was in fact a male of
<taxonomicName id="4C414D7C615CFFB358CBFDAB095DFDD5" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cryptica" status="sp.n.">
<emphasis id="B935EAED615CFFB358CBFDAB095DFDD5" italics="true" pageId="3" pageNumber="86">L. cryptica</emphasis>
</taxonomicName>
<emphasis id="B935EAED615CFFB35B2AFD8D0913FDD5" bold="true" box="[274,322,609,630]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="A2065796615CFFB35B2AFD8D0913FDD5" box="[274,322,609,630]" pageId="3" pageNumber="86" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, unfortunately the specimen has now been lost and at present it is not possible to confirm the presence of
<taxonomicName id="4C414D7C615CFFB358CBFD90095DFD0F" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cryptica" status="sp.n.">
<emphasis id="B935EAED615CFFB358CBFD90095DFD0F" italics="true" pageId="3" pageNumber="86">L. cryptica</emphasis>
</taxonomicName>
<emphasis id="B935EAED615CFFB35B2BFD7A0912FD08" bold="true" box="[275,323,662,683]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="A2065796615CFFB35B2BFD7A0912FD08" box="[275,323,662,683]" pageId="3" pageNumber="86" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
in the British Isles.
</paragraph>
<caption id="DF3E6677615DFFB25ADAF85D0B01F7B2" ID-DOI="http://doi.org/10.5281/zenodo.13177708" ID-Zenodo-Dep="13177708" httpUri="https://zenodo.org/record/13177708/files/figure.png" pageId="2" pageNumber="85" startId="2.[226,263,1969,1988]" targetBox="[197,1362,1192,1942]" targetPageId="2" targetType="figure">
<paragraph id="8BFE36FF615DFFB25ADAF85D0B59F87F" blockId="2.[189,1398,1969,2065]" pageId="2" pageNumber="85">
Figs 59.
<taxonomicName id="4C414D7C615DFFB25B02F85D098AF867" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" box="[314,475,1969,1988]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="cryptica" status="sp. nov.">
<emphasis id="B935EAED615DFFB25B02F85D098AF867" box="[314,475,1969,1988]" italics="true" pageId="2" pageNumber="85">Lonchaea cryptica</emphasis>
</taxonomicName>
<emphasis id="B935EAED615DFFB25BDAF85D0A5CF867" bold="true" box="[482,525,1969,1988]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A2065796615DFFB25BDAF85D0A5CF867" box="[482,525,1969,1988]" pageId="2" pageNumber="85" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 5 — epandrium and associated structures; 6 — inner surface of the surstylus; 7 — phallus; 89 — detail of apex of phallus; 5, 78 — lateral view; 9 — ventral view.
</paragraph>
<paragraph id="8BFE36FF615DFFB25ADAF80A0B01F7B2" blockId="2.[189,1398,1969,2065]" pageId="2" pageNumber="85">
Рис. 59.
<taxonomicName id="4C414D7C615DFFB25B01F8090989F85B" authorityName="MacGowan &amp; Ruchin" authorityYear="2022" box="[313,472,2021,2040]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="cryptica" status="sp. nov.">
<emphasis id="B935EAED615DFFB25B01F8090989F85B" box="[313,472,2021,2040]" italics="true" pageId="2" pageNumber="85">Lonchaea cryptica</emphasis>
</taxonomicName>
<emphasis id="B935EAED615DFFB25BE6F8090A58F85B" bold="true" box="[478,521,2021,2040]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A2065796615DFFB25BE6F8090A58F85B" box="[478,521,2021,2040]" pageId="2" pageNumber="85" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
: 5 — Эпандрий и прилежаЩие структуры; 6 — внутреннЯЯ поверхность сурстилей; 7 — фаллус; 89 — детали строениЯ верШины фаллуса; 5, 78 — сбоку; 9 — сниЗу.
</paragraph>
</caption>
<paragraph id="8BFE36FF615CFFB35ADAFD2709D0FCB4" blockId="3.[189,775,715,791]" pageId="3" pageNumber="86">
<emphasis id="B935EAED615CFFB35ADAFD2709F8FD42" bold="true" box="[226,425,715,737]" pageId="3" pageNumber="86">Acknowledgements</emphasis>
. The authors express their gratitude to M. Esin (
<collectingRegion id="4985F81D615CFFB35B0BFD0A099DFD58" box="[307,460,742,764]" country="Russia" name="Mordoviya" pageId="3" pageNumber="86">Mordovia State</collectingRegion>
Nature Reserve) for his help in processing samples.
</paragraph>
</subSubSection>
<subSubSection id="C35B6574615CFFB35ADAFCDA0944FCC4" pageId="3" pageNumber="86" type="description">
<paragraph id="8BFE36FF615CFFB35ADAFCDA0944FCC4" blockId="3.[189,775,822,871]" pageId="3" pageNumber="86">
<emphasis id="B935EAED615CFFB35ADAFCDA09E0FCEF" bold="true" box="[226,433,822,844]" pageId="3" pageNumber="86">Competing interests</emphasis>
. The authors declare no competing interests.
</paragraph>
</subSubSection>
</treatment>
</document>