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<document id="B9E1142F54B92752FF5FDAD102571641" ID-CLB-Dataset="124672" ID-DOI="10.11646/zootaxa.5175.3.4" ID-GBIF-Dataset="c6e5ff11-f11b-4b27-8510-b3f173bc6888" ID-ISSN="1175-5326" ID-Zenodo-Dep="7003688" ID-ZooBank="80B8E445-EB53-418F-9DEC-5654FF0EBCEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660737329920" checkinUser="plazi" docAuthor="Makarkin, Vladimir N." docDate="2022" docId="03E07209FFFEFF862CAEFD40389518F7" docLanguage="en" docName="zootaxa.5175.3.4.pdf" docOrigin="Zootaxa 5175 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Olindanymphes Martins-Neto 2005" docType="treatment" docVersion="8" lastPageNumber="380" masterDocId="FFD90A71FFFFFF852C39FFBA3A071A72" masterDocTitle="A new important species of Nymphidae (Neuroptera) from the Lower Cretaceous Crato Formation of Brazil" masterLastPageNumber="382" masterPageNumber="377" pageNumber="378" updateTime="1699367368640" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9576718E4ACC0557652733B030DDE8BD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A558BD3E80CCE386C56444079CEFC377">
<mods:title id="308ABA6DA0A5900692F33239EE4C7C2E">A new important species of Nymphidae (Neuroptera) from the Lower Cretaceous Crato Formation of Brazil</mods:title>
</mods:titleInfo>
<mods:name id="EC80CD8C875EE29854E578C92E911E71" type="personal">
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<mods:roleTerm id="4F7EF74ECA9DBBA955D2D3E65011B673">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2AD230C19C862E075A5B02F0775991A1">Makarkin, Vladimir N.</mods:namePart>
</mods:name>
<mods:typeOfResource id="26DA6CB16248BA8E951D20AABEEB4040">text</mods:typeOfResource>
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<mods:title id="AFC6B7ED888D25A36B4C87D1C6221D8F">Zootaxa</mods:title>
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<mods:part id="F189187DD190E1DF269420AA5D5793C9">
<mods:date id="A6ED9AD553887ACFDCABFB47613D6902">2022</mods:date>
<mods:detail id="F4C5A85FF247B99A4AB7A4EBFB30AA4B" type="pubDate">
<mods:number id="D825F74D0D82CF2186E03F674845F70A">2022-08-17</mods:number>
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<mods:number id="FAA6EFCEB9B94975FEE4EDA37EFB837D">5175</mods:number>
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<mods:number id="CF2726FCAB10CD8A13E43E47EBA97111">3</mods:number>
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<mods:classification id="A940CC6C615E6DCC5FB9DB4D8F324259">journal article</mods:classification>
<mods:identifier id="A944EDC96B98272E206CB1B0C66BEE3C" type="CLB-Dataset">124672</mods:identifier>
<mods:identifier id="70D17F6D593115946E7A3BACF6E6C415" type="DOI">10.11646/zootaxa.5175.3.4</mods:identifier>
<mods:identifier id="0F4EC5572B93E7061CC6B8ACDB0C76A2" type="GBIF-Dataset">c6e5ff11-f11b-4b27-8510-b3f173bc6888</mods:identifier>
<mods:identifier id="C3E1CC028B99E2E8F83C00D3B4721A1B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E07209FFFEFF862CAEFD40389518F7" ID-DOI="http://doi.org/10.5281/zenodo.7003557" ID-GBIF-Taxon="199505136" ID-Zenodo-Dep="7003557" LSID="urn:lsid:plazi:treatment:03E07209FFFEFF862CAEFD40389518F7" httpUri="http://treatment.plazi.org/id/03E07209FFFEFF862CAEFD40389518F7" lastPageId="3" lastPageNumber="380" pageId="1" pageNumber="378">
<subSubSection id="C3539094FFFEFF842CAEFD4038271966" box="[151,544,762,789]" pageId="1" pageNumber="378" type="nomenclature">
<paragraph id="8BF6C31FFFFEFF842CAEFD4038271966" blockId="1.[151,544,762,825]" box="[151,544,762,789]" pageId="1" pageNumber="378">
<heading id="D0BE7473FFFEFF842CAEFD4038271966" bold="true" box="[151,544,762,789]" fontSize="11" level="1" pageId="1" pageNumber="378" reason="1">
<emphasis id="B93D1F0DFFFEFF842CAEFD40381D1967" bold="true" box="[151,538,762,789]" italics="true" pageId="1" pageNumber="378">
<taxonomicName id="4C49B89CFFFEFF842CAEFD403B5F1967" ID-CoL="67S5" authorityName="Martins-Neto" authorityYear="2005" box="[151,344,762,789]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="genus">Olindanymphes</taxonomicName>
? headsi sp. nov
</emphasis>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3539094FFFEFF842CAEFCA53AFF194B" box="[151,248,799,825]" pageId="1" pageNumber="378" type="description">
<paragraph id="8BF6C31FFFFEFF842CAEFCA53AFF194B" blockId="1.[151,544,762,825]" box="[151,248,799,825]" pageId="1" pageNumber="378">
<figureCitation id="1372DF9AFFFEFF842CAEFCA53AFF194B" box="[151,248,799,825]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1981,2005]" captionStartId-1="2.[151,250,1205,1229]" captionStartId-2="2.[151,250,1722,1746]" captionTargetBox-0="[239,1348,849,1958]" captionTargetBox-1="[177,1410,669,1182]" captionTargetBox-2="[157,1431,1298,1690]" captionTargetId-0="figure-111@1.[239,1348,849,1958]" captionTargetId-1="figure-216@2.[177,1410,669,1182]" captionTargetId-2="figure-261@2.[151,1436,1289,1698]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. Olindanymphes? headsi sp. nov., holotype INHSP 1510 (wetted with ethanol). Scale bar = 5 mm." captionText-1="FIGURE 2. Olindanymphes? headsi sp. nov., holotype INHSP 1510 (wetted with ethanol). A, head, pronotum (dorsal view); B, apex of abdomen (lateral view?). e, eye; he, head; pr, pronotum; 9T, ninth tergite. Scale bars = 1 mm." captionText-2="FIGURE 3. Hindwing venation of Olindanymphes? headsi sp. nov., holotype INHSP 1510. Scale bar = 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7003690" figureDoi-1="http://doi.org/10.5281/zenodo.7003692" figureDoi-2="http://doi.org/10.5281/zenodo.7003694" httpUri-0="https://zenodo.org/record/7003690/files/figure.png" httpUri-1="https://zenodo.org/record/7003692/files/figure.png" httpUri-2="https://zenodo.org/record/7003694/files/figure.png" pageId="1" pageNumber="378">Figs 1–3</figureCitation>
</paragraph>
</subSubSection>
<caption id="DF369397FFFEFF842CAEF8073EFF1DA7" ID-DOI="http://doi.org/10.5281/zenodo.7003690" ID-Zenodo-Dep="7003690" box="[151,1272,1981,2005]" httpUri="https://zenodo.org/record/7003690/files/figure.png" pageId="1" pageNumber="378" startId="1.[151,250,1981,2005]" targetBox="[239,1348,849,1958]" targetPageId="1">
<paragraph id="8BF6C31FFFFEFF842CAEF8073EFF1DA7" blockId="1.[151,1272,1981,2005]" box="[151,1272,1981,2005]" pageId="1" pageNumber="378">
<emphasis id="B93D1F0DFFFEFF842CAEF8073B141DA7" bold="true" box="[151,275,1981,2005]" pageId="1" pageNumber="378">FIGURE 1.</emphasis>
<emphasis id="B93D1F0DFFFEFF842D20F807380A1DA7" box="[281,525,1981,2005]" italics="true" pageId="1" pageNumber="378">
<taxonomicName id="4C49B89CFFFEFF842D20F8073BBC1DA7" authorityName="Martins-Neto" authorityYear="2005" box="[281,443,1981,2005]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="genus">Olindanymphes</taxonomicName>
? headsi
</emphasis>
<emphasis id="B93D1F0DFFFEFF842E2DF80738631DA7" bold="true" box="[532,612,1981,2005]" pageId="1" pageNumber="378">sp. nov.</emphasis>
, holotype INHSP 1510 (wetted with ethanol). Scale bar = 5 mm.
</paragraph>
</caption>
<subSubSection id="C3539094FFFDFF872CAEFF2D39931B17" pageId="2" pageNumber="379" type="materials_examined">
<paragraph id="8BF6C31FFFFDFF872CAEFF2D3B9D1B6F" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="379">
<emphasis id="B93D1F0DFFFDFF872CAEFF2D3B4F1AC2" bold="true" box="[151,328,150,177]" pageId="2" pageNumber="379">Type material.</emphasis>
<materialsCitation id="3B21C942FFFDFF872D68FF2D3B9D1B6F" ID-GBIF-Occurrence="3890522302" collectorName="S. W. Heads" country="United States of America" location="Illinois" pageId="2" pageNumber="379" specimenCount="1" stateProvince="Illinois" typeStatus="holotype">
<typeStatus id="54F27DBDFFFDFF872D68FF2D3BB91AC3" box="[337,446,151,177]" pageId="2" pageNumber="379" type="holotype">Holotype</typeStatus>
: INHSP 1510, currently deposited in the
<collectingRegion id="498D0DFDFFFDFF872FA2FF2D39ED1AC3" box="[923,1002,151,177]" country="United States of America" name="Illinois" pageId="2" pageNumber="379">Illinois</collectingRegion>
Natural History Survey Paleontology Collection, University of
<collectingRegion id="498D0DFDFFFDFF872D8EFF0138011AA7" box="[439,518,187,213]" country="United States of America" name="Illinois" pageId="2" pageNumber="379">Illinois</collectingRegion>
at Urbana-Champaign,
<collectingCountry id="F35E838FFFFDFF872F2CFF0139641AA7" box="[789,867,187,213]" name="United States of America" pageId="2" pageNumber="379">U.S.A.</collectingCountry>
, but will be repatriated to an appropriate Brazilian institution to comply with Brazilian laws (
<collectorName id="26BCA6C9FFFDFF872EBDFF65390A1A8B" box="[644,781,223,249]" pageId="2" pageNumber="379">S.W. Heads</collectorName>
, pers. comm.). A body with one complete hindwing and fragments of hind legs.
</materialsCitation>
</paragraph>
<paragraph id="8BF6C31FFFFDFF872CFEFE9D39931B17" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="379">
<materialsCitation id="3B21C942FFFDFF872CFEFE9D39971B17" ID-GBIF-Occurrence="3890522301" country="Brazil" county="upper Aptian" location="Northeastern" municipality="Lower Cretaceous" pageId="2" pageNumber="379" specimenCount="1" typeStatus="holotype">
<emphasis id="B93D1F0DFFFDFF872CFEFE9D38081B32" bold="true" box="[199,527,294,321]" pageId="2" pageNumber="379">
<typeStatus id="54F27DBDFFFDFF872CFEFE9D3B051B33" box="[199,258,295,321]" pageId="2" pageNumber="379">Type</typeStatus>
locality and horizon.
</emphasis>
<location id="8E9695C4FFFDFF872E22FE9D38AB1B33" LSID="urn:lsid:plazi:treatment:03E07209FFFEFF862CAEFD40389518F7:8E9695C4FFFDFF872E22FE9D38AB1B33" box="[539,684,295,321]" country="Brazil" county="upper Aptian" municipality="Lower Cretaceous" name="Northeastern" pageId="2" pageNumber="379">Northeastern</location>
<collectingCountry id="F35E838FFFFDFF872E81FE9D39061B33" box="[696,769,295,321]" name="Brazil" pageId="2" pageNumber="379">Brazil</collectingCountry>
:
<location id="8E9695C4FFFDFF872F28FE9D39FB1B33" LSID="urn:lsid:plazi:treatment:03E07209FFFEFF862CAEFD40389518F7:8E9695C4FFFDFF872F28FE9D39FB1B33" box="[785,1020,295,321]" country="Brazil" county="upper Aptian" municipality="Lower Cretaceous" name="Chapada do Araripe" pageId="2" pageNumber="379">Chapada do Araripe</location>
(precise locality unknown).
<collectingMunicipality id="6B925965FFFDFF87296AFE9D3B1F1B17" pageId="2" pageNumber="379">Lower Cretaceous</collectingMunicipality>
:
<collectingCounty id="6297BB93FFFDFF872D1BFEF13BB31B17" box="[290,436,331,357]" pageId="2" pageNumber="379">upper Aptian</collectingCounty>
(Crato Formation: Nova Olinda Member)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3539094FFFDFF872CFEFED538111BDF" pageId="2" pageNumber="379" type="etymology">
<paragraph id="8BF6C31FFFFDFF872CFEFED538111BDF" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="379">
<emphasis id="B93D1F0DFFFDFF872CFEFED53B491BFB" bold="true" box="[199,334,367,393]" pageId="2" pageNumber="379">Etymology.</emphasis>
The specific epithet is formed from the surname of Sam W. Heads in recognition of his contributions to the study of Cretaceous insects.
</paragraph>
</subSubSection>
<subSubSection id="C3539094FFFDFF872CFEFE0C3E5C1B86" pageId="2" pageNumber="379" type="diagnosis">
<paragraph id="8BF6C31FFFFDFF872CFEFE0C3E5C1B86" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="379">
<emphasis id="B93D1F0DFFFDFF872CFEFE0C3B451BA2" bold="true" box="[199,322,438,464]" pageId="2" pageNumber="379">Diagnosis.</emphasis>
Differs from
<taxonomicName id="4C49B89CFFFDFF872DDAFE0D39081BA3" authorityName="Martins-Neto" authorityYear="2005" box="[483,783,439,465]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFDFF872DDAFE0D39081BA3" box="[483,783,439,465]" italics="true" pageId="2" pageNumber="379">Olindanymphes makarkini</emphasis>
</taxonomicName>
by absence of crossveins between distal branches of
<collectingRegion id="498D0DFDFFFDFF872943FE0D3F9B1BA3" box="[1402,1436,439,465]" country="Brazil" name="Para" pageId="2" pageNumber="379">PR</collectingRegion>
[present in
<taxonomicName id="4C49B89CFFFDFF872D2CFE613BAE1B87" authorityName="Martins-Neto" authorityYear="2005" box="[277,425,475,501]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFDFF872D2CFE613BAE1B87" box="[277,425,475,501]" italics="true" pageId="2" pageNumber="379">O. makarkini</emphasis>
</taxonomicName>
], and more acute wing apex [more rounded in
<taxonomicName id="4C49B89CFFFDFF872F8FFE613E4C1B87" authorityName="Martins-Neto" authorityYear="2005" box="[950,1099,475,501]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFDFF872F8FFE613E4C1B87" box="[950,1099,475,501]" italics="true" pageId="2" pageNumber="379">O. makarkini</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C3539094FFFDFF862CFEFE44390A1B33" lastPageId="3" lastPageNumber="380" pageId="2" pageNumber="379" type="description">
<paragraph id="8BF6C31FFFFDFF872CFEFE4439D218F7" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="379">
<emphasis id="B93D1F0DFFFDFF872CFEFE443B5E186A" bold="true" box="[199,345,510,536]" pageId="2" pageNumber="379">Description.</emphasis>
Female (?). Body
<emphasis id="B93D1F0DFFFDFF872E09FE45384D186A" box="[560,586,511,536]" italics="true" pageId="2" pageNumber="379">ca</emphasis>
.
<quantity id="4CB16EFAFFFDFF872E63FE4538AD186B" box="[602,682,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="379" unit="mm" value="27.0">27 mm</quantity>
long; poorly preserved. Details of head not discernible. Pronotum transverse in dorsal view (
<emphasis id="B93D1F0DFFFDFF872D87FD993BDF184E" box="[446,472,547,572]" italics="true" pageId="2" pageNumber="379">ca</emphasis>
. 0.6 as long as wide); covered with dense, probably long setae. Meso- and metanotum dark. Abdomen: apical segments covered with long, dense setae; presumable 9th tergite not extending laterally to ventral part of abdomen, not dilated ventrally (if it is seen in lateral view).
</paragraph>
<caption id="DF369397FFFDFF872CAEFB0F3EB11E83" ID-DOI="http://doi.org/10.5281/zenodo.7003692" ID-Zenodo-Dep="7003692" httpUri="https://zenodo.org/record/7003692/files/figure.png" pageId="2" pageNumber="379" startId="2.[151,250,1205,1229]" targetBox="[177,1410,669,1182]" targetPageId="2">
<paragraph id="8BF6C31FFFFDFF872CAEFB0F3EB11E83" blockId="2.[151,1436,1205,1266]" pageId="2" pageNumber="379">
<emphasis id="B93D1F0DFFFDFF872CAEFB0F3B131EBF" bold="true" box="[151,276,1205,1229]" pageId="2" pageNumber="379">FIGURE 2.</emphasis>
<emphasis id="B93D1F0DFFFDFF872D22FB0F38171EBF" box="[283,528,1205,1229]" italics="true" pageId="2" pageNumber="379">
<taxonomicName id="4C49B89CFFFDFF872D22FB0F3BBA1EBF" authorityName="Martins-Neto" authorityYear="2005" box="[283,445,1205,1229]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">Olindanymphes</taxonomicName>
? headsi
</emphasis>
<emphasis id="B93D1F0DFFFDFF872E2EFB0F386E1EBF" bold="true" box="[535,617,1205,1229]" pageId="2" pageNumber="379">sp. nov.</emphasis>
, holotype INHSP 1510 (wetted with ethanol). A, head, pronotum (dorsal view); B, apex of abdomen (lateral view?). e, eye; he, head; pr, pronotum; 9T, ninth tergite. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF369397FFFDFF872CAEF9003EFA1CA0" ID-DOI="http://doi.org/10.5281/zenodo.7003694" ID-Zenodo-Dep="7003694" box="[151,1277,1722,1746]" httpUri="https://zenodo.org/record/7003694/files/figure.png" pageId="2" pageNumber="379" startId="2.[151,250,1722,1746]" targetBox="[157,1431,1298,1690]" targetPageId="2">
<paragraph id="8BF6C31FFFFDFF872CAEF9003EFA1CA0" blockId="2.[151,1277,1722,1746]" box="[151,1277,1722,1746]" pageId="2" pageNumber="379">
<emphasis id="B93D1F0DFFFDFF872CAEF9003B141CA0" bold="true" box="[151,275,1722,1746]" pageId="2" pageNumber="379">FIGURE 3.</emphasis>
Hindwing venation of
<emphasis id="B93D1F0DFFFDFF872DC7F90038F51CA0" box="[510,754,1722,1746]" italics="true" pageId="2" pageNumber="379">
<taxonomicName id="4C49B89CFFFDFF872DC7F90038A71CA0" authorityName="Martins-Neto" authorityYear="2005" box="[510,672,1722,1746]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">Olindanymphes</taxonomicName>
? headsi
</emphasis>
<emphasis id="B93D1F0DFFFDFF872EC0F900394E1CA0" bold="true" box="[761,841,1722,1746]" pageId="2" pageNumber="379">sp. nov.</emphasis>
, holotype INHSP 1510. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8BF6C31FFFFDFF862CFEF8BA390A1B33" blockId="2.[151,1437,1792,2035]" lastBlockId="3.[151,1437,150,645]" lastPageId="3" lastPageNumber="380" pageId="2" pageNumber="379">
Hindwing elongate with sub-acute apex,
<emphasis id="B93D1F0DFFFDFF872EA1F8BB38B51D68" box="[664,690,1793,1818]" italics="true" pageId="2" pageNumber="379">ca</emphasis>
.
<quantity id="4CB16EFAFFFDFF872EF8F8BA39161D69" box="[705,785,1792,1819]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="379" unit="mm" value="26.0">26 mm</quantity>
as preserved (estimated complete length
<quantity id="4CB16EFAFFFDFF8728DEF8BB3F621D69" box="[1255,1381,1793,1819]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="2" pageNumber="379" unit="mm" value="27.5" valueMax="28.0" valueMin="27.0">27‒28 mm</quantity>
),
<quantity id="4CB16EFAFFFDFF872940F8BB3AC41D4D" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="379" unit="mm" value="7.1">7.1 mm</quantity>
wide. Costal space relatively narrow. All preserved subcostal veinlets simple. Subcostal space relatively broad: no crossveins detected. RP originates relatively close to wing base at acute angle to RA, with 10 pectinate branches; RP1 twice very shallowly forked; most other branches dichotomously forked. Crossveins between branches of RP irregularly spaced; between distal branches absent, becoming denser proximad (seven detected between RP1, RP2). Ten crossveins between RP/RP1 and MA. Anterior trace of MA slightly convex, with three rather short branches, which once or twice shallowly forked. Crossveins between MA, MP numerous (11 detected). MP with six pectinate branches; MP1, MP3 deeply forked; MP2, MP4, MP5 relatively shallowly forked; RP1 simple. One to two crossveins between branches of MP. Anterior trace of CuA terminally not forked, pectinately forked, with three long simple branches; no crossvein between these branches not detected. Anterior trace of CuP terminally not forked, pectinately branched, with two long simple branches. Presumable A1 deeply forked. Crossveins in basal part of wing indiscernible or absent. Trichosors not detected.
</paragraph>
</subSubSection>
<subSubSection id="C3539094FFFCFF862CFEFEF1389518F7" pageId="3" pageNumber="380" type="discussion">
<paragraph id="8BF6C31FFFFCFF862CFEFEF13EE5184E" blockId="3.[151,1437,150,645]" pageId="3" pageNumber="380">
<emphasis id="B93D1F0DFFFCFF862CFEFEF13B331B17" bold="true" box="[199,308,331,357]" pageId="3" pageNumber="380">Remarks</emphasis>
. Crossvein arrangement and the shape of wing apex in most
<taxonomicName id="4C49B89CFFFCFF86282DFEF13E901B17" authorityName="Rambur" authorityYear="1842" box="[1044,1175,331,357]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
(except some derived Myiodactylinae) are more or less similar in fore- and hindwings (see
<emphasis id="B93D1F0DFFFCFF862F9EFED539C81BFA" box="[935,975,367,392]" italics="true" pageId="3" pageNumber="380">e.g.</emphasis>
, She
<emphasis id="B93D1F0DFFFCFF862829FED53E451BFB" box="[1040,1090,367,393]" italics="true" pageId="3" pageNumber="380">et al</emphasis>
. 2013: Fig. 8; She
<emphasis id="B93D1F0DFFFCFF862925FED53F521BFB" box="[1308,1365,367,393]" italics="true" pageId="3" pageNumber="380">et al.</emphasis>
2015:
<figureCitation id="1372DF9AFFFCFF862CAEFE293AD81BDF" box="[151,223,403,429]" captionStart="FIGURE 2" captionStartId="2.[151,250,1205,1229]" captionTargetBox="[177,1410,669,1182]" captionTargetId="figure-216@2.[177,1410,669,1182]" captionTargetPageId="2" captionText="FIGURE 2. Olindanymphes? headsi sp. nov., holotype INHSP 1510 (wetted with ethanol). A, head, pronotum (dorsal view); B, apex of abdomen (lateral view?). e, eye; he, head; pr, pronotum; 9T, ninth tergite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7003692" httpUri="https://zenodo.org/record/7003692/files/figure.png" pageId="3" pageNumber="380">Figs 2</figureCitation>
, 11, 15). It can therefore, reasonably be assumed that these character states in the undescribed hindwing of
<taxonomicName id="4C49B89CFFFCFF862CAEFE0D3BB81BA3" authorityName="Martins-Neto" authorityYear="2005" box="[151,447,439,465]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFCFF862CAEFE0D3BB81BA3" box="[151,447,439,465]" italics="true" pageId="3" pageNumber="380">Olindanymphes makarkini</emphasis>
</taxonomicName>
are more or less similar to those of its described forewing. These features in
<taxonomicName id="4C49B89CFFFCFF862933FE0D3F9A1BA3" authorityName="Martins-Neto" authorityYear="2005" box="[1290,1437,439,465]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFCFF862933FE0D3F9A1BA3" box="[1290,1437,439,465]" italics="true" pageId="3" pageNumber="380">O. makarkini</emphasis>
</taxonomicName>
and
<emphasis id="B93D1F0DFFFCFF862CF0FE613B431B87" box="[201,324,475,501]" italics="true" pageId="3" pageNumber="380">O.? headsi</emphasis>
<emphasis id="B93D1F0DFFFCFF862D74FE613BAF1B87" bold="true" box="[333,424,475,501]" pageId="3" pageNumber="380">sp. nov.</emphasis>
are different: crossveins are present between distal branches of
<collectingRegion id="498D0DFDFFFCFF8628B0FE613EAC1B87" box="[1161,1195,475,501]" country="Brazil" name="Para" pageId="3" pageNumber="380">PR</collectingRegion>
in
<taxonomicName id="4C49B89CFFFCFF8628EAFE613F6E1B87" authorityName="Martins-Neto" authorityYear="2005" box="[1235,1385,475,501]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFCFF8628EAFE613F6E1B87" box="[1235,1385,475,501]" italics="true" pageId="3" pageNumber="380">O. makarkini</emphasis>
</taxonomicName>
, but absent in
<emphasis id="B93D1F0DFFFCFF862D3CFE453B87186B" box="[261,384,511,537]" italics="true" pageId="3" pageNumber="380">O.? headsi</emphasis>
<emphasis id="B93D1F0DFFFCFF862DB0FE453BE3186B" bold="true" box="[393,484,511,537]" pageId="3" pageNumber="380">sp. nov.</emphasis>
and crossvenation is generally denser in the radial space of
<taxonomicName id="4C49B89CFFFCFF8628ADFE453F2D186B" authorityName="Martins-Neto" authorityYear="2005" box="[1172,1322,511,537]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFCFF8628ADFE453F2D186B" box="[1172,1322,511,537]" italics="true" pageId="3" pageNumber="380">O. makarkini</emphasis>
</taxonomicName>
; the wing apex of
<taxonomicName id="4C49B89CFFFCFF862CC8FD993B81184F" authorityName="Martins-Neto" authorityYear="2005" box="[241,390,547,573]" class="Insecta" family="Nymphidae" genus="Olindanymphes" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="makarkini">
<emphasis id="B93D1F0DFFFCFF862CC8FD993B81184F" box="[241,390,547,573]" italics="true" pageId="3" pageNumber="380">O. makarkini</emphasis>
</taxonomicName>
is obviously more rounded than in
<emphasis id="B93D1F0DFFFCFF862F2AFD99398B184F" box="[787,908,547,573]" italics="true" pageId="3" pageNumber="380">O.? headsi</emphasis>
<emphasis id="B93D1F0DFFFCFF862FAAFD9939EB184F" bold="true" box="[915,1004,547,573]" pageId="3" pageNumber="380">sp. nov.</emphasis>
, which is more acute.
</paragraph>
<paragraph id="8BF6C31FFFFCFF862CFEFDFD389518F7" blockId="3.[151,1437,150,645]" pageId="3" pageNumber="380">This specimen is most probably a female judging from the general view of apical segments of the abdomen, although their structures are mostly not clear.</paragraph>
</subSubSection>
</treatment>
</document>
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