treatments-xml/data/03/E0/F7/03E0F739A661CA272694FE43FA6AECC4.xml
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<document id="5109A2F616C05B872C8755367790653A" ID-CLB-Dataset="9013" ID-DOI="10.11646/zootaxa.4861.3.2" ID-GBIF-Dataset="67893006-2912-4134-835f-37b022607840" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416363" ID-ZooBank="2C2BE3D2-6286-4F36-90A6-920B60367D2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609798826267" checkinUser="plazi" docAuthor="Ashrafi, Hossein, Sari, Alireza &amp; Naderloo, Reza" docDate="2020" docId="03E0F739A661CA272694FE43FA6AECC4" docLanguage="en" docName="zootaxa.4861.3.2.pdf" docOrigin="Zootaxa 4861 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Synalpheus pentaspinosus Ashrafi &amp; Sari &amp; Naderloo 2020, n. sp." docType="treatment" docVersion="8" lastPageNumber="347" masterDocId="FFD98F41A660CA2E2603FFF4FFFAE82D" masterDocTitle="A new sponge-dwelling species of Synalpheus Spence Bate, 1888 (Decapoda Caridea: Alpheidae) from the Persian Gulf" masterLastPageNumber="348" masterPageNumber="338" pageNumber="339" updateTime="1698852170587" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0C214751120F10EEF0E6A2A0164E65FB">A new sponge-dwelling species of Synalpheus Spence Bate, 1888 (Decapoda Caridea: Alpheidae) from the Persian Gulf</mods:title>
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<mods:namePart id="EA77EC90D2591AD3CA6480DE8C1E9A51">Ashrafi, Hossein</mods:namePart>
<mods:affiliation id="9D88FDF30A01E9DE165B1F98E5FD26A3">School of Biology and Centre of Excellence in Phylogeny of Living Organisms, College of Science, University of Tehran, Tehran, Iran Department of Biology and Ecology, and Institute of Environmental Technologies, Faculty of Science, University of Ostrava, Ostrava, Czech Republic</mods:affiliation>
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<mods:namePart id="E85542B6E8221F8D6185912F48EDA2F3">Sari, Alireza</mods:namePart>
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<mods:affiliation id="4AAC9EDBFA3D248B76CEF680FBD4E375">School of Biology and Centre of Excellence in Phylogeny of Living Organisms, College of Science, University of Tehran, Tehran, Iran Department of Biology and Ecology, and Institute of Environmental Technologies, Faculty of Science, University of Ostrava, Ostrava, Czech Republic &amp; https: // orcid. org / 0000 - 0003 - 2294 - 7070</mods:affiliation>
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<mods:namePart id="41A2A7689BDDC1A1556ACFB342B5DF14">Naderloo, Reza</mods:namePart>
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<mods:affiliation id="331D403CF0495FA580BB72B1B1BAE67A">School of Biology and Centre of Excellence in Phylogeny of Living Organisms, College of Science, University of Tehran, Tehran, Iran Department of Biology and Ecology, and Institute of Environmental Technologies, Faculty of Science, University of Ostrava, Ostrava, Czech Republic &amp; https: // orcid. org / 0000 - 0002 - 5020 - 0962</mods:affiliation>
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<treatment id="03E0F739A661CA272694FE43FA6AECC4" ID-DOI="http://doi.org/10.5281/zenodo.4416495" ID-GBIF-Taxon="176442210" ID-Zenodo-Dep="4416495" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4" httpUri="http://treatment.plazi.org/id/03E0F739A661CA272694FE43FA6AECC4" lastPageId="9" lastPageNumber="347" pageId="1" pageNumber="339">
<subSubSection id="C35315A4A661CA2F2694FE43FDDAE9FC" box="[151,544,439,466]" pageId="1" pageNumber="339" type="nomenclature">
<paragraph id="8BF6462FA661CA2F2694FE43FDDAE9FC" blockId="1.[151,544,439,501]" box="[151,544,439,466]" pageId="1" pageNumber="339">
<heading id="D0BEF143A661CA2F2694FE43FDDAE9FC" bold="true" box="[151,544,439,466]" fontSize="11" level="1" pageId="1" pageNumber="339" reason="1">
<emphasis id="B93D9A3DA661CA2F2694FE43FDDAE9FC" bold="true" box="[151,544,439,466]" pageId="1" pageNumber="339">
<taxonomicName id="4C493DACA661CA2F2694FE43FE2CE9FC" ID-CoL="874J7" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[151,470,439,466]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="339" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA661CA2F2694FE43FE2CE9FC" bold="true" box="[151,470,439,466]" italics="true" pageId="1" pageNumber="339">Synalpheus pentaspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E2746A661CA2F27DEFE43FDDAE9FC" box="[477,544,439,465]" pageId="1" pageNumber="339" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C35315A4A661CA2F2694FE2FFEE8E9D8" box="[151,274,475,501]" pageId="1" pageNumber="339" type="description">
<paragraph id="8BF6462FA661CA2F2694FE2FFEE8E9D8" blockId="1.[151,544,439,501]" box="[151,274,475,501]" pageId="1" pageNumber="339">
(
<figureCitation id="13725AAAA661CA2F269CFE2FFEF0E9D8" box="[159,266,475,501]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1943,1967]" captionStartId-1="4.[151,250,1943,1967]" captionStartId-2="5.[151,250,1943,1967]" captionStartId-3="6.[151,250,1943,1967]" captionTargetBox-0="[204,1384,192,1905]" captionTargetBox-1="[217,1356,196,1919]" captionTargetBox-2="[237,1349,192,1911]" captionTargetBox-3="[179,1408,181,1920]" captionTargetId-0="figure-30@3.[191,1396,181,1919]" captionTargetId-1="figure-17@4.[217,1371,181,1919]" captionTargetId-2="figure-30@5.[231,1355,181,1919]" captionTargetId-3="figure-17@6.[179,1408,181,1920]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." captionText-1="FIGURE 2. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) major cheliped, dorso-lateral view; (B) same, palm and fingers, mesial view; (C) same, lateral view; (D) minor cheliped, mesial view; (E) same, lateral view; (F) same, palm and fingers, mesial view." captionText-2="FIGURE 3. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) second pereiopod; (B) third pereiopod; (C) fourth pereiopod; (D) fifth pereiopod; (E) same, propodus and dactylus, mesial view." captionText-3="FIGURE 4. Synalpheus pentaspinosus n. sp., (A) paratype, male (CL 6.7 mm) from Larak Island, Iran (MNHN-IU-2014- 1223); (B) paratype, female (CL 8.4 mm) from the same locality (MNHN-IU-2014-1223); (C) allotype, female (CL 7.9 mm) from the same locality (MNHN-IU-2014-1222). Photographs by Rashed Abdollahi." figureDoi-0="http://doi.org/10.5281/zenodo.4416365" figureDoi-1="http://doi.org/10.5281/zenodo.4416367" figureDoi-2="http://doi.org/10.5281/zenodo.4416369" figureDoi-3="http://doi.org/10.5281/zenodo.4416371" httpUri-0="https://zenodo.org/record/4416365/files/figure.png" httpUri-1="https://zenodo.org/record/4416367/files/figure.png" httpUri-2="https://zenodo.org/record/4416369/files/figure.png" httpUri-3="https://zenodo.org/record/4416371/files/figure.png" pageId="1" pageNumber="339">Figs. 1 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35315A4A661CA2F2694FDD7FDB9ECC4" pageId="1" pageNumber="339" type="materials_examined">
<paragraph id="8BF6462FA661CA2F2694FDD7FC74EB8B" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="339">
<emphasis id="B93D9A3DA661CA2F2694FDD7FEB2EA10" bold="true" box="[151,328,547,573]" pageId="1" pageNumber="339">Type material.</emphasis>
<materialsCitation id="3B214C72A661CA2F2752FDD7FD43EAA8" ID-GBIF-Occurrence="3013446301" collectingDate="2019-05-11" collectionCode="R" country="Iran" county="Keshti-Sukhteh" latitude="26.868717" location="Persian Gulf" longLatPrecision="1" longitude="56.325546" municipality="western Larak Island" pageId="1" pageNumber="339" specimenCount="1" typeStatus="holotype">
<emphasis id="B93D9A3DA661CA2F2752FDD7FE3DEA10" bold="true" box="[337,455,547,573]" pageId="1" pageNumber="339">
<typeStatus id="54F2F88DA661CA2F2752FDD7FE38EA10" box="[337,450,547,573]" pageId="1" pageNumber="339" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (CL
<quantity id="4CB1EBCAA661CA2F2445FDD7FD59EA13" box="[582,675,547,574]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="339" unit="mm" value="7.1">7.1 mm</quantity>
), MNHN-IU-2014-1221,
<collectingCountry id="F35E06BFA661CA2F25CBFDD0FC02EA13" box="[968,1016,548,574]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
,
<location id="8E9610F4A661CA2F2205FDD7FB6DEA10" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4:8E9610F4A661CA2F2205FDD7FB6DEA10" box="[1030,1175,547,573]" country="Iran" county="Keshti-Sukhteh" latitude="26.868717" longLatPrecision="1" longitude="56.325546" municipality="western Larak Island" name="Persian Gulf" pageId="1" pageNumber="339">Persian Gulf</location>
,
<collectingMunicipality id="6B92DC55A661CA2F22A6FDD0FA62EA10" box="[1189,1432,547,574]" pageId="1" pageNumber="339">western Larak Island</collectingMunicipality>
,
<collectingCounty id="62973EA3A661CA2F2694FDB3FEB2EA4C" box="[151,328,583,609]" pageId="1" pageNumber="339">Keshti-Sukhteh</collectingCounty>
,
<geoCoordinate id="EE7D20E8A661CA2F2754FDB3FE0DEA4F" box="[343,503,583,610]" degrees="26" direction="north" minutes="52" orientation="latitude" pageId="1" pageNumber="339" precision="1" seconds="7.38" value="26.868717">26°527.38”N</geoCoordinate>
,
<geoCoordinate id="EE7D20E8A661CA2F2404FDB3FD48EA4F" box="[519,690,583,610]" degrees="56" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="339" precision="1" seconds="31.97" value="56.325546">56°1931.97”E</geoCoordinate>
, sandy with rubbles and many corals, shallow subtidal,
<date id="FFF760EFA661CA2F2340FDB3FF28EAA8" pageId="1" pageNumber="339" value="2019-05-11">
<collectingDate id="EFB39907A661CA2F2340FDB3FF28EAA8" pageId="1" pageNumber="339" value="2019-05-11">11 May 2019</collectingDate>
</date>
, coll.
<collectorName id="26BC23F9A661CA2F2716FD98FE77EAA8" box="[277,397,619,646]" pageId="1" pageNumber="339">H. Ashrafi</collectorName>
, S. Pazoki &amp;
<collectionCode id="ED58DEEAA661CA2F2425FD98FDC6EAAB" box="[550,572,620,646]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="339">R</collectionCode>
.
<collectorName id="26BC23F9A661CA2F2446FD9FFD43EAA8" box="[581,697,619,645]" pageId="1" pageNumber="339">Abdollahi</collectorName>
</materialsCitation>
.
<materialsCitation id="3B214C72A661CA2F24DBFD9FFCFCEA87" ID-GBIF-Occurrence="3013446305" collectingDate="2019-05-11" collectionCode="R" country="Iran" county="Keshti-Sukhteh" latitude="26.868717" location="Persian Gulf" longLatPrecision="1" longitude="56.325546" municipality="western Larak Island" pageId="1" pageNumber="339" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<emphasis id="B93D9A3DA661CA2F24DBFD9FFCBCEAA8" bold="true" box="[728,838,619,645]" pageId="1" pageNumber="339">
<typeStatus id="54F2F88DA661CA2F24DBFD9FFCBBEAA8" box="[728,833,619,645]" pageId="1" pageNumber="339" type="allotype">Allotype</typeStatus>
:
</emphasis>
<specimenCount id="9D4F8DA6A661CA2F254EFD9FFC54EAA8" box="[845,942,619,645]" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
(CL
<quantity id="4CB1EBCAA661CA2F25EAFD9FFBB9EAAB" box="[1001,1091,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="339" unit="mm" value="7.9">7.9 mm</quantity>
), MNHN-IU-2014-1222, same collection data as for holotype, (photo voucher 4135)
</materialsCitation>
.
<emphasis id="B93D9A3DA661CA2F252FFD7BFC57EA84" bold="true" box="[812,941,655,681]" pageId="1" pageNumber="339">
<typeStatus id="54F2F88DA661CA2F252FFD7BFC52EA84" box="[812,936,655,681]" pageId="1" pageNumber="339" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D4F8DA6A661CA2F25BBFD7BFBFCEA84" box="[952,1030,655,681]" pageId="1" pageNumber="339" type="male">1 male</specimenCount>
(CL
<quantity id="4CB1EBCAA661CA2F2244FD7BFB5EEA87" box="[1095,1188,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="339" unit="mm" value="6.7">6.7 mm</quantity>
),
<materialsCitation id="3B214C72A661CA2F22B8FD7BFF26EADC" ID-GBIF-Occurrence="3013446308" collectingDate="2019-05-10" collectionCode="R" country="Iran" county="Keshti-Sukhteh" latitude="26.868717" location="Persian Gulf" longLatPrecision="1" longitude="56.325546" municipality="western Larak Island" pageId="1" pageNumber="339" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D4F8DA6A661CA2F22B8FD7BFAE5EA84" box="[1211,1311,655,681]" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
(
<quantity id="4CB1EBCAA661CA2F2331FD7BFA6AEA87" box="[1330,1424,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="1" pageNumber="339" unit="mm" value="8.4">8.4 mm</quantity>
), MNHN-IU-2014-1223, same collection data as for holotype,
<date id="FFF760EFA661CA2F2555FD47FC08EAE0" box="[854,1010,691,718]" pageId="1" pageNumber="339" value="2019-05-10">
<collectingDate id="EFB39907A661CA2F2555FD47FC08EAE0" box="[854,1010,691,718]" pageId="1" pageNumber="339" value="2019-05-10">10 May 2019</collectingDate>
</date>
, (photo vouchers: female 3932, male 3933)
</materialsCitation>
;
<materialsCitation id="3B214C72A661CA2F26EBFD23FD58EB70" ID-GBIF-Occurrence="3013446306" collectingDate="2016-05-08" collectingDateMax="2019-07-19" collectingDateMin="2016-05-08" collectionCode="ZUTC" collectorName="H. Ashrafi" country="Iran" county="Qadir Park" location="Persian Gulf" longitude="55.073055" municipality="Abu-Musa" pageId="1" pageNumber="339" specimenCode="ZUTC 6495" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
1 ovigerous female (CL
<quantity id="4CB1EBCAA661CA2F27FAFD23FDAEEADF" box="[505,596,727,754]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="339" unit="mm" value="7.2">7.2 mm</quantity>
), MNHN-IU-2014-1224,
<collectingCountry id="F35E06BFA661CA2F2577FD2CFC5EEADF" box="[884,932,728,754]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
,
<location id="8E9610F4A661CA2F25ACFD23FBC5EADC" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4:8E9610F4A661CA2F25ACFD23FBC5EADC" box="[943,1087,727,753]" country="Iran" county="Qadir Park" longitude="55.073055" municipality="Abu-Musa" name="Persian Gulf" pageId="1" pageNumber="339">Persian Gulf</location>
,
<collectingMunicipality id="6B92DC55A661CA2F224AFD23FB3BEADC" box="[1097,1217,727,753]" pageId="1" pageNumber="339">Abu-Musa</collectingMunicipality>
Island,
<collectingCounty id="62973EA3A661CA2F2318FD23FA62EADC" box="[1307,1432,727,753]" pageId="1" pageNumber="339">Qadir Park</collectingCounty>
, 25°8958”N,
<geoCoordinate id="EE7D20E8A661CA2F272EFD0FFE4CEB3B" box="[301,438,763,790]" degrees="55" direction="east" minutes="04" orientation="longitude" pageId="1" pageNumber="339" precision="15" seconds="23" value="55.073055">55°0423”E</geoCoordinate>
, sandy/rocky with live corals,
<date id="FFF760EFA661CA2F2504FD0FFC68EB38" box="[775,914,763,789]" pageId="1" pageNumber="339" value="2019-07-19">
<collectingDate id="EFB39907A661CA2F2504FD0FFC68EB38" box="[775,914,763,789]" pageId="1" pageNumber="339" value="2019-07-19">19 Jul. 2019</collectingDate>
</date>
, coll.
<collectorName id="26BC23F9A661CA2F25D0FD08FBB3EB38" box="[979,1097,763,790]" pageId="1" pageNumber="339">H. Ashrafi</collectorName>
, (photo voucher 5028);
<specimenCount id="9D4F8DA6A661CA2F2351FD0FFA66EB38" box="[1362,1436,763,789]" pageId="1" pageNumber="339" type="male">1 male</specimenCount>
(Cl
<quantity id="4CB1EBCAA661CA2F26C2FCEBFEE0EB17" box="[193,282,799,826]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="1" pageNumber="339" unit="mm" value="6.9">6.9 mm</quantity>
),
<specimenCode id="DBEFEE54A661CA2F272EFCEBFE4DEB17" box="[301,439,799,826]" collectionCode="ZUTC" pageId="1" pageNumber="339">ZUTC 6495</specimenCode>
,
<collectingCountry id="F35E06BFA661CA2F27C2FCD4FE0BEB17" box="[449,497,800,826]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
, Persian Gulf, Abu-Musa Island, Qadir Park, 25°8958”N,
<geoCoordinate id="EE7D20E8A661CA2F2283FCEBFAF3EB17" box="[1152,1289,799,826]" degrees="55" direction="east" minutes="04" orientation="longitude" pageId="1" pageNumber="339" precision="15" seconds="23" value="55.073055">55°0423”E</geoCoordinate>
, sandy/rocky with live corals,
<date id="FFF760EFA661CA2F2752FCB7FE12EB70" box="[337,488,835,862]" pageId="1" pageNumber="339" value="2016-05-08">
<collectingDate id="EFB39907A661CA2F2752FCB7FE12EB70" box="[337,488,835,862]" pageId="1" pageNumber="339" value="2016-05-08">08 May 2016</collectingDate>
</date>
, coll.
<collectorName id="26BC23F9A661CA2F2429FCB0FD58EB70" box="[554,674,835,862]" pageId="1" pageNumber="339">H. Ashrafi</collectorName>
</materialsCitation>
;
<specimenCount id="9D4F8DA6A661CA2F24AEFCB7FD0DEB70" box="[685,759,835,861]" pageId="1" pageNumber="339" type="male">1 male</specimenCount>
(CL
<quantity id="4CB1EBCAA661CA2F2532FCB7FC71EB73" box="[817,907,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="339" unit="mm" value="7.1">7.1 mm</quantity>
),
<materialsCitation id="3B214C72A661CA2F259EFCB7FEBEEB88" ID-GBIF-Occurrence="3013446304" collectingDate="2018-02-02" collectionCode="ZUTC" collectorName="H. Ashrafi" country="Iran" latitude="26.884167" location="Persian Gulf" longLatPrecision="20" longitude="56.400833" municipality="Larak Island" pageId="1" pageNumber="339" specimenCode="ZUTC 6653" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D4F8DA6A661CA2F259EFCB7FC04EB70" box="[925,1022,835,861]" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
(
<quantity id="4CB1EBCAA661CA2F220FFCB7FB9DEB73" box="[1036,1127,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="339" unit="mm" value="6.8">6.8 mm</quantity>
),
<specimenCode id="DBEFEE54A661CA2F2279FCB7FAFEEB73" box="[1146,1284,835,862]" collectionCode="ZUTC" pageId="1" pageNumber="339">ZUTC 6653</specimenCode>
,
<collectingCountry id="F35E06BFA661CA2F230CFCB0FAC5EB73" box="[1295,1343,836,862]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
,
<location id="8E9610F4A661CA2F2349FCB7FF37EBAC" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4:8E9610F4A661CA2F2349FCB7FF37EBAC" country="Iran" latitude="26.884167" longLatPrecision="20" longitude="56.400833" municipality="Larak Island" name="Persian Gulf" pageId="1" pageNumber="339">Persian Gulf</location>
, northeast of
<collectingMunicipality id="6B92DC55A661CA2F276BFC93FE03EBAC" box="[360,505,871,897]" pageId="1" pageNumber="339">Larak Island</collectingMunicipality>
,
<geoCoordinate id="EE7D20E8A661CA2F2405FC93FD6BEBAF" box="[518,657,871,898]" degrees="26" direction="north" minutes="53" orientation="latitude" pageId="1" pageNumber="339" precision="15" seconds="03" value="26.884167">26°5303”N</geoCoordinate>
,
<geoCoordinate id="EE7D20E8A661CA2F249DFC93FCDCEBAF" box="[670,806,871,898]" degrees="56" direction="east" minutes="24" orientation="longitude" pageId="1" pageNumber="339" precision="15" seconds="03" value="56.400833">56°2403”E</geoCoordinate>
, sandy/rubble with corals, low intertidal,
<date id="FFF760EFA661CA2F2303FC93FA62EBAC" box="[1280,1432,871,897]" pageId="1" pageNumber="339" value="2018-02-02">
<collectingDate id="EFB39907A661CA2F2303FC93FA62EBAC" box="[1280,1432,871,897]" pageId="1" pageNumber="339" value="2018-02-02">02 Feb. 2018</collectingDate>
</date>
, coll.
<collectorName id="26BC23F9A661CA2F26CCFC78FEBEEB88" box="[207,324,907,934]" pageId="1" pageNumber="339">H. Ashrafi</collectorName>
</materialsCitation>
(preserved together with small part of host sponge).
</paragraph>
<paragraph id="8BF6462FA661CA2F26C4FC5BFDB9ECC4" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="339">
<emphasis id="B93D9A3DA661CA2F26C4FC5BFD6DEBE7" bold="true" box="[199,663,943,970]" pageId="1" pageNumber="339">
Comparative material.
<taxonomicName id="4C493DACA661CA2F27E0FC5BFD6DEBE7" authorityName="Tattersall" authorityYear="1921" box="[483,663,943,970]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="339" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA661CA2F27E0FC5BFD6DEBE7" bold="true" box="[483,663,943,970]" italics="true" pageId="1" pageNumber="339">S. quinquedens</emphasis>
</taxonomicName>
</emphasis>
.
<specimenCount id="9D4F8DA6A661CA2F24A4FC5BFD0EEBE4" box="[679,756,943,969]" pageId="1" pageNumber="339" type="male">1 male</specimenCount>
(CL
<quantity id="4CB1EBCAA661CA2F2537FC5BFC64EBE7" box="[820,926,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="5.52" pageId="1" pageNumber="339" unit="mm" value="5.52">5.52 mm</quantity>
),
<materialsCitation id="3B214C72A661CA2F25B0FC5BFABCEBC0" ID-GBIF-Occurrence="3013446307" collectingDate="2010-05-12" country="Madagascar" elevation="6" latitude="-25.153334" location="Atimo-Vatae Expedition" longLatPrecision="124" longitude="46.756668" municipality="Sta." pageId="1" pageNumber="339" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D4F8DA6A661CA2F25B0FC5BFBEDEBE4" box="[947,1047,943,969]" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
(CL
<quantity id="4CB1EBCAA661CA2F2255FC5BFB3AEBE7" box="[1110,1216,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="6.24" pageId="1" pageNumber="339" unit="mm" value="6.24">6.24 mm</quantity>
), MNHN-IU-2019- 3150, south
<collectingCountry id="F35E06BFA661CA2F2722FC27FE50EBC0" box="[289,426,979,1005]" name="Madagascar" pageId="1" pageNumber="339">Madagascar</collectingCountry>
,
<location id="8E9610F4A661CA2F27B0FC27FD3DEBC0" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4:8E9610F4A661CA2F27B0FC27FD3DEBC0" box="[435,711,979,1005]" country="Madagascar" latitude="-25.153334" longLatPrecision="124" longitude="46.756668" municipality="Sta." name="Atimo-Vatae Expedition" pageId="1" pageNumber="339">Atimo-Vatae Expedition</location>
,
<collectingMunicipality id="6B92DC55A661CA2F24D1FC27FD07EBC0" box="[722,765,979,1005]" pageId="1" pageNumber="339">Sta.</collectingMunicipality>
TB-11,
<geoCoordinate id="EE7D20E8A661CA2F255AFC27FC37EBC3" box="[857,973,979,1006]" degrees="25" direction="south" minutes="09.2" orientation="latitude" pageId="1" pageNumber="339" precision="92" value="-25.153334">25°09.2S</geoCoordinate>
,
<geoCoordinate id="EE7D20E8A661CA2F25DBFC27FBB4EBC3" box="[984,1102,979,1006]" degrees="46" direction="east" minutes="45.4" orientation="longitude" pageId="1" pageNumber="339" precision="92" value="46.756668">46°45.4E</geoCoordinate>
,
<quantity id="4CB1EBCAA661CA2F225BFC27FB5BEBC3" box="[1112,1185,979,1006]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="339" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">
<elevation id="0064A11CA661CA2F225BFC27FB5BEBC3" box="[1112,1185,979,1006]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="339" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">56 m</elevation>
</quantity>
,
<date id="FFF760EFA661CA2F22AEFC27FABCEBC0" box="[1197,1350,979,1006]" pageId="1" pageNumber="339" value="2010-05-12">
<collectingDate id="EFB39907A661CA2F22AEFC27FABCEBC0" box="[1197,1350,979,1006]" pageId="1" pageNumber="339" value="2010-05-12">12 May 2010</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D4F8DA6A661CA2F2352FC27FA66EBC0" box="[1361,1436,979,1005]" pageId="1" pageNumber="339" type="male">1 male</specimenCount>
(CL
<quantity id="4CB1EBCAA661CA2F26CFFC03FECFEC3F" box="[204,309,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="7.38" pageId="1" pageNumber="339" unit="mm" value="7.38">7.38 mm</quantity>
),
<materialsCitation id="3B214C72A661CA2F2749FC03FAC6EC18" ID-GBIF-Occurrence="3013446309" collectingDate="2017-12-29" collectorName="H. Ashrafi" country="Iran" latitude="26.295452" location="Persian Gulf" longLatPrecision="1" longitude="54.48478" municipality="Farur Island" pageId="1" pageNumber="339" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D4F8DA6A661CA2F2749FC03FE56EC3C" box="[330,428,1015,1041]" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
(CL
<quantity id="4CB1EBCAA661CA2F27EAFC03FDA9EC3F" box="[489,595,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="8.08" pageId="1" pageNumber="339" unit="mm" value="8.08">8.08 mm</quantity>
), MNHN-IU-2014-1225,
<collectingCountry id="F35E06BFA661CA2F2575FC0CFC5CEC3F" box="[886,934,1016,1042]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
,
<location id="8E9610F4A661CA2F25B1FC03FBB9EC3C" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4:8E9610F4A661CA2F25B1FC03FBB9EC3C" box="[946,1091,1015,1041]" country="Iran" latitude="26.295452" longLatPrecision="1" longitude="54.48478" municipality="Farur Island" name="Persian Gulf" pageId="1" pageNumber="339">Persian Gulf</location>
,
<collectingMunicipality id="6B92DC55A661CA2F2253FC0CFB26EC3C" box="[1104,1244,1015,1042]" pageId="1" pageNumber="339">Farur Island</collectingMunicipality>
,
<geoCoordinate id="EE7D20E8A661CA2F22EAFC03FA62EC3C" box="[1257,1432,1015,1041]" degrees="26" direction="north" minutes="17" orientation="latitude" pageId="1" pageNumber="339" precision="1" seconds="43.63" value="26.295452">26°1743.63”N</geoCoordinate>
,
<geoCoordinate id="EE7D20E8A661CA2F2694FBEFFECFEC1B" box="[151,309,1051,1078]" degrees="54" direction="east" minutes="29" orientation="longitude" pageId="1" pageNumber="339" precision="1" seconds="5.21" value="54.48478">54°295.21”E</geoCoordinate>
, shallow subtidal,
<date id="FFF760EFA661CA2F2406FBEFFD64EC18" box="[517,670,1051,1078]" pageId="1" pageNumber="339" value="2017-12-29">
<collectingDate id="EFB39907A661CA2F2406FBEFFD64EC18" box="[517,670,1051,1078]" pageId="1" pageNumber="339" value="2017-12-29">29 Dec. 2017</collectingDate>
</date>
, coll.
<collectorName id="26BC23F9A661CA2F24E2FBE8FCACEC18" box="[737,854,1051,1078]" pageId="1" pageNumber="339">H. Ashrafi</collectorName>
(photo vouchers: male 4393, female 4407)
</materialsCitation>
;
<materialsCitation id="3B214C72A661CA2F2344FBEFFEC7EC8F" ID-GBIF-Occurrence="3013446310" collectingDate="2017-02-08" collectorName="H. Ashrafi &amp; A. Sari" country="Iran" latitude="25.898102" location="Persian Gulf" longLatPrecision="1" longitude="54.522842" municipality="Siri Island" pageId="1" pageNumber="339" specimenCount="1">
3 malefemale pairs (CL of males 6.5, 6.9, and
<quantity id="4CB1EBCAA661CA2F245FFBB4FD5AEC77" box="[604,672,1088,1114]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="339" unit="mm" value="7.0">7 mm</quantity>
, CL of females 9.4, 9.6, and
<quantity id="4CB1EBCAA661CA2F25E8FBCBFBBCEC77" box="[1003,1094,1087,1114]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="339" unit="mm" value="9.7">9.7 mm</quantity>
), MNHN-IU-2014-1226,
<collectingCountry id="F35E06BFA661CA2F236BFBB4FA62EC77" box="[1384,1432,1088,1114]" name="Iran" pageId="1" pageNumber="339">Iran</collectingCountry>
,
<location id="8E9610F4A661CA2F2694FB97FEDFEC50" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4:8E9610F4A661CA2F2694FB97FEDFEC50" box="[151,293,1123,1149]" country="Iran" latitude="25.898102" longLatPrecision="1" longitude="54.522842" municipality="Siri Island" name="Persian Gulf" pageId="1" pageNumber="339">Persian Gulf</location>
,
<collectingMunicipality id="6B92DC55A661CA2F272CFB97FE5FEC50" box="[303,421,1123,1149]" pageId="1" pageNumber="339">Siri Island</collectingMunicipality>
,
<geoCoordinate id="EE7D20E8A661CA2F27ACFB97FD9EEC53" box="[431,612,1123,1150]" degrees="25" direction="north" minutes="53" orientation="latitude" pageId="1" pageNumber="339" precision="1" seconds="53.17" value="25.898102">25°5353.17” N</geoCoordinate>
,
<geoCoordinate id="EE7D20E8A661CA2F246DFB97FCD2EC53" box="[622,808,1123,1150]" degrees="54" direction="east" minutes="31" orientation="longitude" pageId="1" pageNumber="339" precision="1" seconds="22.24" value="54.522842">54° 31 22.24” E</geoCoordinate>
, subtidal,
<date id="FFF760EFA661CA2F259BFB97FBD6EC50" box="[920,1068,1123,1149]" pageId="1" pageNumber="339" value="2017-02-08">
<collectingDate id="EFB39907A661CA2F259BFB97FBD6EC50" box="[920,1068,1123,1149]" pageId="1" pageNumber="339" value="2017-02-08">08 Feb. 2017</collectingDate>
</date>
, coll.
<collectorName id="26BC23F9A661CA2F226FFB90FB1AEC50" box="[1132,1248,1123,1150]" pageId="1" pageNumber="339">H. Ashrafi</collectorName>
&amp;
<collectorName id="26BC23F9A661CA2F22FCFB97FAB7EC50" box="[1279,1357,1123,1149]" pageId="1" pageNumber="339">A. Sari</collectorName>
(photo voucher 1595)
</materialsCitation>
;
<materialsCitation id="3B214C72A661CA2F2749FB73FA78EC8C" ID-GBIF-Occurrence="3013446302" box="[330,1410,1159,1186]" collectingDate="2017-02-08" collectionCode="ZUTC" collectorName="H. Ashrafi &amp; A. Sari" country="Iran" latitude="25.898102" location="Persian Gulf" longLatPrecision="1" longitude="54.522842" municipality="Siri Island" pageId="1" pageNumber="339" specimenCode="ZUTC 6908" specimenCount="6" specimenCount-female="4" specimenCount-male="2">
<specimenCount id="9D4F8DA6A661CA2F2749FB73FE5AEC8C" box="[330,416,1159,1185]" pageId="1" pageNumber="339" type="male">2 males</specimenCount>
and
<specimenCount id="9D4F8DA6A661CA2F27DBFB73FDBFEC8C" box="[472,581,1159,1185]" pageId="1" pageNumber="339" type="female">4 females</specimenCount>
(CL indet.),
<specimenCode id="DBEFEE54A661CA2F24D4FB73FC98EC8C" box="[727,866,1159,1185]" collectionCode="ZUTC" pageId="1" pageNumber="339">ZUTC 6908</specimenCode>
, same collection data as for previous specimens
</materialsCitation>
;
<materialsCitation id="3B214C72A661CA2F238DFB73FF39ECC4" ID-GBIF-Occurrence="3013446303" collectingDate="2014-12-24" collectionCode="OUMNH" collectorName="M. Al-Kandari" country="Kuwait" location="eastern Failaka Island" pageId="1" pageNumber="339" specimenCode="ZC 2020.01" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D4F8DA6A661CA2F238DFB73FF1CECE8" pageId="1" pageNumber="339" type="female">1 female</specimenCount>
,
<collectionCode id="ED58DEEAA661CA2F26F2FB5FFEA4ECE8" box="[241,350,1195,1221]" pageId="1" pageNumber="339">OUMNH</collectionCode>
.
<specimenCode id="DBEFEE54A661CA2F2762FB5FFE11ECE8" box="[353,491,1195,1221]" collectionCode="ZC" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="339" type="University or college">ZC 2020.01</specimenCode>
.022,
<collectingCountry id="F35E06BFA661CA2F2425FB5FFD81ECE8" box="[550,635,1195,1221]" name="Kuwait" pageId="1" pageNumber="339">Kuwait</collectingCountry>
,
<location id="8E9610F4A661CA2F2485FB58FC86ECE8" LSID="urn:lsid:plazi:treatment:03E0F739A661CA272694FE43FA6AECC4:8E9610F4A661CA2F2485FB58FC86ECE8" box="[646,892,1195,1222]" country="Kuwait" name="eastern Failaka Island" pageId="1" pageNumber="339">eastern Failaka Island</location>
, rocky substrate,
<date id="FFF760EFA661CA2F223CFB5FFB2DECE8" box="[1087,1239,1195,1222]" pageId="1" pageNumber="339" value="2014-12-24">
<collectingDate id="EFB39907A661CA2F223CFB5FFB2DECE8" box="[1087,1239,1195,1222]" pageId="1" pageNumber="339" value="2014-12-24">24 Dec. 2014</collectingDate>
</date>
, coll.
<collectorName id="26BC23F9A661CA2F2319FB58FF39ECC4" pageId="1" pageNumber="339">M. Al-Kandari</collectorName>
</materialsCitation>
<emphasis id="B93D9A3DA661CA2F26C9FB24FF01ECC4" box="[202,251,1231,1257]" italics="true" pageId="1" pageNumber="339">et al</emphasis>
. (examined by S. De Grave).
</paragraph>
</subSubSection>
<subSubSection id="C35315A4A661CA2C26C4FB07FDF6EFB8" lastPageId="2" lastPageNumber="340" pageId="1" pageNumber="339" type="description">
<paragraph id="8BF6462FA661CA2F26C4FB07FB03EDEF" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="339">
<emphasis id="B93D9A3DA661CA2F26C4FB07FEA3ED20" bold="true" box="[199,345,1267,1293]" pageId="1" pageNumber="339">Description.</emphasis>
Medium-sized species of
<taxonomicName id="4C493DACA661CA2F2481FB07FCF8ED20" authorityName="Spence Bate" authorityYear="1888" box="[642,770,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="339" phylum="Arthropoda" rank="genus">
<emphasis id="B93D9A3DA661CA2F2481FB07FCF8ED20" box="[642,770,1267,1293]" italics="true" pageId="1" pageNumber="339">Synalpheus</emphasis>
</taxonomicName>
. Carapace (
<figureCitation id="13725AAAA661CA2F258AFB07FC29ED20" box="[905,979,1267,1293]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="1" pageNumber="339">Figs. 1</figureCitation>
AC) sparsely setose, with deep cardiac notch; lower margin with some short setae; rostrum subacute, slightly tapering distally, about 2.5 times as long as wide, reaching almost to middle of first antennular article, slightly up-turned in lateral view, with two setae at tip; orbital teeth subtriangular, subacute, about two times as wide as rostrum at base, slightly falling short of rostrum, nearly as long as wide, with two or three setae at tip; frontal margin between rostrum and orbital teeth deeply concave, curved on orbital tooth side, straight on rostrum side; pterygostomial angle bluntly protruding.
</paragraph>
<paragraph id="8BF6462FA661CA2F26C4FA3FFB85EDC8" blockId="1.[151,1437,547,2014]" box="[199,1151,1483,1510]" pageId="1" pageNumber="339">
Each eye (
<figureCitation id="13725AAAA661CA2F273FFA3FFE6BEDC8" box="[316,401,1483,1509]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="1" pageNumber="339">Fig. 1B</figureCitation>
) with well-developed cornea and small blunt projection anteriorly.
</paragraph>
<paragraph id="8BF6462FA661CA2F26C4FA1BFF2FEF28" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="339">
Pleon showing sexual dimorphism on first to fourth pleonites, with subacute projections, as described below, in males, rounded antero- and postero-ventrally in females. Male pleon (
<figureCitation id="13725AAAA661CA2F25A2F9E7FC0CEE00" box="[929,1014,1555,1581]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="1" pageNumber="339">Fig. 1C</figureCitation>
) with first pleuron bearing small subacute projection antero-ventrally and ventrally pointing, hook-shaped projection postero-ventrally; second pleuron slightly angled antero-ventrally, with subacute triangular projection postero-ventrally; third pleuron bearing small blunt projection antero-ventrally, slightly projecting postero-ventrally; fourth pleuron rounded antero-ventrally, angular postero-ventrally; fifth pleuron rounded antero- and postero-ventrally. Sixth pleonite in both sexes (
<figureCitation id="13725AAAA661CA2F233CF957FF56EECF" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="1" pageNumber="339">Fig. 1C, D</figureCitation>
) armed with five strong sharp teeth on posterior margin, most-lateral teeth flanking telson stronger than remaining teeth.
</paragraph>
<paragraph id="8BF6462FA661CA2C26C4F8FBFF0AE8D4" blockId="1.[151,1437,547,2014]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="340" pageId="1" pageNumber="339">
Telson (
<figureCitation id="13725AAAA661CA2F2722F8FBFE82EF04" box="[289,376,1807,1833]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="1" pageNumber="339">Fig. 1D</figureCitation>
) subrectangular, tapering distally, with shallow mid-dorsal depression starting from 0.3 length of telson and almost reaching posterior end of telson, broadening posteriorly; dorsal surface with two pairs of spiniform setae on 0.3 and 0.7 length of telson, respectively; posterolateral angles slightly projecting; posterior margin broad, rounded, with two pairs of spiniform setae, one pair near each posterolateral angle, mesial spiniform setae about twice as long as lateral ones. Uropod with protopod armed with two sharp teeth, lateral longer than mesial; exopod bearing stout disto-lateral tooth; diaeresis sinuous, lateral margin with stout acute tooth, almost as long as disto-lateral tooth of exopod; disto-lateral spiniform seta adjacent to diaresis stout, overreaching both teeth, but not reaching beyond posterior margin of exopod; endopod narrowly ovate, with several clusters of setae on dorsal surface.
</paragraph>
<paragraph id="8BF6462FA662CA2C26C4FEF7FE2FE9FC" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
Antennule (
<figureCitation id="13725AAAA662CA2C274FFEF7FE3FE930" box="[332,453,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="2" pageNumber="340">Fig. 1A, B</figureCitation>
) relatively slender; first article longest, about twice as long as wide, with several short setae on distodorsal and mesial margins; stylocerite subacute, extending well beyond distal margin of first article, but not reaching mid-length of second article; second article nearly 0.8 times as long as first, about 1.5 times as long as wide, bearing short setae disto-dorsally, with scattered setae mesially; third article slightly shorter than second; lateral flagellum biramous; fused portion consisting of seven or eight units; aesthetascs extending from fifth fused unit to end of shorter ramus.
</paragraph>
<paragraph id="8BF6462FA662CA2C26C4FE2FFB8DEA4C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
Antenna (
<figureCitation id="13725AAAA662CA2C273BFE2FFE4CE9D8" box="[312,438,475,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="2" pageNumber="340">Fig. 1A, B</figureCitation>
) with very robust basicerite, armed with stout subacute disto-dorsal tooth and stouter, longer disto-ventral tooth, latter reaching to tip of antennular stylocerite; lateral tooth of scaphocerite very strong, exceeding antennular peduncle, blade narrow, reaching to about 0.7 length of disto-lateral tooth, with short marginal setae. Carpocerite slender, extending well beyond scaphocerite and antennular peduncle.
</paragraph>
<paragraph id="8BF6462FA662CA2C26C4FD9FFE65EB14" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
Mouthparts not dissected. Third maxillipeds (
<figureCitation id="13725AAAA662CA2C24C5FD9FFCE3EAA8" box="[710,793,619,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[204,1384,192,1905]" captionTargetId="figure-30@3.[191,1396,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="2" pageNumber="340">Fig. 1E</figureCitation>
) pediform; coxa and basis fused; lateral plate with small, laterally pointing, acute projection, mesial surface with short setae; antepenultimate article trigonal in cross section, about four times as long as wide, concave proximo-mesially, margins furnished with long setae; penultimate article short, about one fourth as long as antepenultimate article; ultimate article slender, nearly as long as antepenultimate article; tip with crown of eight or so slender spiniform setae; exopod slender, approximately reaching to end of antepenultimate article.
</paragraph>
<paragraph id="8BF6462FA662CA2C26C4FCB7FA9EECC4" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
First pereiopods very unequal in size and asymmetrical in shape. Major cheliped (
<figureCitation id="13725AAAA662CA2C2286FCB7FB30EB70" box="[1157,1226,835,861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[217,1356,196,1919]" captionTargetId="figure-17@4.[217,1371,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) major cheliped, dorso-lateral view; (B) same, palm and fingers, mesial view; (C) same, lateral view; (D) minor cheliped, mesial view; (E) same, lateral view; (F) same, palm and fingers, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4416367" httpUri="https://zenodo.org/record/4416367/files/figure.png" pageId="2" pageNumber="340">Fig. 2</figureCitation>
AC) robust, with ischium short, about 1.5 times as wide as long; merus robust, somewhat swollen, about twice as long as high, with small subacute projection disto-dorsally, shallowly concave ventrally; carpus short, cup-shaped; palm tumid, subcylindrical, smooth, with prominent rounded lobe and three small bumps disto-dorsally; dactylus 1.3 times as long as pollex and 0.4 times as long as palm, curved distally. Minor cheliped (
<figureCitation id="13725AAAA662CA2C25B0FC27FC0FEBC0" box="[947,1013,979,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[217,1356,196,1919]" captionTargetId="figure-17@4.[217,1371,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) major cheliped, dorso-lateral view; (B) same, palm and fingers, mesial view; (C) same, lateral view; (D) minor cheliped, mesial view; (E) same, lateral view; (F) same, palm and fingers, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4416367" httpUri="https://zenodo.org/record/4416367/files/figure.png" pageId="2" pageNumber="340">Fig. 2</figureCitation>
DF) with basis short, slightly longer than wide, with few long setae on ventral margin; ischium oblique distally, bearing some long setae ventrally; merus approximately six times as long as ischium, tapering distally, about 3 times as long as its widest part, furnished with long setae along ventral margin and short setae on dorsal margin, shallowly concave ventrally, terminating in small obtuse projection disto-dorsally; carpus vase-shaped, about 0.4 as long as merus; chela nearly as long as merus; palm ovoid, subcylindrical, about 1.2 times as long as fingers; fingers subequal in length; pollex with one short row of setae mesially and one laterally, with simple, non-bidentate tip; dactylus with medium-sized depression on cutting edge, with simple, non-bidentate tip; two prominent rows of setae present, one mesially and one dorsally.
</paragraph>
<paragraph id="8BF6462FA662CA2C26CCFB07FD86ED54" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
Second pereiopod (
<figureCitation id="13725AAAA662CA2C27AAFB07FDF8ED20" box="[425,514,1267,1293]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[237,1349,192,1911]" captionTargetId="figure-30@5.[231,1355,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) second pereiopod; (B) third pereiopod; (C) fourth pereiopod; (D) fifth pereiopod; (E) same, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4416369" httpUri="https://zenodo.org/record/4416369/files/figure.png" pageId="2" pageNumber="340">Fig. 3A</figureCitation>
) relatively slender; coxa with small tooth distodorsally; basis short; ischium slen- der, about 3.5 times as long as widest part; merus slender, about 1.2 times longer than ischium, five times as long as broad, with long setae ventrally; carpus slightly longer than merus, with five units (subarticles) approximate ratio 5: 1: 1: 1: 2; chela about half of merus length.
</paragraph>
<paragraph id="8BF6462FA662CA2C26C4FA77FBBCEE7C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
Third pereiopod (
<figureCitation id="13725AAAA662CA2C2790FA77FE16EDB0" box="[403,492,1411,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[237,1349,192,1911]" captionTargetId="figure-30@5.[231,1355,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) second pereiopod; (B) third pereiopod; (C) fourth pereiopod; (D) fifth pereiopod; (E) same, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4416369" httpUri="https://zenodo.org/record/4416369/files/figure.png" pageId="2" pageNumber="340">Fig. 3B</figureCitation>
) more robust than other walking legs; coxa with small distodorsal lobe; ischium slightly longer than basis, about 1.3 as long as its widest part; merus robust, broadened, slightly spindle-shaped, three times as long as widest part, about 2.2 as long as carpus; carpus about 2.3 times as long as wide, with one spiniform seta distoventrally; propodus about 1.8 times longer than carpus, bearing seven or eight spiniform setae along ventral margin, not including one distal pair, adjacent to dactylus; dactylus biunguiculate, with dorsal unguis slightly longer than ventral unguis, latter slightly broader at base than dorsal unguis.
</paragraph>
<paragraph id="8BF6462FA662CA2C26C4F9AFFC37EE90" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
Fourth pereiopod (
<figureCitation id="13725AAAA662CA2C2798F9AFFE08EE58" box="[411,498,1627,1653]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[237,1349,192,1911]" captionTargetId="figure-30@5.[231,1355,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) second pereiopod; (B) third pereiopod; (C) fourth pereiopod; (D) fifth pereiopod; (E) same, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4416369" httpUri="https://zenodo.org/record/4416369/files/figure.png" pageId="2" pageNumber="340">Fig. 3C</figureCitation>
) generally similar to third pereiopod, less robust; coxa with one disto-dorsal lobe; merus about 2.3 times as long as ischium; carpus with one spiniform seta disto-ventrally; propodus with seven spiniform setae on ventral margin and one pair distally, adjacent to dactylus.
</paragraph>
<paragraph id="8BF6462FA662CA2C26C4F933FDF6EFB8" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
Fifth pereiopod (
<figureCitation id="13725AAAA662CA2C2789F933FE37EECC" box="[394,461,1735,1761]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[237,1349,192,1911]" captionTargetId="figure-30@5.[231,1355,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Synalpheus pentaspinosus n. sp., holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) second pereiopod; (B) third pereiopod; (C) fourth pereiopod; (D) fifth pereiopod; (E) same, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4416369" httpUri="https://zenodo.org/record/4416369/files/figure.png" pageId="2" pageNumber="340">Fig. 3</figureCitation>
DE) more slender than other walking legs; coxa without disto-dorsal lobe; ischium about twice as long as widest part, approximately as long as basis; merus about four times as long as broad, nearly 2.3 times as long as ischium; carpus slender, about 0.7 as long as merus, without disto-ventral spiniform seta; propodus slightly longer than carpus, with three spiniform setae ventrally and one pair distally, adjacent to dactylus, and with five transverse rows of serrulate setae on distal half of ventro-mesial margin forming cleaning brush; dactylus similar to that of third pereiopod.
</paragraph>
</subSubSection>
<subSubSection id="C35315A4A662CA2C26C4F86BFD24EFF0" pageId="2" pageNumber="340" type="etymology">
<paragraph id="8BF6462FA662CA2C26C4F86BFD24EFF0" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="340">
<emphasis id="B93D9A3DA662CA2C26C4F86BFEB4EF94" bold="true" box="[199,334,1951,1977]" pageId="2" pageNumber="340">Etymology.</emphasis>
The new species name refers to the presence of five sharp spine-like teeth on the sixth pleonite, one of its main diagnostic features; used as an adjective.
</paragraph>
</subSubSection>
<caption id="DF3616A7A663CA2D2694F863FEA7EFD5" ID-DOI="http://doi.org/10.5281/zenodo.4416365" ID-Zenodo-Dep="4416365" httpUri="https://zenodo.org/record/4416365/files/figure.png" pageId="3" pageNumber="341" startId="3.[151,250,1943,1967]" targetBox="[204,1384,192,1905]" targetPageId="3">
<paragraph id="8BF6462FA663CA2D2694F863FEA7EFD5" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="341">
<emphasis id="B93D9A3DA663CA2D2694F863FEF5EF82" bold="true" box="[151,271,1943,1967]" pageId="3" pageNumber="341">FIGURE 1</emphasis>
.
<taxonomicName id="4C493DACA663CA2D271DF86CFDD0EF82" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[286,554,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="341" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA663CA2D271DF86CFDD0EF82" box="[286,554,1944,1968]" italics="true" pageId="3" pageNumber="341">Synalpheus pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA663CA2D2430F86CFD94EF9D" bold="true" box="[563,622,1944,1968]" pageId="3" pageNumber="341">
<taxonomicNameLabel id="A20E2746A663CA2D2430F86CFD94EF9D" box="[563,622,1944,1968]" pageId="3" pageNumber="341" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) anterior region, dorsal view; (B) anterior region, lateral view; (C) pleon, lateral view; (D) telson and uropods; (E) third maxilliped, ventro-lateral view.
</paragraph>
</caption>
<caption id="DF3616A7A664CA2A2694F863FD02EFD5" ID-DOI="http://doi.org/10.5281/zenodo.4416367" ID-Zenodo-Dep="4416367" httpUri="https://zenodo.org/record/4416367/files/figure.png" pageId="4" pageNumber="342" startId="4.[151,250,1943,1967]" targetBox="[217,1356,196,1919]" targetPageId="4">
<paragraph id="8BF6462FA664CA2A2694F863FD02EFD5" blockId="4.[151,1437,1943,2040]" pageId="4" pageNumber="342">
<emphasis id="B93D9A3DA664CA2A2694F863FEE8EF82" bold="true" box="[151,274,1943,1967]" pageId="4" pageNumber="342">FIGURE 2.</emphasis>
<taxonomicName id="4C493DACA664CA2A271BF86CFDDBEF82" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[280,545,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="342" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA664CA2A271BF86CFDDBEF82" box="[280,545,1944,1968]" italics="true" pageId="4" pageNumber="342">Synalpheus pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA664CA2A2425F86CFDA5EF9D" bold="true" box="[550,607,1944,1968]" pageId="4" pageNumber="342">
<taxonomicNameLabel id="A20E2746A664CA2A2425F86CFDA5EF9D" box="[550,607,1944,1968]" pageId="4" pageNumber="342" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) major cheliped, dorso-lateral view; (B) same, palm and fingers, mesial view; (C) same, lateral view; (D) minor cheliped, mesial view; (E) same, lateral view; (F) same, palm and fingers, mesial view.
</paragraph>
</caption>
<caption id="DF3616A7A665CA2B2694F863FAA5EFF9" ID-DOI="http://doi.org/10.5281/zenodo.4416369" ID-Zenodo-Dep="4416369" httpUri="https://zenodo.org/record/4416369/files/figure.png" pageId="5" pageNumber="343" startId="5.[151,250,1943,1967]" targetBox="[237,1349,192,1911]" targetPageId="5">
<paragraph id="8BF6462FA665CA2B2694F863FAA5EFF9" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="343">
<emphasis id="B93D9A3DA665CA2B2694F863FEECEF82" bold="true" box="[151,278,1943,1967]" pageId="5" pageNumber="343">FIGURE 3.</emphasis>
<taxonomicName id="4C493DACA665CA2B271CF86CFDD6EF82" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[287,556,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="343" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA665CA2B271CF86CFDD6EF82" box="[287,556,1944,1968]" italics="true" pageId="5" pageNumber="343">Synalpheus pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA665CA2B2436F86CFD8BEF9D" bold="true" box="[565,625,1944,1968]" pageId="5" pageNumber="343">
<taxonomicNameLabel id="A20E2746A665CA2B2436F86CFD8BEF9D" box="[565,625,1944,1968]" pageId="5" pageNumber="343" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male from Larak Island, Iran (MNHN-IU-2014-1221), (A) second pereiopod; (B) third pereiopod; (C) fourth pereiopod; (D) fifth pereiopod; (E) same, propodus and dactylus, mesial view.
</paragraph>
</caption>
<caption id="DF3616A7A666CA282694F863FC1CEFD5" ID-DOI="http://doi.org/10.5281/zenodo.4416371" ID-Zenodo-Dep="4416371" httpUri="https://zenodo.org/record/4416371/files/figure.png" pageId="6" pageNumber="344" startId="6.[151,250,1943,1967]" targetBox="[179,1408,181,1920]" targetPageId="6">
<paragraph id="8BF6462FA666CA282694F863FC1CEFD5" blockId="6.[151,1436,1943,2040]" pageId="6" pageNumber="344">
<emphasis id="B93D9A3DA666CA282694F863FEEAEF82" bold="true" box="[151,272,1943,1967]" pageId="6" pageNumber="344">FIGURE 4</emphasis>
.
<taxonomicName id="4C493DACA666CA28271CF86CFDD6EF82" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[287,556,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="344" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA666CA28271CF86CFDD6EF82" box="[287,556,1944,1968]" italics="true" pageId="6" pageNumber="344">Synalpheus pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA666CA282436F86CFD8DEF9D" bold="true" box="[565,631,1944,1968]" pageId="6" pageNumber="344">
<taxonomicNameLabel id="A20E2746A666CA282436F86CFD88EF9D" box="[565,626,1944,1968]" pageId="6" pageNumber="344" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
(A) paratype, male (CL 6.7 mm) from Larak Island, Iran (MNHN-IU-2014- 1223); (B) paratype, female (CL 8.4 mm) from the same locality (MNHN-IU-2014-1223); (C) allotype, female (CL 7.9 mm) from the same locality (MNHN-IU-2014-1222). Photographs by Rashed Abdollahi.
</paragraph>
</caption>
<caption id="DF3616A7A667CA292694F9C0FB0FEE5D" ID-DOI="http://doi.org/10.5281/zenodo.4416373" ID-Zenodo-Dep="4416373" httpUri="https://zenodo.org/record/4416373/files/figure.png" pageId="7" pageNumber="345" startId="7.[151,250,1588,1612]" targetBox="[151,1437,181,1564]" targetPageId="7">
<paragraph id="8BF6462FA667CA292694F9C0FB0FEE5D" blockId="7.[151,1436,1588,1649]" pageId="7" pageNumber="345">
<emphasis id="B93D9A3DA667CA292694F9C0FEF6EE60" bold="true" box="[151,268,1588,1613]" pageId="7" pageNumber="345">FIGURE 5</emphasis>
.
<taxonomicName id="4C493DACA667CA29271BF9C0FD4FEE60" authority="Tattersall, 1921" authorityName="Tattersall" authorityYear="1921" box="[280,693,1588,1613]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="345" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA667CA29271BF9C0FDEBEE61" box="[280,529,1588,1612]" italics="true" pageId="7" pageNumber="345">Synalpheus quinquedens</emphasis>
<bibRefCitation id="EFD83BDEA667CA292415F9C0FD4FEE60" author="Tattersall, W. M." box="[534,693,1588,1613]" pageId="7" pageNumber="345" pagination="345 - 398" refId="ref5073" refString="Tattersall, W. M. (1921) Report on the stomatopoda and macrurous decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of the Linnean Society of London, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall, 1921</bibRefCitation>
</taxonomicName>
, (A) male (CL 7.38 mm) from Farur Island, Iran (MNHN-IU-2014-1225); (B) female (CL 8.08 mm) from Farur Island, Iran (MNHN-IU-2014-1225). Photographs by Rashed Abdollahi.
</paragraph>
</caption>
<subSubSection id="C35315A4A667CA2926C4F96BFD01EF67" pageId="7" pageNumber="345" type="biology_ecology">
<paragraph id="8BF6462FA667CA2926C4F96BFD01EF67" blockId="7.[151,1437,1695,2009]" pageId="7" pageNumber="345">
<emphasis id="B93D9A3DA667CA2926C4F96BFED6EE94" bold="true" box="[199,300,1695,1721]" pageId="7" pageNumber="345">Ecology.</emphasis>
All specimens were found inside sponge canals in the low intertidal and shallow subtidal areas. The host sponges were either partly overgrowing on hard corals or were growing within a mixture of cobble, rocks, coral and coral rubble. The same sponges were also found in the intertidal and shallow subtidal areas without corals. However, these sponges were occupied by another, presumably undescribed species of
<taxonomicName id="4C493DACA667CA29229AF8FFFAE3EF08" authorityName="Spence Bate" authorityYear="1888" box="[1177,1305,1803,1829]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B93D9A3DA667CA29229AF8FFFAE3EF08" box="[1177,1305,1803,1829]" italics="true" pageId="7" pageNumber="345">Synalpheus</emphasis>
</taxonomicName>
from the
<taxonomicName id="4C493DACA667CA292384F8FFFED4EF64" baseAuthorityName="Paul'son" baseAuthorityYear="1875" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="345" phylum="Arthropoda" rank="species" species="tumidomanus">
<emphasis id="B93D9A3DA667CA292384F8FFFED4EF64" italics="true" pageId="7" pageNumber="345">S. tumidomanus</emphasis>
</taxonomicName>
species complex (H. Ashrafi, pers. obs.).
</paragraph>
</subSubSection>
<subSubSection id="C35315A4A667CA2726C4F8A7FA6AECC4" lastPageId="9" lastPageNumber="347" pageId="7" pageNumber="345" type="discussion">
<paragraph id="8BF6462FA667CA2726C4F8A7FBF0E948" blockId="7.[151,1437,1695,2009]" lastBlockId="9.[151,1437,151,1257]" lastPageId="9" lastPageNumber="347" pageId="7" pageNumber="345">
<emphasis id="B93D9A3DA667CA2926C4F8A7FEC1EF40" bold="true" box="[199,315,1875,1901]" pageId="7" pageNumber="345">Remarks.</emphasis>
The new species belongs to a small morphologically distinctive group of
<taxonomicName id="4C493DACA667CA2922A5F8A7FADCEF40" authorityName="Spence Bate" authorityYear="1888" box="[1190,1318,1875,1901]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B93D9A3DA667CA2922A5F8A7FADCEF40" box="[1190,1318,1875,1901]" italics="true" pageId="7" pageNumber="345">Synalpheus</emphasis>
</taxonomicName>
, in which the posterior margin of the sixth pleonite is armed with strong teeth. Whether this group is monophyletic remains to be shown. Based on the shape of the rostrum and orbital teeth, this group can be further subdivided into three smaller subgroups. The first subgroup includes
<taxonomicName id="4C493DACA667CA2924C3F84BFBF0EFF4" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[704,1034,1983,2009]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="345" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B93D9A3DA667CA2924C3F84BFCAFEFF4" box="[704,853,1983,2009]" italics="true" pageId="7" pageNumber="345">S. bispinosus</emphasis>
<bibRefCitation id="EFD83BDEA667CA292563F84BFBF0EFF4" author="De Man, J. G." box="[864,1034,1983,2009]" pageId="7" pageNumber="345" pagination="287 - 319" refId="ref4804" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 11, 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C493DACA667CA29221AF84BFA62EFF4" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[1049,1432,1983,2009]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="345" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B93D9A3DA667CA29221AF84BFB19EFF4" box="[1049,1251,1983,2009]" italics="true" pageId="7" pageNumber="345">S. quadrispinosus</emphasis>
<bibRefCitation id="EFD83BDEA667CA2922EDF84BFA62EFF4" author="De Man, J. G." box="[1262,1432,1983,2009]" pageId="7" pageNumber="345" pagination="287 - 319" refId="ref4804" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 11, 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C493DACA669CA272694FF63FE0EE89C" authority="Tattersall, 1921" authorityName="Tattersall" authorityYear="1921" box="[151,500,151,177]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272694FF63FEBBE89C" box="[151,321,151,177]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
<bibRefCitation id="EFD83BDEA669CA272744FF63FE0EE89C" author="Tattersall, W. M." box="[327,500,151,177]" pageId="9" pageNumber="347" pagination="345 - 398" refId="ref5073" refString="Tattersall, W. M. (1921) Report on the stomatopoda and macrurous decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of the Linnean Society of London, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall, 1921</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C493DACA669CA272429FF63FD1FE89D" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[554,741,151,177]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA272429FF63FD1FE89D" box="[554,741,151,177]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E2746A669CA2724EFFF63FCD0E89C" box="[748,810,151,177]" pageId="9" pageNumber="347" rank="species">
<emphasis id="B93D9A3DA669CA2724EFFF63FCD9E89C" bold="true" box="[748,803,151,177]" pageId="9" pageNumber="347">n. sp</emphasis>
.
</taxonomicNameLabel>
, all with the rostrum and orbital teeth being subequal in length and with the orbital teeth distinctly wider than the rostrum. The second subgroup is comprised of
<taxonomicName id="4C493DACA669CA272325FF4FFE8AE8D4" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="triacanthus">
<emphasis id="B93D9A3DA669CA272325FF4FFF3CE8D4" italics="true" pageId="9" pageNumber="347">S. triacanthus</emphasis>
<bibRefCitation id="EFD83BDEA669CA2726CEFF2BFE8AE8D4" author="De Man, J. G." box="[205,368,223,249]" pageId="9" pageNumber="347" pagination="287 - 319" refId="ref4804" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 11, 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C493DACA669CA272778FF2BFD40E8D4" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[379,698,223,249]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="B93D9A3DA669CA272778FF2BFDE8E8D5" box="[379,530,223,249]" italics="true" pageId="9" pageNumber="347">S. trispinosus</emphasis>
<bibRefCitation id="EFD83BDEA669CA27241AFF2BFD40E8D4" author="De Man, J. G." box="[537,698,223,249]" pageId="9" pageNumber="347" pagination="287 - 319" refId="ref4804" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 11, 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C493DACA669CA2724F3FF2BFB53E8D4" authority="Banner &amp; Banner, 1979" authorityName="Banner &amp; Banner" authorityYear="1979" box="[752,1193,223,249]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="cretoculatus">
<emphasis id="B93D9A3DA669CA2724F3FF2BFC6FE8D4" box="[752,917,223,249]" italics="true" pageId="9" pageNumber="347">S. cretoculatus</emphasis>
<bibRefCitation id="EFD83BDEA669CA27259FFF2BFB53E8D4" author="Banner, A. H. &amp; Banner, D. M." box="[924,1193,223,249]" pageId="9" pageNumber="347" pagination="25 - 35" refId="ref4596" refString="Banner, A. H. &amp; Banner, D. M. (1979) Some small collections of alpheid shrimp from the Indian Ocean, including two new species of the genus Synalpheus. Pacific Science, 33, 25 - 35." type="journal article" year="1979">Banner &amp; Banner, 1979</bibRefCitation>
</taxonomicName>
, in which the rostrum is significantly longer and much more slender than the orbital teeth (cf.
<bibRefCitation id="EFD83BDEA669CA2725B1FEF7FBB7E930" author="De Man, J. G." box="[946,1101,259,285]" pageId="9" pageNumber="347" pagination="133 - 465" refId="ref4842" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition, Part II, family Alpheidae. Monographic, 39 a, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="book chapter" year="1911">De Man 1911</bibRefCitation>
;
<bibRefCitation id="EFD83BDEA669CA27225BFEF7FAA5E930" author="Banner, A. H. &amp; Banner, D. M." box="[1112,1375,259,285]" pageId="9" pageNumber="347" pagination="25 - 35" refId="ref4596" refString="Banner, A. H. &amp; Banner, D. M. (1979) Some small collections of alpheid shrimp from the Indian Ocean, including two new species of the genus Synalpheus. Pacific Science, 33, 25 - 35." type="journal article" year="1979">Banner &amp; Banner 1979</bibRefCitation>
). The third subgroup includes only
<taxonomicName id="4C493DACA669CA2727E4FED3FC9FE96C" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[487,869,295,321]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="septemspinosus">
<emphasis id="B93D9A3DA669CA2727E4FED3FD49E96D" box="[487,691,295,321]" italics="true" pageId="9" pageNumber="347">S. septemspinosus</emphasis>
<bibRefCitation id="EFD83BDEA669CA2724BEFED3FC9FE96C" author="De Man, J. G." box="[701,869,295,321]" pageId="9" pageNumber="347" pagination="287 - 319" refId="ref4804" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 11, 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
, which is characterised by small orbital teeth and rostrum, the latter also being longer relative to orbital teeth (cf.
<bibRefCitation id="EFD83BDEA669CA272563FEBFFC07E948" author="De Man, J. G." box="[864,1021,331,357]" pageId="9" pageNumber="347" pagination="133 - 465" refId="ref4842" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition, Part II, family Alpheidae. Monographic, 39 a, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="book chapter" year="1911">De Man 1911</bibRefCitation>
).
</paragraph>
<caption id="DF3616A7A668CA262694F863FE58EFD5" ID-DOI="http://doi.org/10.5281/zenodo.4416375" ID-Zenodo-Dep="4416375" httpUri="https://zenodo.org/record/4416375/files/figure.png" pageId="8" pageNumber="346" startId="8.[151,250,1943,1967]" targetBox="[166,1410,195,1912]" targetPageId="8">
<paragraph id="8BF6462FA668CA262694F863FE58EFD5" blockId="8.[151,1437,1943,2040]" pageId="8" pageNumber="346">
<emphasis id="B93D9A3DA668CA262694F863FEF4EF82" bold="true" box="[151,270,1943,1967]" pageId="8" pageNumber="346">FIGURE 6</emphasis>
.
<taxonomicName id="4C493DACA668CA262718F86CFD47EF9D" authority="Tattersall, 1921" authorityName="Tattersall" authorityYear="1921" box="[283,701,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="346" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA668CA262718F86CFDEFEF9D" box="[283,533,1944,1968]" italics="true" pageId="8" pageNumber="346">Synalpheus quinquedens</emphasis>
<bibRefCitation id="EFD83BDEA668CA26241FF86CFD47EF9D" author="Tattersall, W. M." box="[540,701,1944,1968]" pageId="8" pageNumber="346" pagination="345 - 398" refId="ref5073" refString="Tattersall, W. M. (1921) Report on the stomatopoda and macrurous decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of the Linnean Society of London, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall, 1921</bibRefCitation>
</taxonomicName>
, female from Farur Island, Iran (CL 12.3 mm) (MNHN-IU-2014-1225), (A) anterior region, dorsal view; (B) third maxilliped, ventrolateral view; (C) minor cheliped, mesial view; (D) same, palm and fingers, dorsomesial view.
</paragraph>
</caption>
<paragraph id="8BF6462FA669CA2726C4FE9BFA6FEAE0" blockId="9.[151,1437,151,1257]" pageId="9" pageNumber="347">
Apart from the difference in the number of teeth on the posterior margin of sixth pleonite (two in
<taxonomicName id="4C493DACA669CA272300FE9BFA6FE9A4" authorityName="De Man" authorityYear="1910" box="[1283,1429,367,393]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B93D9A3DA669CA272300FE9BFA6FE9A4" box="[1283,1429,367,393]" italics="true" pageId="9" pageNumber="347">S. bispinosus</emphasis>
</taxonomicName>
, four in
<taxonomicName id="4C493DACA669CA2726EBFE67FE55E980" authorityName="De Man" authorityYear="1910" box="[232,431,403,429]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B93D9A3DA669CA2726EBFE67FE55E980" box="[232,431,403,429]" italics="true" pageId="9" pageNumber="347">S. quadrispinosus</emphasis>
</taxonomicName>
and five in
<taxonomicName id="4C493DACA669CA272431FE67FD21E980" authorityName="Tattersall" authorityYear="1921" box="[562,731,403,429]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272431FE67FD21E980" box="[562,731,403,429]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C493DACA669CA272513FE67FC30E981" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[784,970,403,429]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA272513FE67FC30E981" box="[784,970,403,429]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E2746A669CA2725D3FE67FBF6E980" box="[976,1036,403,429]" pageId="9" pageNumber="347" rank="species">
<emphasis id="B93D9A3DA669CA2725D3FE67FBFDE980" bold="true" box="[976,1031,403,429]" pageId="9" pageNumber="347">n. sp</emphasis>
.
</taxonomicNameLabel>
), the members of the first subgroup are very similar to each other (cf.
<bibRefCitation id="EFD83BDEA669CA27240DFE43FD53E9FC" author="De Man, J. G." box="[526,681,439,465]" pageId="9" pageNumber="347" pagination="133 - 465" refId="ref4842" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition, Part II, family Alpheidae. Monographic, 39 a, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="book chapter" year="1911">De Man 1911</bibRefCitation>
;
<bibRefCitation id="EFD83BDEA669CA2724B0FE43FCA7E9FC" author="Tattersall, W. M." box="[691,861,439,465]" pageId="9" pageNumber="347" pagination="345 - 398" refId="ref5073" refString="Tattersall, W. M. (1921) Report on the stomatopoda and macrurous decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of the Linnean Society of London, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall 1921</bibRefCitation>
). Nevertheless,
<taxonomicName id="4C493DACA669CA27220DFE43FB32E9FD" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[1038,1224,439,465]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA27220DFE43FB32E9FD" box="[1038,1224,439,465]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E2746A669CA2722CDFE43FAF7E9FC" box="[1230,1293,439,465]" pageId="9" pageNumber="347" rank="species">
<emphasis id="B93D9A3DA669CA2722CDFE43FAFFE9FC" bold="true" box="[1230,1285,439,465]" pageId="9" pageNumber="347">n. sp</emphasis>
.
</taxonomicNameLabel>
can be separated from
<taxonomicName id="4C493DACA669CA272712FE2FFE58E9D8" authorityName="De Man" authorityYear="1910" box="[273,418,475,501]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B93D9A3DA669CA272712FE2FFE58E9D8" box="[273,418,475,501]" italics="true" pageId="9" pageNumber="347">S. bispinosus</emphasis>
</taxonomicName>
, in addition to the difference in the armature on the sixth pleonite, by disto-dorsal projection of the major cheliped palm which is less robust than
<taxonomicName id="4C493DACA669CA2724E6FE0BFC8DEA34" authorityName="De Man" authorityYear="1910" box="[741,887,511,537]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B93D9A3DA669CA2724E6FE0BFC8DEA34" box="[741,887,511,537]" italics="true" pageId="9" pageNumber="347">S. bispinosus</emphasis>
</taxonomicName>
. Furthermore, according to
<bibRefCitation id="EFD83BDEA669CA2722B2FE0BFAA5EA34" author="De Man, J. G." box="[1201,1375,511,537]" pageId="9" pageNumber="347" pagination="133 - 465" refId="ref4842" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition, Part II, family Alpheidae. Monographic, 39 a, 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="book chapter" year="1911">De Man (1911)</bibRefCitation>
, in
<taxonomicName id="4C493DACA669CA272384FE0BFEF7EA10" authorityName="De Man" authorityYear="1910" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B93D9A3DA669CA272384FE0BFEF7EA10" italics="true" pageId="9" pageNumber="347">S. bispinosus</emphasis>
</taxonomicName>
, the “outer margin of the dactylus [of the minor chela is] fringed with rather long, stiff setae from the articulation to the tip”. In contrast, in
<taxonomicName id="4C493DACA669CA272423FDB3FD21EA4D" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[544,731,583,609]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
S.
<emphasis id="B93D9A3DA669CA27243FFDB3FD21EA4D" box="[572,731,583,608]" italics="true" pageId="9" pageNumber="347">pentaspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E2746A669CA2724E2FDB3FCE4EA4C" box="[737,798,583,609]" pageId="9" pageNumber="347" rank="species">
<emphasis id="B93D9A3DA669CA2724E2FDB3FCEDEA4C" bold="true" box="[737,791,583,609]" pageId="9" pageNumber="347">n. sp</emphasis>
.
</taxonomicNameLabel>
, this margin bears two rows of setae. Similarly, the minor chela dactylus of
<taxonomicName id="4C493DACA669CA27275EFD9FFDDEEAA8" authorityName="De Man" authorityYear="1910" box="[349,548,619,645]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B93D9A3DA669CA27275EFD9FFDDEEAA8" box="[349,548,619,645]" italics="true" pageId="9" pageNumber="347">S. quadrispinosus</emphasis>
</taxonomicName>
seems to bear only one distinct row of setae, not two, as in the new species. In
<taxonomicName id="4C493DACA669CA272694FD7BFEA4EA84" authorityName="De Man" authorityYear="1910" box="[151,350,655,681]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B93D9A3DA669CA272694FD7BFEA4EA84" box="[151,350,655,681]" italics="true" pageId="9" pageNumber="347">S. quadrispinosus</emphasis>
</taxonomicName>
, the color of the major chela is pink in its distal half (
<bibRefCitation id="EFD83BDEA669CA2725BAFD7BFB2FEA84" author="Anker, A. &amp; De Grave, S." box="[953,1237,655,681]" pageId="9" pageNumber="347" pagination="343 - 454" refId="ref4479" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016</bibRefCitation>
: fig. 49), whilst it is brownish orange and brownish green in females and males of
<taxonomicName id="4C493DACA669CA27256BFD47FBD9EAE1" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[872,1059,691,717]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA27256BFD47FBD9EAE1" box="[872,1059,691,717]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA669CA272229FD47FB92EAE0" bold="true" box="[1066,1128,691,717]" pageId="9" pageNumber="347">
<taxonomicNameLabel id="A20E2746A669CA272229FD47FB92EAE0" box="[1066,1128,691,717]" pageId="9" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, respectively (
<figureCitation id="13725AAAA669CA27230EFD47FA7DEAE0" box="[1293,1415,691,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Synalpheus pentaspinosus n. sp., (A) paratype, male (CL 6.7 mm) from Larak Island, Iran (MNHN-IU-2014- 1223); (B) paratype, female (CL 8.4 mm) from the same locality (MNHN-IU-2014-1223); (C) allotype, female (CL 7.9 mm) from the same locality (MNHN-IU-2014-1222). Photographs by Rashed Abdollahi." figureDoi="http://doi.org/10.5281/zenodo.4416371" httpUri="https://zenodo.org/record/4416371/files/figure.png" pageId="9" pageNumber="347">Fig. 4A, B</figureCitation>
).
</paragraph>
<paragraph id="8BF6462FA669CA2726C4FD23FA6AECC4" blockId="9.[151,1437,151,1257]" pageId="9" pageNumber="347">
In having the same number of teeth (five) on the sixth pleonite,
<taxonomicName id="4C493DACA669CA2725A0FD23FB9AEADD" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[931,1120,727,753]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA2725A0FD23FB9AEADD" box="[931,1120,727,753]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA669CA27226BFD23FB52EADC" bold="true" box="[1128,1192,727,753]" pageId="9" pageNumber="347">
<taxonomicNameLabel id="A20E2746A669CA27226BFD23FB52EADC" box="[1128,1192,727,753]" pageId="9" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
appears to be closest to
<taxonomicName id="4C493DACA669CA2726B5FD0FFE9BEB38" authorityName="Tattersall" authorityYear="1921" box="[182,353,763,789]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA2726B5FD0FFE9BEB38" box="[182,353,763,789]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
(
<figureCitation id="13725AAAA669CA272771FD0FFE42EB38" box="[370,440,762,789]" captionStart="FIGURE 5" captionStartId="7.[151,250,1588,1612]" captionTargetBox="[151,1437,181,1564]" captionTargetId="figure-30@7.[151,1437,181,1564]" captionTargetPageId="7" captionText="FIGURE 5. Synalpheus quinquedens Tattersall, 1921, (A) male (CL 7.38 mm) from Farur Island, Iran (MNHN-IU-2014-1225); (B) female (CL 8.08 mm) from Farur Island, Iran (MNHN-IU-2014-1225). Photographs by Rashed Abdollahi." figureDoi="http://doi.org/10.5281/zenodo.4416373" httpUri="https://zenodo.org/record/4416373/files/figure.png" pageId="9" pageNumber="347">Fig. 5</figureCitation>
). However, the two species can be discriminated by some morphological characters, as well as by the color of the ovaries / eggs. In the new species, the minor chela dactylus is furnished with two rows of setae, one mesial row and one dorsal row, this configuration being different from that of
<taxonomicName id="4C493DACA669CA272279FCB7FADEEB70" authorityName="Tattersall" authorityYear="1921" box="[1146,1316,835,861]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272279FCB7FADEEB70" box="[1146,1316,835,861]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
, with only one prominent mesial row of setae (
<figureCitation id="13725AAAA669CA27242FFC93FD58EBAC" box="[556,674,871,897]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[166,1410,195,1912]" captionTargetId="figure-17@8.[155,1431,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Synalpheus quinquedens Tattersall, 1921, female from Farur Island, Iran (CL 12.3 mm) (MNHN-IU-2014-1225), (A) anterior region, dorsal view; (B) third maxilliped, ventrolateral view; (C) minor cheliped, mesial view; (D) same, palm and fingers, dorsomesial view." figureDoi="http://doi.org/10.5281/zenodo.4416375" httpUri="https://zenodo.org/record/4416375/files/figure.png" pageId="9" pageNumber="347">Fig 6C, D</figureCitation>
). In
<taxonomicName id="4C493DACA669CA2724D6FC93FC6AEBAD" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[725,912,871,897]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA2724D6FC93FC6AEBAD" box="[725,912,871,897]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA669CA27259BFC93FC2DEBAC" bold="true" box="[920,983,871,897]" pageId="9" pageNumber="347">
<taxonomicNameLabel id="A20E2746A669CA27259BFC93FC2DEBAC" box="[920,983,871,897]" pageId="9" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the minor chela dactylus bears a small, but rather obvious concavity on the cutting edge, which is not distinct in
<taxonomicName id="4C493DACA669CA2725BCFC7FFB93EB88" authorityName="Tattersall" authorityYear="1921" box="[959,1129,907,933]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA2725BCFC7FFB93EB88" box="[959,1129,907,933]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
. It is important to note here that
<bibRefCitation id="EFD83BDEA669CA2726CBFC5BFE6EEBE4" author="Tattersall, W. M." box="[200,404,943,969]" pageId="9" pageNumber="347" pagination="345 - 398" refId="ref5073" refString="Tattersall, W. M. (1921) Report on the stomatopoda and macrurous decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of the Linnean Society of London, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersalls (1921)</bibRefCitation>
figures of
<taxonomicName id="4C493DACA669CA272413FC5BFD41EBE4" authorityName="Tattersall" authorityYear="1921" box="[528,699,943,969]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272413FC5BFD41EBE4" box="[528,699,943,969]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
appear to be somewhat diagrammatic and may therefore be inaccurate. For instance, according to
<bibRefCitation id="EFD83BDEA669CA272416FC27FD30EBC0" author="Tattersall, W. M." box="[533,714,979,1005]" pageId="9" pageNumber="347" pagination="345 - 398" refId="ref5073" refString="Tattersall, W. M. (1921) Report on the stomatopoda and macrurous decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of the Linnean Society of London, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall (1921</bibRefCitation>
: pl. 28, fig. 1), the orbital teeth of the
<typeStatus id="54F2F88DA669CA272274FC27FB5DEBC0" box="[1143,1191,979,1005]" pageId="9" pageNumber="347">type</typeStatus>
of
<taxonomicName id="4C493DACA669CA2722CEFC27FA89EBC0" box="[1229,1395,979,1005]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquidens">
<emphasis id="B93D9A3DA669CA2722CEFC27FA89EBC0" box="[1229,1395,979,1005]" italics="true" pageId="9" pageNumber="347">S. quinquidens</emphasis>
</taxonomicName>
are very broad and in fact appear to be broader than in the new species. However, our examination of the material of
<taxonomicName id="4C493DACA669CA272384FC03FEDFEC18" authorityName="Tattersall" authorityYear="1921" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272384FC03FEDFEC18" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
from
<collectingCountry id="F35E06BFA669CA27276AFBEFFE6CEC18" box="[361,406,1051,1077]" name="Iran" pageId="9" pageNumber="347">Iran</collectingCountry>
and
<collectingCountry id="F35E06BFA669CA2727CEFBEFFDAEEC18" box="[461,596,1051,1077]" name="Madagascar" pageId="9" pageNumber="347">Madagascar</collectingCountry>
showed that this character cannot be used in discrimination of
<taxonomicName id="4C493DACA669CA27231BFBEFFF2DEC75" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA27231BFBEFFF2DEC75" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA669CA2726DCFBCBFEE5EC74" bold="true" box="[223,287,1087,1113]" pageId="9" pageNumber="347">
<taxonomicNameLabel id="A20E2746A669CA2726DCFBCBFEE5EC74" box="[223,287,1087,1113]" pageId="9" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C493DACA669CA272764FBCBFDE8EC74" authorityName="Tattersall" authorityYear="1921" box="[359,530,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272764FBCBFDE8EC74" box="[359,530,1087,1113]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
, as smaller specimens of the latter species bear relatively narrower orbital teeth (
<figureCitation id="13725AAAA669CA2726A3FB97FF03EC53" box="[160,249,1123,1150]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[166,1410,195,1912]" captionTargetId="figure-17@8.[155,1431,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Synalpheus quinquedens Tattersall, 1921, female from Farur Island, Iran (CL 12.3 mm) (MNHN-IU-2014-1225), (A) anterior region, dorsal view; (B) third maxilliped, ventrolateral view; (C) minor cheliped, mesial view; (D) same, palm and fingers, dorsomesial view." figureDoi="http://doi.org/10.5281/zenodo.4416375" httpUri="https://zenodo.org/record/4416375/files/figure.png" pageId="9" pageNumber="347">Fig. 6A</figureCitation>
). In life,
<taxonomicName id="4C493DACA669CA27275DFB97FDE3EC51" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[350,537,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA27275DFB97FDE3EC51" box="[350,537,1123,1149]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA669CA272423FB97FDA4EC50" bold="true" box="[544,606,1123,1149]" pageId="9" pageNumber="347">
<taxonomicNameLabel id="A20E2746A669CA272423FB97FDA4EC50" box="[544,606,1123,1149]" pageId="9" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C493DACA669CA272496FB97FCC5EC50" authorityName="Tattersall" authorityYear="1921" box="[661,831,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272496FB97FCC5EC50" box="[661,831,1123,1149]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
can be distinguished by the color of ovaries, eggs and possibly also by corneal pigments. Females of
<taxonomicName id="4C493DACA669CA27249FFB73FCACEC8D" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[668,854,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA27249FFB73FCACEC8D" box="[668,854,1159,1185]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA669CA27255FFB73FC60EC8C" bold="true" box="[860,922,1159,1185]" pageId="9" pageNumber="347">
<taxonomicNameLabel id="A20E2746A669CA27255FFB73FC60EC8C" box="[860,922,1159,1185]" pageId="9" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
show yellow to yellow-orange ovaries and orange-red eggs (
<figureCitation id="13725AAAA669CA272742FB5FFE42ECE8" box="[321,440,1195,1221]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Synalpheus pentaspinosus n. sp., (A) paratype, male (CL 6.7 mm) from Larak Island, Iran (MNHN-IU-2014- 1223); (B) paratype, female (CL 8.4 mm) from the same locality (MNHN-IU-2014-1223); (C) allotype, female (CL 7.9 mm) from the same locality (MNHN-IU-2014-1222). Photographs by Rashed Abdollahi." figureDoi="http://doi.org/10.5281/zenodo.4416371" httpUri="https://zenodo.org/record/4416371/files/figure.png" pageId="9" pageNumber="347">Fig. 4B, C</figureCitation>
), whilst females of
<taxonomicName id="4C493DACA669CA272490FB5FFCC6ECE8" authorityName="Tattersall" authorityYear="1921" box="[659,828,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272490FB5FFCC6ECE8" box="[659,828,1195,1221]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
have greenish ovaries and eggs (
<figureCitation id="13725AAAA669CA2722A9FB5FFAF8ECE8" box="[1194,1282,1194,1221]" captionStart="FIGURE 5" captionStartId="7.[151,250,1588,1612]" captionTargetBox="[151,1437,181,1564]" captionTargetId="figure-30@7.[151,1437,181,1564]" captionTargetPageId="7" captionText="FIGURE 5. Synalpheus quinquedens Tattersall, 1921, (A) male (CL 7.38 mm) from Farur Island, Iran (MNHN-IU-2014-1225); (B) female (CL 8.08 mm) from Farur Island, Iran (MNHN-IU-2014-1225). Photographs by Rashed Abdollahi." figureDoi="http://doi.org/10.5281/zenodo.4416373" httpUri="https://zenodo.org/record/4416373/files/figure.png" pageId="9" pageNumber="347">Fig. 5B</figureCitation>
). The corneas of
<taxonomicName id="4C493DACA669CA2726B5FB3BFE8BECC5" authority="Ashrafi &amp; Sari &amp; Naderloo, 2020" authorityName="Ashrafi &amp; Sari &amp; Naderloo" authorityYear="2020" box="[182,369,1231,1257]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaspinosus" status="sp. nov.">
<emphasis id="B93D9A3DA669CA2726B5FB3BFE8BECC5" box="[182,369,1231,1257]" italics="true" pageId="9" pageNumber="347">S. pentaspinosus</emphasis>
</taxonomicName>
<emphasis id="B93D9A3DA669CA272774FB3BFE4CECC4" bold="true" box="[375,438,1231,1257]" pageId="9" pageNumber="347">
<taxonomicNameLabel id="A20E2746A669CA272774FB3BFE4CECC4" box="[375,438,1231,1257]" pageId="9" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
vary from pinkish to reddish (
<figureCitation id="13725AAAA669CA272508FB3BFCAAECC4" box="[779,848,1231,1257]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Synalpheus pentaspinosus n. sp., (A) paratype, male (CL 6.7 mm) from Larak Island, Iran (MNHN-IU-2014- 1223); (B) paratype, female (CL 8.4 mm) from the same locality (MNHN-IU-2014-1223); (C) allotype, female (CL 7.9 mm) from the same locality (MNHN-IU-2014-1222). Photographs by Rashed Abdollahi." figureDoi="http://doi.org/10.5281/zenodo.4416371" httpUri="https://zenodo.org/record/4416371/files/figure.png" pageId="9" pageNumber="347">Fig. 4</figureCitation>
), being darker brownish in
<taxonomicName id="4C493DACA669CA272286FB3BFAD5ECC4" authorityName="Tattersall" authorityYear="1921" box="[1157,1327,1231,1257]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="347" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B93D9A3DA669CA272286FB3BFAD5ECC4" box="[1157,1327,1231,1257]" italics="true" pageId="9" pageNumber="347">S. quinquedens</emphasis>
</taxonomicName>
(
<figureCitation id="13725AAAA669CA27233DFB3BFA78ECC4" box="[1342,1410,1230,1257]" captionStart="FIGURE 5" captionStartId="7.[151,250,1588,1612]" captionTargetBox="[151,1437,181,1564]" captionTargetId="figure-30@7.[151,1437,181,1564]" captionTargetPageId="7" captionText="FIGURE 5. Synalpheus quinquedens Tattersall, 1921, (A) male (CL 7.38 mm) from Farur Island, Iran (MNHN-IU-2014-1225); (B) female (CL 8.08 mm) from Farur Island, Iran (MNHN-IU-2014-1225). Photographs by Rashed Abdollahi." figureDoi="http://doi.org/10.5281/zenodo.4416373" httpUri="https://zenodo.org/record/4416373/files/figure.png" pageId="9" pageNumber="347">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>