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<document id="C738DEC18ED1FAEA3E4D2D16948394FB" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFC1FFD13C2C6307FB5875CC" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Epaphiotrechus fortipes 1999" docType="treatment" docVersion="2" lastPageNumber="429" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="425" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8F788DF581095E07F295434E8B9B889C">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="95F20964DADA5C114CFE48F455C9843A">Deuve, Thierry</mods:namePart>
<mods:affiliation id="E7474E1C02845253D0E35E45E2021930">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="76CF38C51B0AB74965EA030749431FD5">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5F1747966B626DC3402FD2A40C6EBE1E">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="144A064CEA2A15786831F509796281DF">Liang, Hongbin</mods:namePart>
<mods:affiliation id="1923587C18EBD27FDF68BE3F3D8AC148">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:date id="F8F42EB5ABDCF04D5138433551660C3B">2016</mods:date>
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<paragraph id="8BC0368BFFC1FFDD3C2C6307FDD574DE" blockId="84.[123,539,885,940]" box="[123,539,885,910]" pageId="84" pageNumber="425">
<heading id="D08881E7FFC1FFDD3C2C6307FDD574DE" box="[123,539,885,910]" fontSize="10" level="2" pageId="84" pageNumber="425" reason="4">
<taxonomicName id="4C7F4D08FFC1FFDD3C2C6307FDD574DE" ID-CoL="8MMR7" authority="(Ueno), 1999" authorityName="" authorityYear="1999" baseAuthorityName="Ueno" baseAuthorityYear="1999" box="[123,539,885,910]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFC1FFDD3C2C6307FDD574DE" bold="true" box="[123,539,885,910]" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3C2C6307FE4A74DE" bold="true" box="[123,388,886,910]" italics="true" pageId="84" pageNumber="425">Epaphiotrechus fortipes</emphasis>
(Uéno), 1999
</emphasis>
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<subSubSection id="C3656500FFC1FFDD3C2C63E7FEDA74FC" box="[123,276,918,940]" pageId="84" pageNumber="425" type="description">
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(
<figureCitation id="13442A0EFFC1FFDD3CD563E7FF0774FB" box="[130,201,918,940]" captionStart="FIGURE" captionStartId="85.[699,711,1580,1599]" captionTargetBox="[123,1176,216,1723]" captionTargetId="figure-308@85.[648,1176,216,1567]" captionTargetPageId="85" captionText="FIGURE 32. Epaphiotrechus fortipes (Uéno); Map of locality record (red circle) in the Gaoligong Shan region. Scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155344" httpUri="https://zenodo.org/record/13155344/files/figure.png" pageId="84" pageNumber="425">Fig. 32</figureCitation>
,
<figureCitation id="13442A0EFFC1FFDD3C8363E7FEC374FC" box="[212,269,918,940]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="84" pageNumber="425">46-48</figureCitation>
)
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<paragraph id="8BC0368BFFC1FFDD3C2C63CFFDD77484" blockId="84.[123,633,958,1012]" box="[123,537,958,980]" pageId="84" pageNumber="425">
<treatmentCitationGroup id="AB6F11A5FFC1FFDD3C2C63CFFDD77484" box="[123,537,958,980]" pageId="84" pageNumber="425">
<treatmentCitation id="0ADE109AFFC1FFDD3C2C63CFFDDB7484" author="UENO, S. I." box="[123,533,958,980]" page="219" pageId="84" pageNumber="425" year="1999">
<taxonomicName id="4C7F4D08FFC1FFDD3C2C63CFFDDB7484" ID-CoL="8LP5V" authority="Ueno, 1999 a: 219" authorityName="Ueno" authorityPageNumber="219" authorityYear="1999" box="[123,533,958,980]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFC1FFDD3C2C63CFFF087483" box="[123,198,958,979]" italics="true" pageId="84" pageNumber="425">Trechus</emphasis>
(
<emphasis id="B90BEA99FFC1FFDD3C8363CEFEC67484" box="[212,264,959,980]" italics="true" pageId="84" pageNumber="425">s. str.</emphasis>
)
<emphasis id="B90BEA99FFC1FFDD3D4163CFFE917483" box="[278,351,958,979]" italics="true" pageId="84" pageNumber="425">fortipes</emphasis>
<bibRefCitation id="EFEE4B7AFFC1FFDD3D3163CFFDDB7484" author="UENO, S. I." box="[358,533,958,980]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno, 1999a: 219</bibRefCitation>
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.
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<taxonomicName id="4C7F4D08FFC1FFDD3C2C63AFFE6174A4" ID-CoL="8MMR7" authority="(Ueno)" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2016" box="[123,431,990,1012]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes" status="comb. nov.">
<emphasis id="B90BEA99FFC1FFDD3C2C63AFFEAF74A3" box="[123,353,990,1011]" italics="true" pageId="84" pageNumber="425">Epaphiotrechus fortipes</emphasis>
(Uéno)
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,
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<emphasis id="B90BEA99FFC1FFDD3DEE63AEFE2674A2" bold="true" box="[441,488,991,1012]" pageId="84" pageNumber="425">NEW</emphasis>
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<emphasis id="B90BEA99FFC1FFDD3DB963AFFDB774A3" bold="true" box="[494,633,990,1012]" pageId="84" pageNumber="425">COMBINATION</emphasis>
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.—
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<typeStatus id="54C48829FFC1FFDD3DD06474FE23734D" box="[391,493,1029,1053]" pageId="84" pageNumber="425" type="holotype">Holotype</typeStatus>
, a male, in NSMT. Type locality:
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,
<collectingRegion id="49BBF869FFC1FFDD3F956474FBD8734C" box="[962,1046,1029,1052]" country="China" name="Yunnan" pageId="84" pageNumber="425">Yunnan</collectingRegion>
,
<location id="8EA06050FFC1FFDD38726474FF7D736D" LSID="urn:lsid:plazi:treatment:03D6879DFFC1FFD13C2C6307FB5875CC:8EA06050FFC1FFDD38726474FF7D736D" country="China" county="Baoshan County" latitude="24.95" longLatPrecision="1249" longitude="98.75" name="Gaoligong Shan" pageId="84" pageNumber="425" stateProvince="Yunnan">Gaoligong Shan</location>
,
<collectingCounty id="62A14E07FFC1FFDD3CE96454FEBF736D" box="[190,369,1061,1085]" pageId="84" pageNumber="425">Baoshan County</collectingCounty>
,
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/
<geoCoordinate id="EE4B504CFFC1FFDD3DB36452FDF2736C" box="[484,572,1059,1084]" degrees="98" direction="east" minutes="45" orientation="longitude" pageId="84" pageNumber="425" precision="925" value="98.75">E98°45</geoCoordinate>
,
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</paragraph>
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</emphasis>
.— We have not had an opportunity to study the
<typeStatus id="54C48829FFC1FFDD3FBD6434FB89730D" box="[1002,1095,1093,1117]" pageId="84" pageNumber="425" type="holotype">holotype</typeStatus>
or any
<typeStatus id="54C48829FFC1FFDD3C2C6414FEE3732D" box="[123,301,1125,1149]" isOtherTypeStatus="true" pageId="84" pageNumber="425">other specimens</typeStatus>
of this species. Features noted below are based on
<bibRefCitation id="EFEE4B7AFFC1FFDD3FDC6414FBF8732D" author="UENO, S. I." box="[907,1078,1125,1149]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uénos (1999a)</bibRefCitation>
original description and illustrations.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC1FFDF3CFC64D4FD957565" lastPageId="86" lastPageNumber="427" pageId="84" pageNumber="425" type="diagnosis">
<paragraph id="8BC0368BFFC1FFDD3CFC64D4FC22738C" blockId="84.[123,1174,1029,1726]" pageId="84" pageNumber="425">
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<emphasis id="B90BEA99FFC1FFDD3CFC64D4FEEE73EB" bold="true" box="[171,288,1189,1213]" pageId="84" pageNumber="425">DIAGNOSIS</emphasis>
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.— Adults of this species (see Uéno 199a,
<figureCitation id="13442A0EFFC1FFDD3EB964D4FCE073ED" box="[750,814,1189,1213]" captionStart="FIGURES" captionStartId="4.[171,183,1157,1176]" captionTargetBox="[120,1176,216,1144]" captionTargetId="figure-163@4.[120,1176,216,1144]" captionTargetPageId="4" captionText="FIGURES 24. Fig. 2. Map showing natural extent of study area, colored in green (however, sampling was permitted only in those portions in Yunnan Province. Fig. 3. Map showing location of core sampling areas. Fig. 4. Map showing locations of all entomological sampling sites." figureDoi="http://doi.org/10.5281/zenodo.13155287" httpUri="https://zenodo.org/record/13155287/files/figure.png" pageId="84" pageNumber="425">Fig. 4</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states:
</paragraph>
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Size large (BL =
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), apterous; female
<typeStatus id="54C48829FFC1FFDD3ECD6494FD3973AD" box="[666,759,1253,1277]" pageId="84" pageNumber="425" type="holotype">holotype</typeStatus>
with unusually broad protarsomeres; body color dark piceous, slightly paler reddish on elytral interval 1; right mandible tridentate; mentum with median tooth short and simple; pronotum slightly narrowed (ratio PW/PL = 1.26), with basal angles small and rectangular; elytra elongate, slightly flattened medially, striae finely impressed, medial three or four striae distinctly crenulate (due to strial punctures), more lateral striae slightly effaced, but striae 6 and 7 evident, stria 5 deepened at its base; parascutellar striole present, slightly elongate; recurrent stria short, terminated near apex of stria 5; two discal setiferous pores present, inserted at anterior one-sixth and slightly posterior to middle, respectively, on interval 3 near stria 3; preapical seta present, inserted on interval 3 next to stria 2 and about equidistant from apical and sutural elytral margins; median lobe of male aedeagus (see
<bibRefCitation id="EFEE4B7AFFC1FFDD3FEF6674FBF0714D" author="UENO, S. I." box="[952,1086,1541,1565]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno 1999a</bibRefCitation>
,
<figureCitation id="13442A0EFFC1FFDD381C6674FB5B714D" box="[1099,1173,1541,1565]" captionStart="FIGURE" captionStartId="8.[171,183,1613,1632]" captionTargetBox="[120,1176,224,1597]" captionTargetId="figure-11@8.[120,1176,224,1597]" captionTargetPageId="8" captionText="FIGURE 5. Perileptus imaicus Jeannel; a. Dorsal habitus (CASENT1036287). b. Median lobe of aedeagus of male (CASENT10363020), left lateral aspect. c. Map of locality records (red circles) for P. imaicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155291" httpUri="https://zenodo.org/record/13155291/files/figure.png" pageId="84" pageNumber="425">Figs. 5</figureCitation>
and
<figureCitation id="13442A0EFFC1FFDD3CFE6654FF79716D" box="[169,183,1573,1597]" captionStart="FIGURE" captionStartId="10.[171,183,1609,1628]" captionTargetBox="[144,1176,216,1593]" captionTargetId="figure-11@10.[120,1176,216,1593]" captionTargetPageId="10" captionText="FIGURE 6. Perileptus pusilloides sp. nov.; a. Dorsal habitus (CASENT1039260). b. Median lobe of aedeagus of male (CASENT1039259), left lateral aspect. c. Map of locality records (red circles) for P. pusilloides in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155293" httpUri="https://zenodo.org/record/13155293/files/figure.png" pageId="84" pageNumber="425">6</figureCitation>
) with apex, expanded, securiform in lateral view; endophallus with distinct, elongate-triangular sclerite.
</paragraph>
<paragraph id="8BC0368BFFC1FFDD3CFC6614FB5871ED" blockId="84.[123,1174,1029,1726]" pageId="84" pageNumber="425">
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</emphasis>
.— No members of this species were found during field collecting for this project, and no habitat information accompanied the
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material.
<bibRefCitation id="EFEE4B7AFFC1FFDD3FDA66F4FBEA71CD" author="UENO, S. I." box="[909,1060,1669,1693]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno (1999a)</bibRefCitation>
suggested that the
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locality was located “near the lower edge of the
<taxonomicName id="4C7F4D08FFC1FFDD3F4F66D4FC7871ED" box="[792,950,1701,1725]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="84" pageNumber="425" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFC1FFDD3F4F66D4FC7871ED" box="[792,950,1701,1725]" italics="true" pageId="84" pageNumber="425">Rhododendron</emphasis>
</taxonomicName>
zone. He visited the
</paragraph>
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32.
<taxonomicName id="4C7F4D08FFC0FFDC3F7F665DFB8D716F" authority="(Ueno)" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2016" box="[808,1091,1580,1599]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="426" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFC0FFDC3F7F665DFC34716F" box="[808,1018,1580,1599]" italics="true" pageId="85" pageNumber="426">Epaphiotrechus fortipes</emphasis>
(Uéno)
</taxonomicName>
; Map of locality record (red circle) in the Gaoligong Shan region.
</paragraph>
<paragraph id="8BC0368BFFC0FFDC3EDC662EFCF37122" blockId="85.[651,1175,1580,1650]" box="[651,829,1631,1650]" pageId="85" pageNumber="426">Scale line = 100 km.</paragraph>
</caption>
<paragraph id="8BC0368BFFC3FFDF3CFC60AAFD957565" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">
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<emphasis id="B90BEA99FFC3FFDF3CFC60AAFEEE77A1" bold="true" box="[171,288,219,243]" pageId="86" pageNumber="427">DIAGNOSIS</emphasis>
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.— Adults of this species (
<figureCitation id="13442A0EFFC3FFDF3E1060AAFD6D77A3" box="[583,675,219,243]" captionStart="FIGURE" captionStartId="87.[171,183,1591,1610]" captionTargetBox="[120,1176,216,1574]" captionTargetId="figure-15@87.[120,1176,216,1575]" captionTargetPageId="87" captionText="FIGURE 33. Epaphiotrechus fortipesoides sp. nov.; a. Dorsal habitus (CASENT1039089). b. Map of locality record (red circle) for E. fortipesoides in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155346" httpUri="https://zenodo.org/record/13155346/files/figure.png" pageId="86" pageNumber="427">Fig. 33a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL =
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), apterous; body color piceous, sutural and lateral elytral margins slightly paler, reddish; tempora glabrous; median tooth of mentum apically truncate, almost bifurcate; antennae long; pronotum slightly narrowed (ratio PW/PL = 1.28), basal angles small, sharp, rectangular; elytra oblong, slightly flattened along median suture, striae finely impressed, finely punctate, striae 1 to 2 or 3 more deeply impressed, more lateral striae more or less effaced but still evident, stria 5 not more deeply impressed at its base; recurrent stria long, deeply impressed, abruptly terminated on interval 7; two discal setiferous pores present, inserted at anterior one-sixth and slightly posterior to middle, respectively, on interval 3 near stria 3; preapical seta present, inserted 1.5 times as far from apical elytral margin as from sutural margin.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC3FFDF3CFC624CFC8C71CC" pageId="86" pageNumber="427" type="description">
<paragraph id="8BC0368BFFC3FFDF3CFC624CFBB475E6" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">
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<emphasis id="B90BEA99FFC3FFDF3CFC624CFEF47504" bold="true" box="[171,314,573,597]" pageId="86" pageNumber="427">DESCRIPTION</emphasis>
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.— Size large, BL =
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. Body color dark piceous, head, basal and lateral pronotal margins, and lateral and sutural margins of elytra more or less reddish, appendages paler reddish brown, palpi yellowish brown, dorsum smooth, shiny with pronotum and elytra moderately iridescent, head faintly alutaceous due to moderately impressed isodiametric microsculpture.
</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC62CFFEEC7348" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">
Head. Broad, slightly elongate. Eyes slightly convex, their diameter slightly less than twice length of tempora, the latter short but distinctly convex. Frons convex, with frontal furrows sharply impressed, not attenuated posterior to eyes. Two pairs of supraorbital setae present, the anterior pair inserted in slight foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Right mandible (
<figureCitation id="13442A0EFFC3FFDF3DE6634EFDC37407" box="[433,525,831,855]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="86" pageNumber="427">
Fig.
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) with premolar not fused with retinaculum but closely associated with the latter, anterior tip of retinaculum enlarged as a distinct tooth and displaced anteriorly.
<collectingRegion id="49BBF869FFC3FFDF3CE563F1FEC274C7" box="[178,268,896,919]" country="Myanmar" name="Chin" pageId="86" pageNumber="427">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFC3FFDF3E6D63F1FD5A74C7" box="[570,660,896,919]" country="Myanmar" name="Chin" pageId="86" pageNumber="427">Mentum</collectingRegion>
with medial tooth apically truncate, about one-half the length of the lateral lobes. Submentum with six setae anteriorly. Genae with a single ventral seta one each side. Antennae moderate in length, with about 2.5 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 3 and 4 about equal in length, antennomere 2 slightly shorter.
</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6450FE2F726A" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">Pronotum. Slightly narrowed (ratio PW/PL = 1.28), with greatest width anterior to middle, only slightly narrowed posteriorly; lateral margins rounded, straightened for a short distance posteriorly, then slightly sinuate just anterior to basal angles, the latter small, sharp and rectangular; basal margin with median region slightly and convexly projected posteriorly. Disc convex, with median longitudinal impression distinct, markedly deepened in median basal area, not extended anteriorly to apical margin; basal foveae moderate in size and depth; median basal area delimited laterally by a pair of short, oblique furrows, smooth except for a pair of paramedial longitudinal foveae. Single midlateral setae on each side inserted anterior to middle; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6532FB8D716C" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">Elytra. Elongate ovoid, slightly narrowed anteriorly, humeri rounded and only slightly evident. Disc moderately convex, slightly flattened in sutural area; striae finely impressed, finely punctate, striae 1 to 2 or 3 more deeply impressed, more lateral striae more or less effaced but still evident, stria 5 not more deeply impressed at its base. Parascutellar striole present, deeply impressed and long. Recurrent stria long, deeply impressed, abruptly terminated on interval 7 slightly anterior to the insertion point of preapical seta. Two discal setiferous pores present, inserted at anterior one-sixth and near middle, respectively, on interval 3 near stria 3. Preapical seta present, inserted 1.5 times as far from apical elytral margin as from sutural margin on interval 3 next to stria 2.</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6634FD67710C" blockId="86.[123,1174,219,1725]" box="[171,681,1604,1629]" pageId="86" pageNumber="427">Legs. Short; protibiae with longitudinal furrow.</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6615FC8C71CC" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of female apically with two pairs of paramedial setae.</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC3FFD13CFC66D4FB5875CC" lastPageId="88" lastPageNumber="429" pageId="86" pageNumber="427" type="discussion">
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<emphasis id="B90BEA99FFC3FFDF3CFC66D4FEE671EC" bold="true" box="[171,296,1701,1725]" pageId="86" pageNumber="427">COMMENTS</emphasis>
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.— This species appears to be closely related to
<taxonomicName id="4C7F4D08FFC3FFDF3F7266D7FC5A71ED" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2016" box="[805,916,1701,1725]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="427" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFC3FFDF3F7266D7FC5A71ED" box="[805,916,1701,1725]" italics="true" pageId="86" pageNumber="427">E. fortipes</emphasis>
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, but its members can be distinguished from those of the latter by the truncate, almost bifurcate, median tooth of the mentum (simple in
<taxonomicName id="4C7F4D08FFCDFFD13D76608DFE5F7644" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2016" box="[289,401,252,276]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFCDFFD13D76608DFE5F7644" box="[289,401,252,276]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members), elytral stria 5 not more deeply impressed basally (distinctly more deeply impressed in
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<emphasis id="B90BEA99FFCDFFD13DC2616CFDCA7664" box="[405,516,284,308]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members), recurrent stria longer (shorter in
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<emphasis id="B90BEA99FFCDFFD13FB6616CFB9E7664" box="[993,1104,284,308]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members) and terminated on interval 7 (terminated on interval
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<emphasis id="B90BEA99FFCDFFD13F5B614FFCB77605" box="[780,889,317,341]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members), and the preapical seta inserted 1.5 times as far from apical elytral margin as from sutural margin (inserted about equidistant from apical and sutural margins in
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<emphasis id="B90BEA99FFCDFFD13E27610EFD1176C7" box="[624,735,383,407]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members).
</paragraph>
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33.
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<emphasis id="B90BEA99FFC2FFDE3D5C6646FDCA711A" box="[267,516,1591,1610]" italics="true" pageId="87" pageNumber="428">Epaphiotrechus fortipesoides</emphasis>
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<taxonomicNameLabel id="A23857E2FFC2FFDE3E5E6646FD83711A" box="[521,589,1591,1610]" pageId="87" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1039089). b. Map of locality record (red circle) for
<taxonomicName id="4C7F4D08FFC2FFDE3C816620FE927133" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[214,348,1616,1635]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="428" phylum="Arthropoda" rank="species" species="fortipesoides">
<emphasis id="B90BEA99FFC2FFDE3C816620FE927133" box="[214,348,1616,1635]" italics="true" pageId="87" pageNumber="428">E. fortipesoides</emphasis>
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in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km.
</paragraph>
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</emphasis>
.— The unique
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female of this species was found on the ground under a stone or log in a managed conifer forest along a roadcut at an elevation of
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. No other trechines were found in this area.
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.—
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.
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species is known only from a single female specimen from the
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locality in
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in the southern part of the Gaoligong
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. This locality is in
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6
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.
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.— This species currently is known only from Tengchong County in the southern part of the Gaoligong Shan region, western
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,
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.
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</treatment>
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