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<mods:title id="F1EB626F67DA0C54C0D7C4722F3622BC">Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #</mods:title>
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<heading id="D08881CDA73FFFB53A88F996BA654E59" bold="true" box="[907,1403,1552,1577]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="113" reason="2">
<taxonomicName id="4C7F4D22A73FFFB53A88F996BA654E59" ID-CoL="3SYMC" authority="(Redtenbacher, 1867)" baseAuthorityName="Redtenbacher" baseAuthorityYear="1867" box="[907,1403,1552,1577]" class="Insecta" family="Dytiscidae" genus="Leiodytes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="113" phylum="Arthropoda" rank="species" species="nicobaricus">
<emphasis id="B90BEAB3A73FFFB53A88F996BA654E59" bold="true" box="[907,1403,1552,1577]" pageId="16" pageNumber="113">
<emphasis id="B90BEAB3A73FFFB53A88F996BB624E58" bold="true" box="[907,1148,1553,1577]" italics="true" pageId="16" pageNumber="113">Leiodytes nicobaricus</emphasis>
(
<bibRefCitation id="EFEE4B50A73FFFB53D88F997BA6D4E59" author="Redtenbacher, L" box="[1163,1395,1552,1576]" pageId="16" pageNumber="113" refId="ref40351" refString="Redtenbacher, L., 1867. Coleopteren. In: Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Mullerdsdorf-Vobair. Akademie der Wissenschaften Wien, Karl Gerold's Sohn, iv + 249 + 5 pp." type="book" year="1867">Redtenbacher, 1867</bibRefCitation>
)
</emphasis>
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(
<figureCitation id="13442A24A73FFFB53D58F9B6BBB44E39" box="[1115,1194,1585,1608]" captionStart="Figs" captionStartId="15.[113,158,2006,2027]" captionTargetBox="[253,1312,191,1946]" captionTargetPageId="15" captionText="Figs. 18-23. 18.Hydaticus fabricii (MacLeay, 1825) (Fam. Dytiscidae, 10.5 mm); 19. Leiodytes nicobaricus (Redtenbacher, 1867) (Fam. Dytiscidae, 2.1 mm); 20. Hydrovatus rufoniger rufoniger (Clark, 1863) (Fam. Dytiscidae, 3.6 mm); 21. Laccophilus medialis Sharp, 1882 (Fam. Dytiscidae, 3.3 mm); 22. Copelatus minutissimus Balfour-Browne, 1939 (Fam. Dytiscidae, 4.1 mm); 23. Eretes griseus (Fabricius, 1781) (Fam. Dytiscidae, 13.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13244912" httpUri="https://zenodo.org/record/13244912/files/figure.png" pageId="16" pageNumber="113">Fig. 19</figureCitation>
)
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<treatmentCitationGroup id="AB6F118FA73FFFB53A40F9EABADD4EF3" box="[835,1475,1645,1666]" pageId="16" pageNumber="113">
<treatmentCitation id="0ADE10B0A73FFFB53A40F9EABA0C4EF3" author="Redtenbacher, L" box="[835,1298,1645,1666]" page="21" pageId="16" pageNumber="113" year="1867">
<taxonomicName id="4C7F4D22A73FFFB53A40F9EABA0C4EF3" authority="Redtenbacher, 1867: 21" authorityName="Redtenbacher" authorityPageNumber="21" authorityYear="1867" box="[835,1298,1645,1666]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="113" phylum="Arthropoda" rank="species" species="nicobaricus">
<emphasis id="B90BEAB3A73FFFB53A40F9EABB354EF3" box="[835,1067,1645,1666]" italics="true" pageId="16" pageNumber="113">Hydroporus nicobaricus</emphasis>
<bibRefCitation id="EFEE4B50A73FFFB53D33F9EABA0C4EF3" author="Redtenbacher, L" box="[1072,1298,1645,1666]" pageId="16" pageNumber="113" refId="ref40351" refString="Redtenbacher, L., 1867. Coleopteren. In: Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Mullerdsdorf-Vobair. Akademie der Wissenschaften Wien, Karl Gerold's Sohn, iv + 249 + 5 pp." type="book" year="1867">Redtenbacher, 1867: 21</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0ADE10B0A73FFFB53C18F9EABAA14EF3" author="Sharp, D" box="[1307,1471,1645,1666]" page="802" pageId="16" pageNumber="113" year="1882">
<bibRefCitation id="EFEE4B50A73FFFB53C18F9EABAA14EF3" author="Sharp, D" box="[1307,1471,1645,1666]" pageId="16" pageNumber="113" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882: 802</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BC036A1A73FFFB53A40F90DBABE4EEF" blockId="16.[835,1475,1645,1892]" box="[835,1440,1673,1695]" pageId="16" pageNumber="113">
<treatmentCitationGroup id="AB6F118FA73FFFB53A40F90DBABE4EEF" box="[835,1440,1673,1695]" pageId="16" pageNumber="113">
<taxonomicName id="4C7F4D22A73FFFB53A40F90DBBB04EEF" authority="(Redtenbacher)" baseAuthorityName="Redtenbacher" box="[835,1198,1673,1695]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="113" phylum="Arthropoda" rank="species" species="nicobaricus">
<emphasis id="B90BEAB3A73FFFB53A40F90DBB0C4EEE" box="[835,1042,1674,1695]" italics="true" pageId="16" pageNumber="113">Bidessus nicobaricus</emphasis>
(Redtenbacher)
</taxonomicName>
-
<treatmentCitation id="0ADE10B0A73FFFB53DC0F90EBA824EEF" author="Regimbart, M" box="[1219,1436,1673,1694]" page="228" pageId="16" pageNumber="113" year="1899">
<bibRefCitation id="EFEE4B50A73FFFB53DC0F90EBA824EEF" author="Regimbart, M" box="[1219,1436,1673,1694]" pageId="16" pageNumber="113" pagination="186 - 367" refId="ref40540" refString="Regimbart, M., 1899. Revision des Dytiscidae de la region indosino-malaise. Annales de la Societe Entomologique de France, 68: 186 - 367." type="journal article" year="1899">Régimbart, 1899: 228</bibRefCitation>
</treatmentCitation>
.
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<paragraph id="8BC036A1A73FFFB53A40F921BC964EA6" blockId="16.[835,1475,1645,1892]" pageId="16" pageNumber="113">
<treatmentCitationGroup id="AB6F118FA73FFFB53A40F921BC964EA6" pageId="16" pageNumber="113">
<taxonomicName id="4C7F4D22A73FFFB53A40F921BBF54ECB" authority="(Redtenbacher)" baseAuthorityName="Redtenbacher" box="[835,1259,1701,1723]" class="Insecta" family="Dytiscidae" genus="Bidessus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="113" phylum="Arthropoda" rank="species" species="nicobaricus">
<emphasis id="B90BEAB3A73FFFB53A40F921BC894ECA" box="[835,919,1702,1723]" italics="true" pageId="16" pageNumber="113">Bidessus</emphasis>
(s.str.)
<emphasis id="B90BEAB3A73FFFB53ADCF921BB4F4ECA" box="[991,1105,1702,1723]" italics="true" pageId="16" pageNumber="113">nicobaricus</emphasis>
(Redtenbacher)
</taxonomicName>
-
<treatmentCitation id="0ADE10B0A73FFFB53DFEF922BC9A4EA6" author="Zimmermann, A" page="55" pageId="16" pageNumber="113" year="1920">
<bibRefCitation id="EFEE4B50A73FFFB53DFEF922BC9A4EA6" author="Zimmermann, A" pageId="16" pageNumber="113" pagination="1 - 326" refId="ref42405" refString="Zimmermann, A., 1920. Dytiscidae. In: Junk, W. &amp; S. Schenkling (eds.), Coleopterorum Catatalogus, Berlin, 4, pars 71, 1 - 326." type="journal article" year="1920">Zimmermann, 1920: 55</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036A1A73FFFB53A40F959BA8F4E82" blockId="16.[835,1475,1645,1892]" box="[835,1425,1758,1779]" pageId="16" pageNumber="113">
<treatmentCitationGroup id="AB6F118FA73FFFB53A40F959BA8F4E82" box="[835,1425,1758,1779]" pageId="16" pageNumber="113">
<taxonomicName id="4C7F4D22A73FFFB53A40F959BBA14E82" authority="(Redtenbacher)" baseAuthorityName="Redtenbacher" box="[835,1215,1758,1779]" class="Insecta" family="Dytiscidae" genus="Guignotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="113" phylum="Arthropoda" rank="species" species="nicobaricus">
<emphasis id="B90BEAB3A73FFFB53A40F959BB3C4E82" box="[835,1058,1758,1779]" italics="true" pageId="16" pageNumber="113">Guignotus nicobaricus</emphasis>
(Redtenbacher)
</taxonomicName>
-
<treatmentCitation id="0ADE10B0A73FFFB53DD7F959BA934E82" author="Vazirani, T" box="[1236,1421,1758,1779]" page="34" pageId="16" pageNumber="113" year="1977">
<bibRefCitation id="EFEE4B50A73FFFB53DD7F959BA934E82" author="Vazirani, T" box="[1236,1421,1758,1779]" pageId="16" pageNumber="113" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977: 34</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036A1A73FFFB53A40F97DBCCC4F5A" blockId="16.[835,1475,1645,1892]" pageId="16" pageNumber="113">
<treatmentCitationGroup id="AB6F118FA73FFFB53A40F97DBCCC4F5A" pageId="16" pageNumber="113">
<taxonomicName id="4C7F4D22A73FFFB53A40F97DBA534F7E" authority="(Redtenbacher)" baseAuthorityName="Redtenbacher" box="[835,1357,1786,1807]" class="Insecta" family="Dytiscidae" genus="Clypeodytes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="113" phylum="Arthropoda" rank="species" species="nicobaricus" subGenus="Lioclypeus">
<emphasis id="B90BEAB3A73FFFB53A40F97DBCA44F7E" box="[835,954,1786,1807]" italics="true" pageId="16" pageNumber="113">Clypeodytes</emphasis>
(
<emphasis id="B90BEAB3A73FFFB53ACBF97DBB2C4F7E" box="[968,1074,1786,1807]" italics="true" pageId="16" pageNumber="113">Lioclypeus</emphasis>
)
<emphasis id="B90BEAB3A73FFFB53D3CF97DBBAC4F7E" box="[1087,1202,1786,1807]" italics="true" pageId="16" pageNumber="113">nicobaricus</emphasis>
(Redtenbacher)
</taxonomicName>
-
<treatmentCitation id="0ADE10B0A73FFFB53C63F97DBCD04F5A" author="Wewalka, G" page="124" pageId="16" pageNumber="113" year="1982">
<bibRefCitation id="EFEE4B50A73FFFB53C63F97DBCD04F5A" author="Wewalka, G" pageId="16" pageNumber="113" pagination="115 - 125" refId="ref41663" refString="Wewalka, G., 1982. Results of the Austrian-Indian hydrobiological mission 1976, to the Andaman islands part IX, Dytiscidae (Coleoptera). Koleopterologische Rundschau, 56: 115 - 125." type="journal article" year="1982">Wewalka, 1982: 124</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BC036A1A73FFFB53A40F8B4BBC74F15" blockId="16.[835,1475,1645,1892]" pageId="16" pageNumber="113">
<treatmentCitationGroup id="AB6F118FA73FFFB53A40F8B4BBC74F15" pageId="16" pageNumber="113">
<taxonomicName id="4C7F4D22A73FFFB53A40F8B4BBB54F39" ID-CoL="3SYMC" authority="(Redtenbacher)" baseAuthorityName="Redtenbacher" baseAuthorityYear="1867" box="[835,1195,1843,1864]" class="Insecta" family="Dytiscidae" genus="Leiodytes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="113" phylum="Arthropoda" rank="species" species="nicobaricus">
<emphasis id="B90BEAB3A73FFFB53A40F8B4BB0D4F39" box="[835,1043,1843,1864]" italics="true" pageId="16" pageNumber="113">Leiodytes nicobaricus</emphasis>
(Redtenbacher)
</taxonomicName>
-
<treatmentCitation id="0ADE10B0A73FFFB53DBFF8B4BA764F39" author="Bistrom, O" box="[1212,1384,1843,1864]" page="27" pageId="16" pageNumber="113" year="1988">
<bibRefCitation id="EFEE4B50A73FFFB53DBFF8B4BA764F39" author="Bistrom, O" box="[1212,1384,1843,1864]" pageId="16" pageNumber="113" pagination="1 - 41" refId="ref35836" refString="Bistrom, O., 1988. Generic review of the Bidessini (Coleoptera, Dytiscidae). Acta zoologica Fennica, 184: 1 - 41." type="journal article" year="1988">Biström, 1988: 27</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73FFFB53C72F8B4BB124F15" author="Hebauer, F" page="337" pageId="16" pageNumber="113" year="1999">
<bibRefCitation id="EFEE4B50A73FFFB53C72F8B4BB124F15" author="Hebauer, F" pageId="16" pageNumber="113" pagination="333 - 348" refId="ref37686" refString="Hebauer, F., L. Hendrich &amp; M. Balke, 1999. A Contribution to the Knowledge of the Water Beetle Fauna of a Tropical Freshwater Lake: Tasek Cini, Pahang, West Malaysia (Coleoptera: Hydradephaga, Hydrophiloidea and Staphylinoidea). Raffles Bulletin of Zoology, 47: 333 - 348." type="journal article" year="1999">Hebauer et al., 1999: 337</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73FFFB53D14F8C8BBCA4F15" author="Nilsson, A" box="[1047,1236,1871,1892]" page="128" pageId="16" pageNumber="113" year="2001">
<bibRefCitation id="EFEE4B50A73FFFB53D14F8C8BBCA4F15" author="Nilsson, A" box="[1047,1236,1871,1892]" pageId="16" pageNumber="113" refId="ref39046" refString="Nilsson, A. N., 2001. Dytiscidae (Coleoptera). In: World Catalogue of Insects Volume 3. Apollo Books, Stenstrup, Denmark, 395 pp." type="book" year="2001">Nilsson, 2001: 128</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<emphasis id="B90BEAB3A73FFFB53A40F80FBB054FEC" bold="true" box="[835,1051,1928,1949]" italics="true" pageId="16" pageNumber="113">Material examined. </emphasis>
<materialsCitation id="3B173CFCA73FFFB53D21F80FBAAD4FED" box="[1058,1459,1927,1949]" location="Central" pageId="16" pageNumber="113" specimenCount="59">
<specimenCount id="9D79FD28A73FFFB53D21F80FBBB94FEC" box="[1058,1191,1928,1949]" count="59" pageId="16" pageNumber="113" type="generic">59 specimens</specimenCount>
<emphasis id="B90BEAB3A73FFFB53DAEF800BAAD4FED" bold="true" box="[1197,1459,1927,1948]" pageId="16" pageNumber="113">
-
<location id="8EA0607AA73FFFB53DBFF800BA124FED" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73FFFB53DBFF800BA124FED" box="[1212,1292,1927,1948]" name="Central" pageId="16" pageNumber="113">Central</location>
catchment area
</emphasis>
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<emphasis id="B90BEAB3A73FFFB53CB9F800BADC4FED" bold="true" box="[1466,1474,1927,1948]" pageId="16" pageNumber="113">-</emphasis>
<materialsCitation id="3B173CFCA73FFFB53A40F823BCDF4FA4" collectingDate="1992-01-16" collectionCode="ZRC" collectorName="C. M. Yang" location="Nee Soon Swamp Forest" pageId="16" pageNumber="113" specimenCount="10">
<specimenCount id="9D79FD28A73FFFB53A40F823BC9F4FC8" box="[835,897,1956,1977]" count="10" pageId="16" pageNumber="113" type="generic">10 ex.</specimenCount>
,
<location id="8EA0607AA73FFFB53A89F823BB634FC8" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73FFFB53A89F823BB634FC8" box="[906,1149,1956,1977]" name="Nee Soon Swamp Forest" pageId="16" pageNumber="113">Nee Soon Swamp Forest</location>
,
<date id="FFC11061A73FFFB53D85F823BBE34FC8" box="[1158,1277,1956,1977]" pageId="16" pageNumber="113" value="1992-01-16">
<collectingDate id="EF85E989A73FFFB53D85F823BBE34FC8" box="[1158,1277,1956,1977]" pageId="16" pageNumber="113" value="1992-01-16">16 Jan.1992</collectingDate>
</date>
, y 786A, coll.
<collectorName id="268A5377A73FFFB53C88F823BC664FA4" pageId="16" pageNumber="113">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73FFFB53A85F847BCA44FA4" box="[902,954,1984,2005]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="16" pageNumber="113">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73FFFB53ACFF847BCED4F83" collectingDate="1992-05-22" collectionCode="ZRC" collectorName="K. L. Yeo" location="Nee Soon Swamp Forest" pageId="16" pageNumber="113" specimenCount="1">
<specimenCount id="9D79FD28A73FFFB53ACFF847BCE04FA4" box="[972,1022,1984,2005]" count="1" pageId="16" pageNumber="113" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73FFFB53D0BF847BBE04FA4" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73FFFB53D0BF847BBE04FA4" box="[1032,1278,1984,2005]" name="Nee Soon Swamp Forest" pageId="16" pageNumber="113">Nee Soon Swamp Forest</location>
,
<date id="FFC11061A73FFFB53C0BF847BA934FA4" box="[1288,1421,1984,2005]" pageId="16" pageNumber="113" value="1992-05-22">
<collectingDate id="EF85E989A73FFFB53C0BF847BA934FA4" box="[1288,1421,1984,2005]" pageId="16" pageNumber="113" value="1992-05-22">22 May.1992</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73FFFB53A40F85ABCB64F83" box="[835,936,2013,2034]" pageId="16" pageNumber="113">K. L. Yeo</collectorName>
(
<collectionCode id="ED6EAE64A73FFFB53ABBF85ABCF24F83" box="[952,1004,2013,2034]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="16" pageNumber="113">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73FFFB53AFCF85ABB8A407F" collectingDate="1992-05-09" collectionCode="ZRC" collectorName="K. L. Yeo" location="Nee Soon Swamp Forest" pageId="16" pageNumber="113" specimenCount="1">
<specimenCount id="9D79FD28A73FFFB53AFCF85ABB2D4F83" box="[1023,1075,2013,2034]" count="1" pageId="16" pageNumber="113" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73FFFB53D3CF85ABA274F83" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73FFFB53D3CF85ABA274F83" box="[1087,1337,2013,2034]" name="Nee Soon Swamp Forest" pageId="16" pageNumber="113">Nee Soon Swamp Forest</location>
,
<date id="FFC11061A73FFFB53C47F85ABAA14F83" box="[1348,1471,2013,2034]" pageId="16" pageNumber="113" value="1992-05-09">
<collectingDate id="EF85E989A73FFFB53C47F85ABAA14F83" box="[1348,1471,2013,2034]" pageId="16" pageNumber="113" value="1992-05-09">9 May.1992</collectingDate>
</date>
, 795D, coll.
<collectorName id="268A5377A73FFFB53AB5F87EBB08407F" box="[950,1046,2041,2062]" pageId="16" pageNumber="113">K. L. Yeo</collectorName>
et al. (
<collectionCode id="ED6EAE64A73FFFB53D5AF87EBB93407F" box="[1113,1165,2041,2062]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="16" pageNumber="113">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73FFFB53D9DF87EBA16405B" collectingDate="1994-06-23" collectionCode="ZRC" collectorName="C. M. Yang" location="Chestnut Drive" municipality="Gangsa" pageId="16" pageNumber="113" specimenCount="1">
<specimenCount id="9D79FD28A73FFFB53D9DF87EBBD1407F" box="[1182,1231,2041,2062]" count="1" pageId="16" pageNumber="113" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73FFFB53DDAF87EBA6E407F" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73FFFB53DDAF87EBA6E407F" box="[1241,1392,2041,2062]" municipality="Gangsa" name="Chestnut Drive" pageId="16" pageNumber="113">Chestnut Drive</location>
,
<collectingMunicipality id="6BA4ACDBA73FFFB53C7AF87EBADC407F" box="[1401,1474,2041,2062]" pageId="16" pageNumber="113">Gangsa</collectingMunicipality>
stream,
<date id="FFC11061A73FFFB53A91F792BB12405B" box="[914,1036,2069,2090]" pageId="16" pageNumber="113" value="1994-06-23">
<collectingDate id="EF85E989A73FFFB53A91F792BB12405B" box="[914,1036,2069,2090]" pageId="16" pageNumber="113" value="1994-06-23">23 Jun.1994</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73FFFB53D4AF792BBA0405B" box="[1097,1214,2069,2090]" pageId="16" pageNumber="113">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73FFFB53DCFF792BA1E405B" box="[1228,1280,2069,2090]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="16" pageNumber="113">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73FFFB53C10F792BA9B4036" collectingDate="1994-04-21" collectionCode="ZRC" collectorName="K. L. Yeo" location="MacRitchie Reservoir" pageId="16" pageNumber="113" specimenCount="2">
<specimenCount id="9D79FD28A73FFFB53C10F792BA5B405B" box="[1299,1349,2069,2090]" count="2" pageId="16" pageNumber="113" type="generic">2 ex.</specimenCount>
,
<location id="8EA0607AA73FFFB53C53F792BCBA4036" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73FFFB53C53F792BCBA4036" name="MacRitchie Reservoir" pageId="16" pageNumber="113">MacRitchie Reservoir</location>
,
<date id="FFC11061A73FFFB53AAEF7B5BB374036" box="[941,1065,2098,2119]" pageId="16" pageNumber="113" value="1994-04-21">
<collectingDate id="EF85E989A73FFFB53AAEF7B5BB374036" box="[941,1065,2098,2119]" pageId="16" pageNumber="113" value="1994-04-21">21 Apr.1994</collectingDate>
</date>
, NS 124 F/H, coll.
<collectorName id="268A5377A73FFFB53DDCF7B5BA234036" box="[1247,1341,2098,2119]" pageId="16" pageNumber="113">K. L. Yeo</collectorName>
(
<collectionCode id="ED6EAE64A73FFFB53C49F7B5BA604036" box="[1354,1406,2098,2119]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="16" pageNumber="113">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73FFFB43C8DF7B5BE5E4887" collectingDate="1997-04-21" collectionCode="CBH" collectorName="M. Balke &amp; L. Hendrich &amp; C. M. Yang" lastPageId="17" lastPageNumber="114" location="MacRitchie Reservoir" pageId="16" pageNumber="113" specimenCount="4">
<specimenCount id="9D79FD28A73FFFB53C8DF7B5BAA04036" box="[1422,1470,2098,2119]" count="4" pageId="16" pageNumber="113" type="generic">4 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB43972FF45BE5748A6" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB43972FF45BE5748A6" box="[113,329,194,215]" name="MacRitchie Reservoir" pageId="17" pageNumber="114">MacRitchie Reservoir</location>
,
<date id="FFC11061A73EFFB43852FF45BED348A6" box="[337,461,194,215]" pageId="17" pageNumber="114" value="1997-04-21">
<collectingDate id="EF85E989A73EFFB43852FF45BED348A6" box="[337,461,194,215]" pageId="17" pageNumber="114" value="1997-04-21">21 Apr.1997</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73EFFB43B05FF45BD4148A6" box="[518,607,194,215]" pageId="17" pageNumber="114">M. Balke</collectorName>
&amp;
<collectorName id="268A5377A73EFFB43B7FFF45BDEE48A6" box="[636,752,194,215]" pageId="17" pageNumber="114">L. Hendrich</collectorName>
&amp;
<collectorName id="268A5377A73EFFB4398AFF66BFE94887" box="[137,247,225,246]" pageId="17" pageNumber="114">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB43800FF66BE274887" box="[259,313,225,246]" pageId="17" pageNumber="114">CBH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73EFFB4384BFF66BD764964" collectingDate="1997-04-22" collectionCode="ZRC" collectorName="M. Balke &amp; L. Hendrich" location="Nee Soon Swamp Forest" pageId="17" pageNumber="114" specimenCount="5">
<specimenCount id="9D79FD28A73EFFB4384BFF66BE664887" box="[328,376,225,246]" count="5" pageId="17" pageNumber="114" type="generic">5 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB43883FF66BD724887" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB43883FF66BD724887" box="[384,620,225,246]" name="Nee Soon Swamp Forest" pageId="17" pageNumber="114">Nee Soon Swamp Forest</location>
, at blacklight,
<date id="FFC11061A73EFFB43972FE87BFEE4964" box="[113,240,256,277]" pageId="17" pageNumber="114" value="1997-04-22">
<collectingDate id="EF85E989A73EFFB43972FE87BFEE4964" box="[113,240,256,277]" pageId="17" pageNumber="114" value="1997-04-22">22 Apr.1997</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73EFFB4382FFE87BE964964" box="[300,392,256,277]" pageId="17" pageNumber="114">M. Balke</collectorName>
&amp;
<collectorName id="268A5377A73EFFB438ABFE87BD014964" box="[424,543,256,277]" pageId="17" pageNumber="114">L. Hendrich</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB43B2EFE87BD7F4964" box="[557,609,256,277]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="17" pageNumber="114">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73EFFB43B70FE87BE4F4922" collectingDate="1997-04-22" collectionCode="ZRC" collectorName="H. K. Lua" location="Lower Peirce Reservoir" pageId="17" pageNumber="114" specimenCount="5">
<specimenCount id="9D79FD28A73EFFB43B70FE87BDB84964" box="[627,678,256,277]" count="5" pageId="17" pageNumber="114" type="generic">5 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB43BB2FE87BE114945" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB43BB2FE87BE114945" name="Lower Peirce Reservoir" pageId="17" pageNumber="114">Lower Peirce Reservoir</location>
, after the concrete water runoff,
<date id="FFC11061A73EFFB43B38FE98BDAA4945" box="[571,692,287,308]" pageId="17" pageNumber="114" value="1997-04-22">
<collectingDate id="EF85E989A73EFFB43B38FE98BDAA4945" box="[571,692,287,308]" pageId="17" pageNumber="114" value="1997-04-22">22 Apr.1997</collectingDate>
</date>
, 230a, coll.
<collectorName id="268A5377A73EFFB439A0FEB9BE164922" box="[163,264,318,339]" pageId="17" pageNumber="114">H. K. Lua</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB43815FEB9BE554922" box="[278,331,318,339]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="17" pageNumber="114">ZRC</collectionCode>
)
</materialsCitation>
.
<emphasis id="B90BEAB3A73EFFB4385FFEBABEEC4923" bold="true" box="[348,498,317,338]" pageId="17" pageNumber="114">Bukit Timah -</emphasis>
<materialsCitation id="3B173CFCA73EFFB438FAFEB9BEBF49E0" collectingDate="1997-04-09" collectionCode="CBH" collectorName="M. Balke &amp; L. Hendrich &amp; H. K. Lua" location="Bukit Timah" pageId="17" pageNumber="114" specimenCount="1">
<specimenCount id="9D79FD28A73EFFB438FAFEB9BD324922" box="[505,556,318,339]" count="1" pageId="17" pageNumber="114" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB43B34FEB9BDAA4922" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB43B34FEB9BDAA4922" box="[567,692,318,339]" name="Bukit Timah" pageId="17" pageNumber="114">Bukit Timah</location>
<taxonomicNameLabel id="A23857C8A73EFFB43BBFFEB9BDF24922" box="[700,748,318,339]" pageId="17" pageNumber="114" rank="subSpecies">N.R.</taxonomicNameLabel>
, swampy meadow near fishpond,
<date id="FFC11061A73EFFB438BFFEDABD2E4903" box="[444,560,349,370]" pageId="17" pageNumber="114" value="1997-04-09">
<collectingDate id="EF85E989A73EFFB438BFFEDABD2E4903" box="[444,560,349,370]" pageId="17" pageNumber="114" value="1997-04-09">9 Apr.1997</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73EFFB43B73FEDABDD14903" box="[624,719,349,370]" pageId="17" pageNumber="114">M. Balke</collectorName>
,
<collectorName id="268A5377A73EFFB43BD8FEDABFD249E0" pageId="17" pageNumber="114">L. Hendrich</collectorName>
&amp;
<collectorName id="268A5377A73EFFB439ECFEFBBE4A49E0" box="[239,340,380,401]" pageId="17" pageNumber="114">H. K. Lua</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB43860FEFBBE8449E0" box="[355,410,380,401]" pageId="17" pageNumber="114">CBH</collectionCode>
)
</materialsCitation>
.
<emphasis id="B90BEAB3A73EFFB438AEFEFCBD2149E1" bold="true" box="[429,575,379,400]" pageId="17" pageNumber="114">
<materialsCitation id="3B173CFCA73EFFB438AEFEFCBD3449E1" box="[429,554,379,400]" location="Other" pageId="17" pageNumber="114" specimenCount="1">
<location id="8EA0607AA73EFFB438AEFEFCBEF549E1" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB438AEFEFCBEF549E1" box="[429,491,379,400]" name="Other" pageId="17" pageNumber="114">Other</location>
areas
</materialsCitation>
</emphasis>
<materialsCitation id="3B173CFCA73EFFB43B44FEFBBEE349BF" collectingDate="1997-04-21" collectionCode="CBH" collectorName="M. Balke &amp; L. Hendrich &amp; H. K. Lua &amp; C. M. Yang" location="Botanical Garden" pageId="17" pageNumber="114" specimenCount="23">
<specimenCount id="9D79FD28A73EFFB43B44FEFBBD9949E0" box="[583,647,380,401]" count="23" pageId="17" pageNumber="114" type="generic">23 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB43B91FEFBBFA249C1" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB43B91FEFBBFA249C1" name="Botanical Garden" pageId="17" pageNumber="114">Botanical Garden</location>
, small artificial pond,
<date id="FFC11061A73EFFB438ABFE1CBD3449C1" box="[424,554,411,432]" pageId="17" pageNumber="114" value="1997-04-21">
<collectingDate id="EF85E989A73EFFB438ABFE1CBD3449C1" box="[424,554,411,432]" pageId="17" pageNumber="114" value="1997-04-21">21 Apr.1997</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73EFFB43B6FFE1CBDD049C1" box="[620,718,411,432]" pageId="17" pageNumber="114">M. Balke</collectorName>
,
<collectorName id="268A5377A73EFFB43BD8FE1CBFD549BF" pageId="17" pageNumber="114">L. Hendrich</collectorName>
,
<collectorName id="268A5377A73EFFB439D0FE3EBE3049BF" box="[211,302,441,462]" pageId="17" pageNumber="114">H. K. Lua</collectorName>
&amp;
<collectorName id="268A5377A73EFFB4384AFE3EBEAB49BF" box="[329,437,441,462]" pageId="17" pageNumber="114">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB438C2FE3EBEE849BF" box="[449,502,441,462]" pageId="17" pageNumber="114">CBH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73EFFB43B06FE3EBDC4499C" collectingDate="1997-04-23" collectionCode="CBH" collectorName="M. Balke &amp; L. Hendrich" location="Bukit Batok Nature Park" pageId="17" pageNumber="114" specimenCount="1">
<specimenCount id="9D79FD28A73EFFB43B06FE3EBD2D49BF" box="[517,563,441,462]" count="1" pageId="17" pageNumber="114" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB43B38FE3EBFBE499C" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB43B38FE3EBFBE499C" name="Bukit Batok Nature Park" pageId="17" pageNumber="114">Bukit Batok Nature Park</location>
, pond,
<date id="FFC11061A73EFFB439E4FE5FBE7B499C" box="[231,357,472,493]" pageId="17" pageNumber="114" value="1997-04-23">
<collectingDate id="EF85E989A73EFFB439E4FE5FBE7B499C" box="[231,357,472,493]" pageId="17" pageNumber="114" value="1997-04-23">23 Apr.1997</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73EFFB438A3FE5FBEE4499C" box="[416,506,472,493]" pageId="17" pageNumber="114">M. Balke</collectorName>
&amp;
<collectorName id="268A5377A73EFFB43B1AFE5FBD91499C" box="[537,655,472,493]" pageId="17" pageNumber="114">L. Hendrich</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB43B9FFE5FBDCD499C" box="[668,723,472,493]" pageId="17" pageNumber="114">CBH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73EFFB43BE7FE5FBE564A5A" collectingDate="1995-06-29" collectionCode="ZRC" collectorName="H. K. Lua &amp; C. M. Yang" location="Lorong Banir" pageId="17" pageNumber="114" specimenCount="2">
<specimenCount id="9D79FD28A73EFFB43BE7FE5FBF8F4A7D" count="2" pageId="17" pageNumber="114" type="generic">2 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB4399FFE70BE3D4A7D" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB4399FFE70BE3D4A7D" box="[156,291,503,524]" name="Lorong Banir" pageId="17" pageNumber="114">Lorong Banir</location>
, pond 1,
<date id="FFC11061A73EFFB4387CFE70BEE74A7D" box="[383,505,503,524]" pageId="17" pageNumber="114" value="1995-06-29">
<collectingDate id="EF85E989A73EFFB4387CFE70BEE74A7D" box="[383,505,503,524]" pageId="17" pageNumber="114" value="1995-06-29">29 Jun.1995</collectingDate>
</date>
, NS 193, coll.
<collectorName id="268A5377A73EFFB43B8EFE70BDEE4A7D" box="[653,752,503,524]" pageId="17" pageNumber="114">H. K. Lua</collectorName>
&amp;
<collectorName id="268A5377A73EFFB43988FD91BFE14A5A" box="[139,255,534,555]" pageId="17" pageNumber="114">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB4380EFD91BE5F4A5A" box="[269,321,534,555]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="17" pageNumber="114">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73EFFB43850FD91BD214A3B" collectingDate="1995-06-16" collectionCode="ZRC" collectorName="H. K. Lua &amp; C. M. Yang" location="Lorong Banir" pageId="17" pageNumber="114" specimenCount="1">
<specimenCount id="9D79FD28A73EFFB43850FD91BE9B4A5A" box="[339,389,534,555]" count="1" pageId="17" pageNumber="114" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73EFFB43893FD91BD094A5A" LSID="urn:lsid:plazi:treatment:03D687B7A73FFFB43A88F996BE1C4BCF:8EA0607AA73EFFB43893FD91BD094A5A" box="[400,535,534,555]" name="Lorong Banir" pageId="17" pageNumber="114">Lorong Banir</location>
, pond 2,
<date id="FFC11061A73EFFB43B71FD91BDF34A5A" box="[626,749,534,555]" pageId="17" pageNumber="114" value="1995-06-16">
<collectingDate id="EF85E989A73EFFB43B71FD91BDF34A5A" box="[626,749,534,555]" pageId="17" pageNumber="114" value="1995-06-16">16 Jun.1995</collectingDate>
</date>
, NS 192, coll.
<collectorName id="268A5377A73EFFB439FFFDB2BE414A3B" box="[252,351,565,586]" pageId="17" pageNumber="114">H. K. Lua</collectorName>
&amp;
<collectorName id="268A5377A73EFFB43882FDB2BEE84A3B" box="[385,502,565,586]" pageId="17" pageNumber="114">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73EFFB43B07FDB2BD274A3B" box="[516,569,565,586]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="17" pageNumber="114">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73EFFB43972FDF6BE934AB6" pageId="17" pageNumber="114" type="distribution">
<paragraph id="8BC036A1A73EFFB43972FDF6BE934AB6" blockId="17.[113,752,625,711]" pageId="17" pageNumber="114">
<emphasis id="B90BEAB3A73EFFB43972FDF6BE054AF8" bold="true" box="[113,283,625,649]" italics="true" pageId="17" pageNumber="114">Distribution. </emphasis>
India: Nicobar Islands (
<bibRefCitation id="EFEE4B50A73EFFB43B35FDF6BDFD4AF9" author="Vazirani, T" box="[566,739,625,648]" pageId="17" pageNumber="114" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977</bibRefCitation>
); Thailand (Wang, in litt.). First record for the Malaysian Peninsula and
<emphasis id="B90BEAB3A73EFFB43813FD28BE934AB6" bold="true" box="[272,397,687,711]" pageId="17" pageNumber="114">
<collectingCountry id="F3687631A73EFFB43813FD28BE974AB6" box="[272,393,687,711]" name="Singapore" pageId="17" pageNumber="114">Singapore</collectingCountry>
!
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73EFFB43972FD6ABE1C4BCF" pageId="17" pageNumber="114" type="biology_ecology">
<paragraph id="8BC036A1A73EFFB43972FD6ABE1C4BCF" blockId="17.[113,753,749,958]" pageId="17" pageNumber="114">
<emphasis id="B90BEAB3A73EFFB43972FD6ABEFB4B75" box="[113,485,749,773]" italics="true" pageId="17" pageNumber="114">
<emphasis id="B90BEAB3A73EFFB43972FD6ABFF74B74" bold="true" box="[113,233,749,773]" italics="true" pageId="17" pageNumber="114">Ecology. </emphasis>
<taxonomicName id="4C7F4D22A73EFFB439F7FD6ABEFB4B75" baseAuthorityName="Redtenbacher" baseAuthorityYear="1867" box="[244,485,749,772]" class="Insecta" family="Dytiscidae" genus="Leiodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="nicobaricus">Leiodytes nicobaricus</taxonomicName>
</emphasis>
is the most widespread species of
<taxonomicName id="4C7F4D22A73EFFB439ECFC8BBE494B52" authorityName="Sharp" authorityYear="1880" box="[239,343,780,803]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Bidessini">Bidessini</taxonomicName>
in
<collectingCountry id="F3687631A73EFFB43880FC8BBEE64B52" box="[387,504,780,803]" name="Singapore" pageId="17" pageNumber="114">Singapore</collectingCountry>
. The species inhabits exposed or semi-shaded, temporary and permanent water bodies rich in vegetation and plant debris, such as ponds, pools and drainage ditches in primary and cultivated areas. The beetle occupies the very shallow zone just at the edge of the water bodies.
</paragraph>
</subSubSection>
</treatment>
</document>