317 lines
38 KiB
XML
317 lines
38 KiB
XML
<document id="975C2178EC88F7F0756098E143270781" ID-DOI="10.11646/zootaxa.3741.1.2" ID-GBIF-Dataset="6a6ba78e-a9c5-4d11-997a-3d06e5e31520" ID-ISSN="1175-5326" ID-Zenodo-Dep="223773" ID-ZooBank="BF5EA9B8-C6F4-448A-BEF9-1976AB4EC308" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460702326177" checkinUser="plazi" docAuthor="Lazányi, Eszter & Vagalinski, Boyan" docDate="2013" docId="03C887D3FFE2FFAAFF349CA4FB4CACBD" docLanguage="en" docName="zt03741p100.pdf" docOrigin="Zootaxa 3741 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaphyllum hercules Verhoeff 1901" docType="treatment" docVersion="8" lastPageNumber="84" masterDocId="FFF1FFABFFF9FFB7FFA39926FFF2AD11" masterDocTitle="Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)" masterLastPageNumber="100" masterPageNumber="55" pageNumber="82" updateTime="1698356536038" updateUser="plazi">
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<mods:title id="A1B2D4DEFFCEF75E43FF3EFD4B675B26">Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="C3E3BE399157EA49F2C3A49C2314E46E">Lazányi, Eszter</mods:namePart>
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<mods:namePart id="7B41F703FDC0B6F2D7EC7FFF306601E3">Vagalinski, Boyan</mods:namePart>
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<mods:typeOfResource id="A8ABDDE538AC79A5CD81E2B008CD04F2">text</mods:typeOfResource>
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<mods:title id="4EEBBBECBA898931DBE994CD7816927D">Zootaxa</mods:title>
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<mods:part id="D8991CEB5D1BF748A5FA4CA8CF9B3B92">
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<mods:date id="7DD57983188A8CC9D52805969BF496D8">2013</mods:date>
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<mods:number id="26EE783CDDD6FCC1C6C3D2D42F6C87BD">3741</mods:number>
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<mods:identifier id="A39FDF78DAAB4165965061222E91DA90" type="DOI">10.11646/zootaxa.3741.1.2</mods:identifier>
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<treatment id="03C887D3FFE2FFAAFF349CA4FB4CACBD" ID-DOI="http://doi.org/10.5281/zenodo.6151738" ID-GBIF-Taxon="119583540" ID-Zenodo-Dep="6151738" LSID="urn:lsid:plazi:treatment:03C887D3FFE2FFAAFF349CA4FB4CACBD" httpUri="http://treatment.plazi.org/id/03C887D3FFE2FFAAFF349CA4FB4CACBD" lastPageId="29" lastPageNumber="84" pageId="27" pageNumber="82">
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<subSubSection id="C37B654EFFE2FFACFF349CA4FC8AAB4E" pageId="27" pageNumber="82" type="nomenclature">
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<paragraph id="8BDE36C5FFE2FFACFF349CA4FD8EA88C" blockId="27.[151,636,1410,1470]" box="[151,636,1410,1437]" pageId="27" pageNumber="82">
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<heading id="D09681A9FFE2FFACFF349CA4FD8EA88C" bold="true" box="[151,636,1410,1437]" fontSize="11" level="1" pageId="27" pageNumber="82" reason="1">
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<taxonomicName id="4C614D46FFE2FFACFF349CA4FD8EA88C" ID-CoL="8KR4V" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[151,636,1410,1437]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="species" species="hercules">
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<emphasis id="B915EAD7FFE2FFACFF349CA4FD8EA88C" bold="true" box="[151,636,1410,1437]" pageId="27" pageNumber="82">
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<emphasis id="B915EAD7FFE2FFACFF349CA4FE5EA88D" bold="true" box="[151,428,1410,1436]" italics="true" pageId="27" pageNumber="82">Megaphyllum hercules</emphasis>
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(Verhoeff, 1901)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDE36C5FFE2FFACFF349C83FEE0A8AF" blockId="27.[151,636,1410,1470]" box="[151,274,1445,1470]" pageId="27" pageNumber="82">
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<figureCitation id="135A2A40FFE2FFACFF349C83FEE0A8AF" box="[151,274,1445,1470]" captionStart="FIGURES 15 a – g" captionStartId="28.[151,264,1408,1430]" captionTargetBox="[198,1393,271,1384]" captionTargetId="figure@28.[190,1397,265,1386]" captionTargetPageId="28" captionText="FIGURES 15 a – g. Megaphyllum hercules. Male from Kechrokambos, Lekanis Mts, Greece: (15 a) gonopods, posterior view; (15 b) right gonopods, mesal view; (15 c) left promere, mesal view; (15 d) left opisthomere, mesal view. Male from Demir Kapija, Republic of Macedonia: penis, (15 e) posterior and (15 f) right lateral views. (15 g) Female from the Rodopi Mts, Greece: left vulva, antero-mesal view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223787/files/figure.png" pageId="27" pageNumber="82">Figs 15a–g</figureCitation>
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</paragraph>
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<paragraph id="8BDE36C5FFE2FFACFF349CCBFC8AAB4E" blockId="27.[151,1230,1517,1754]" pageId="27" pageNumber="82">
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<taxonomicName id="4C614D46FFE2FFACFF349CCBFCC0AB12" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[151,818,1517,1539]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="subSpecies" species="unilineatus" subGenus="Chromatoiulus" subSpecies="hercules">
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<emphasis id="B915EAD7FFE2FFACFF349CCBFD63AB12" box="[151,657,1517,1539]" italics="true" pageId="27" pageNumber="82">Brachyiulus (Chromatoiulus) unilineatus hercules</emphasis>
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Verhoeff, 1901
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</taxonomicName>
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: Verhoeff 1901a: 97–98,
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<figureCitation id="135A2A40FFE2FFACFB929CCBFB54AB12" box="[1073,1190,1517,1539]" captionStart-0="FIGURES 18 b – d" captionStart-1="FIGURES 19 a – f" captionStart-2="FIGURES 20 a – g" captionStartId-0="32.[151,264,1855,1877]" captionStartId-1="33.[151,264,1333,1355]" captionStartId-2="35.[151,264,1484,1506]" captionTargetBox-0="[310,1276,1254,1831]" captionTargetBox-1="[241,1344,429,1309]" captionTargetBox-2="[266,1320,497,1467]" captionTargetId-0="figure@32.[307,1280,1253,1834]" captionTargetId-1="figure@33.[238,1349,425,1312]" captionTargetId-2="figure@35.[262,1324,493,1468]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="35" captionText-0="FIGURES 18 b – d. Megaphyllum spathulatum after Lohmander (1936): (18 b) right gonopods, mesal view; (18 c) left promere, meso-posterior view; (18 d) probably left opisthomere, lateral view. Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, sh: serrated hump, P: promere." captionText-1="FIGURES 19 a – f. Megaphyllum tauricum. Syntype male from Krim, Alupka, Inv. No. 3134: (19 a) gonopods, posterior view; (19 b) left gonopods, mesal view; (19 c) left promere, meso-posterior view; (19 d) left opisthomere, postero-lateral view. Other male from Krim, ZMB 8652: penis, (19 e) posterior and (19 f) right lateral views. Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, i: indentation, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." captionText-2="FIGURES 20 a – g. Megaphyllum unilineatum. Male from Sminthi, Xanthi county, Greece: (20 a) gonopods, posterior view; (20 b) right gonopods, mesal view; (20 c) left promere, meso-posterior view. Male from the shore of Lake Vistonida, Lagos, Greece: (20 d) left opisthomere, lateral view. Male from near Sminthi, Kamertsi Mts, Greece: penis, (20 e) posterior and (20 f) right lateral views. Female from Nestos Delta, Kavala county, Greece: (20 g) left vulva, latero-posterior view (setae omitted). Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri-0="https://zenodo.org/record/223790/files/figure.png" httpUri-1="https://zenodo.org/record/223791/files/figure.png" httpUri-2="https://zenodo.org/record/223792/files/figure.png" pageId="27" pageNumber="82">Figs 18–20</figureCitation>
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.
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<taxonomicName id="4C614D46FFE2FFACFF349F2AFD54AB33" authority="Lignau 1903: 127" authorityName="Lignau" authorityPageNumber="127" authorityYear="1903" box="[151,678,1548,1570]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="subSpecies" species="unilineatus" subSpecies="hercules">
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<emphasis id="B915EAD7FFE2FFACFF349F2AFE11AB33" box="[151,483,1548,1570]" italics="true" pageId="27" pageNumber="82">Brachyiulus unilineatus hercules</emphasis>
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: Lignau 1903: 127
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</taxonomicName>
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, 55; Verhoeff 1932: 493; 1937: 107–108, 117,
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<figureCitation id="135A2A40FFE2FFACFB239F2AFB38AB33" box="[1152,1226,1548,1570]" captionStart="FIGURES 16 a – g" captionStartId="29.[151,264,1730,1752]" captionTargetBox="[171,1417,489,1688]" captionTargetId="figure@29.[169,1423,483,1688]" captionTargetPageId="29" captionText="FIGURES 16 a – g. Megaphyllum rhodopinum. Male from near Shabran Peak, Slavyanka Mt., Bulgaria: (16 a) gonopods, posterior view; (16 b) right gonopods, mesal view; (16 c) right opisthomere, meso-posterior view; (16 d) right opisthomere, lateral view. Males from near Proastio, Kavala county, Greece: penis form (16 e) posterior and (16 f) left lateral views, (16 f *) another penis, right lateral view. Female from the first locality: (16 g) left vulva, posterior view (setae omitted). Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223788/files/figure.png" pageId="27" pageNumber="82">Fig. 16</figureCitation>
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.
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<taxonomicName id="4C614D46FFE2FFACFF349F0CFD30AB51" authority="Attems 1927: 225" authorityName="Attems" authorityPageNumber="225" authorityYear="1927" box="[151,706,1578,1600]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="subSpecies" species="unilineatus" subSpecies="hercules">
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<emphasis id="B915EAD7FFE2FFACFF349F0CFDF2AB51" box="[151,512,1578,1600]" italics="true" pageId="27" pageNumber="82">Chromatoiulus unilineatus hercules</emphasis>
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: Attems 1927: 225
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</taxonomicName>
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–226, Figs 291–293.
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<taxonomicName id="4C614D46FFE2FFACFF349F6FFC81AB4E" authority="Attems 1959: 305" authorityName="Attems" authorityPageNumber="305" authorityYear="1959" box="[151,883,1609,1631]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="subSpecies" species="unilineatus" subGenus="Chromatoiulus" subSpecies="hercules">
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<emphasis id="B915EAD7FFE2FFACFF349F6FFD5CAB4E" box="[151,686,1609,1631]" italics="true" pageId="27" pageNumber="82">Chromatoiulus (Chromatoiulus) unilineatus hercules</emphasis>
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: Attems 1959: 305
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFE2FFACFF349F4EFD8EABCB" pageId="27" pageNumber="82" type="description">
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<paragraph id="8BDE36C5FFE2FFACFF349F4EFDABAB6F" blockId="27.[151,1230,1517,1754]" box="[151,601,1640,1662]" pageId="27" pageNumber="82">
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<taxonomicName id="4C614D46FFE2FFACFF349F4EFDA7AB6F" authority="Strasser 1967: 285" authorityName="Strasser" authorityPageNumber="285" authorityYear="1967" box="[151,597,1640,1662]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="species" species="hercules">
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<emphasis id="B915EAD7FFE2FFACFF349F4EFE7BAB6F" box="[151,393,1640,1662]" italics="true" pageId="27" pageNumber="82">Chromatoiulus hercules</emphasis>
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: Strasser 1967: 285
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BDE36C5FFE2FFACFF349FA0FCF5AB8D" blockId="27.[151,1230,1517,1754]" box="[151,775,1670,1692]" pageId="27" pageNumber="82">
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<taxonomicName id="4C614D46FFE2FFACFF349FA0FCF1AB8D" authority="Strasser 1969: 159" authorityName="Strasser" authorityPageNumber="159" authorityYear="1969" box="[151,771,1670,1692]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="species" species="hercules" subGenus="Chromatoiulus">
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<emphasis id="B915EAD7FFE2FFACFF349FA0FDC5AB8D" box="[151,567,1670,1692]" italics="true" pageId="27" pageNumber="82">Chromatoiulus (Chromatoiulus) hercules</emphasis>
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: Strasser 1969: 159
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BDE36C5FFE2FFACFF349F83FD8EABCB" blockId="27.[151,1230,1517,1754]" pageId="27" pageNumber="82">
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<taxonomicName id="4C614D46FFE2FFACFF349F83FD80ABAA" authority="Enghoff & Kime 2009" authorityName="Enghoff & Kime" authorityYear="2009" box="[151,626,1701,1723]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="species" species="hercules">
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<emphasis id="B915EAD7FFE2FFACFF349F83FE8FABAA" box="[151,381,1701,1723]" italics="true" pageId="27" pageNumber="82">Megaphyllum hercules</emphasis>
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: Enghoff & Kime 2009
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</taxonomicName>
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; Lazányi
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<emphasis id="B915EAD7FFE2FFACFD779F80FCF5ABAA" box="[724,775,1701,1723]" italics="true" pageId="27" pageNumber="82">et al.</emphasis>
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2012: 13–14, 28, 41.
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<taxonomicName id="4C614D46FFE2FFACFF349FE2FD8AABCB" authority="Enghoff & Kime 2009" authorityName="Enghoff & Kime" authorityYear="2009" box="[151,632,1732,1754]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="27" pageNumber="82" phylum="Arthropoda" rank="species" species="tauricum">
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<emphasis id="B915EAD7FFE2FFACFF349FE2FE71ABCB" box="[151,387,1732,1754]" italics="true" pageId="27" pageNumber="82">Megaphyllum tauricum</emphasis>
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: Enghoff & Kime 2009
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFE2FFABFF349E20FB33ADC5" lastPageId="28" lastPageNumber="83" pageId="27" pageNumber="82" type="materials_examined">
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||
<paragraph id="8BDE36C5FFE2FFABFF349E20FB33ADC5" blockId="27.[151,1437,1798,2003]" lastBlockId="28.[151,1436,151,213]" lastPageId="28" lastPageNumber="83" pageId="27" pageNumber="82">
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||
<emphasis id="B915EAD7FFE2FFACFF349E20FE0FAA0E" bold="true" box="[151,509,1798,1823]" pageId="27" pageNumber="82">
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Material examined.
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<collectingCountry id="F3767655FFE2FFACFE369E20FE0FAA0E" box="[405,509,1798,1823]" name="Bulgaria" pageId="27" pageNumber="82">Bulgaria</collectingCountry>
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</emphasis>
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:
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<specimenCount id="9D67FD4CFFE2FFACFDB09E21FDC5AA0E" box="[531,567,1799,1823]" pageId="27" pageNumber="82" type="generic">1♂</specimenCount>
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, 1♀, Nr. 49, Sakar Mts, steppe-like and rocky grassland
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<quantity id="4C999B20FFE2FFACFB5B9E20FACDAA0E" box="[1272,1343,1798,1823]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="82" unit="km" value="0.5">0.5km</quantity>
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SW of Balgarska Polyana,
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<quantity id="4C999B20FFE2FFACFEDC9E0CFE30AA52" box="[383,450,1834,1859]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="82" unit="m" value="500.0">500m</quantity>
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,
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<geoCoordinate id="EE555002FFE2FFACFE719E0CFD91AA52" box="[466,611,1834,1859]" direction="north" orientation="latitude" pageId="27" pageNumber="82" precision="1" value="42.02252">N42°01.351’</geoCoordinate>
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<geoCoordinate id="EE555002FFE2FFACFDCC9E0CFD0DAA52" box="[623,767,1834,1859]" direction="east" orientation="longitude" pageId="27" pageNumber="82" precision="1" value="26.190834">E26°11.450’</geoCoordinate>
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, leg. Á. Ecsedi, T. Kovács, G. Puskás (HNHM);
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<specimenCount id="9D67FD4CFFE2FFACFAEA9E0DFA9CAA52" box="[1353,1390,1835,1859]" pageId="27" pageNumber="82" type="generic">1♂</specimenCount>
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, nr. Harmanli,
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<date id="FFDF1005FFE2FFACFEB19E68FE60AA76" box="[274,402,1870,1895]" pageId="27" pageNumber="82" value="1986-03-30">30.03.1986</date>
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, leg. P. Beron (NMNHS);
|
||
<emphasis id="B915EAD7FFE2FFACFD689E68FC14AA76" bold="true" box="[715,998,1870,1895]" pageId="27" pageNumber="82">
|
||
<collectingCountry id="F3767655FFE2FFACFD689E68FC14AA76" box="[715,998,1870,1895]" name="Macedonia" pageId="27" pageNumber="82">Republic of Macedonia</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D67FD4CFFE2FFACFC5B9E69FBEFAA76" box="[1016,1053,1871,1895]" pageId="27" pageNumber="82" type="generic">1♂</specimenCount>
|
||
, Kavadarci,
|
||
<date id="FFDF1005FFE2FFACFB129E69FAC0AA76" box="[1201,1330,1871,1895]" pageId="27" pageNumber="82" value="2009-09-17">17.09.2009</date>
|
||
, (4), leg. Barina, Somay, Puskás, between villages Rožden and Maden,
|
||
<quantity id="4C999B20FFE2FFACFCC69E54FC5AAA9A" box="[869,936,1906,1931]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="27" pageNumber="82" unit="m" value="750.0">750m</quantity>
|
||
,
|
||
<geoCoordinate id="EE555002FFE2FFACFC159E55FBC1AA9A" box="[950,1075,1907,1931]" direction="north" orientation="latitude" pageId="27" pageNumber="82" precision="1" value="41.17897">N41.17897</geoCoordinate>
|
||
<geoCoordinate id="EE555002FFE2FFACFB9E9E54FB4BAA9A" box="[1085,1209,1906,1931]" direction="east" orientation="longitude" pageId="27" pageNumber="82" precision="1" value="21.95191">E21.95191</geoCoordinate>
|
||
, dry grassland with junipers (HNHM); 1♀ Kavadarci,
|
||
<date id="FFDF1005FFE2FFACFD8E9EB1FD5CAABE" box="[557,686,1943,1967]" pageId="27" pageNumber="82" value="2009-09-17">17.09.2009</date>
|
||
, (5), leg. Barina, Somay, Puskás, E of village Maden (Majden, Majdan, Alsar)
|
||
<geoCoordinate id="EE555002FFE2FFACFEEA9E9DFE3CAAC2" box="[329,462,1978,2003]" direction="north" orientation="latitude" pageId="27" pageNumber="82" precision="1" value="41.15707">N 41.15707</geoCoordinate>
|
||
<geoCoordinate id="EE555002FFE2FFACFE769E9CFDA1AAC2" box="[469,595,1978,2003]" direction="east" orientation="longitude" pageId="27" pageNumber="82" precision="1" value="21.94648">E21.94648</geoCoordinate>
|
||
, dry stony grassland on hillside,
|
||
<quantity id="4C999B20FFE2FFACFC659E9CFBF4AAC2" box="[966,1030,1978,2003]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="27" pageNumber="82" unit="m" value="850.0">850m</quantity>
|
||
(HNHM); 1♀, Kočani,
|
||
<date id="FFDF1005FFE2FFACFAB49E9DFA6AAAC2" box="[1303,1432,1979,2003]" pageId="27" pageNumber="82" value="2009-09-21">21.09.2009</date>
|
||
, (21), leg. Barina, Somay, Puskás, Osogovska Planina, above town Kočani, c.
|
||
<quantity id="4C999B20FFE5FFABFBB999B1FB99ADA1" box="[1050,1131,151,176]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="28" pageNumber="83" unit="km" value="5.5">5.5 km</quantity>
|
||
NW of the town, towards village Ponikva,
|
||
<geoCoordinate id="EE555002FFE5FFABFEF7999AFE23ADC4" box="[340,465,188,213]" direction="north" orientation="latitude" pageId="28" pageNumber="83" precision="1" value="41.96677">N41.96677</geoCoordinate>
|
||
<geoCoordinate id="EE555002FFE5FFABFE7B999AFDA7ADC4" box="[472,597,188,213]" direction="east" orientation="longitude" pageId="28" pageNumber="83" precision="1" value="22.37568">E22.37568</geoCoordinate>
|
||
, disturbed grassland and stony slope,
|
||
<quantity id="4C999B20FFE5FFABFC5F999AFBB9ADC5" box="[1020,1099,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="83" unit="m" value="1000.0">1000m</quantity>
|
||
(HNHM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1E664DFFE5FFABFF349CA6FDABAB3E" httpUri="https://zenodo.org/record/223787/files/figure.png" pageId="28" pageNumber="83" targetBox="[198,1393,271,1384]" targetPageId="28">
|
||
<paragraph id="8BDE36C5FFE5FFABFF349CA6FDABAB3E" blockId="28.[151,1437,1408,1583]" pageId="28" pageNumber="83">
|
||
<emphasis id="B915EAD7FFE5FFABFF349CA6FEA7A887" bold="true" box="[151,341,1408,1430]" pageId="28" pageNumber="83">FIGURES 15a–g.</emphasis>
|
||
<taxonomicName id="4C614D46FFE5FFABFEFF9CA6FDB6A887" box="[348,580,1408,1430]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="28" pageNumber="83" phylum="Arthropoda" rank="species" species="hercules">
|
||
<emphasis id="B915EAD7FFE5FFABFEFF9CA6FDB6A887" box="[348,580,1408,1430]" italics="true" pageId="28" pageNumber="83">Megaphyllum hercules</emphasis>
|
||
</taxonomicName>
|
||
. Male from Kechrokambos, Lekanis Mts, Greece:
|
||
<emphasis id="B915EAD7FFE5FFABFBF09CA6FB7BA887" bold="true" box="[1107,1161,1408,1430]" pageId="28" pageNumber="83">(15a)</emphasis>
|
||
gonopods, posterior view;
|
||
<emphasis id="B915EAD7FFE5FFABFF349CB8FF3DA8A5" bold="true" box="[151,207,1438,1460]" pageId="28" pageNumber="83">(15b)</emphasis>
|
||
right gonopods, mesal view;
|
||
<emphasis id="B915EAD7FFE5FFABFDAF9CB8FDB0A8A5" bold="true" box="[524,578,1438,1460]" pageId="28" pageNumber="83">(15c)</emphasis>
|
||
left promere, mesal view;
|
||
<emphasis id="B915EAD7FFE5FFABFCC09CB8FC69A8A5" bold="true" box="[867,923,1438,1460]" pageId="28" pageNumber="83">(15d)</emphasis>
|
||
left opisthomere, mesal view. Male from Demir Kapija, Republic of Macedonia: penis,
|
||
<emphasis id="B915EAD7FFE5FFABFD829C9BFDA5A8C2" bold="true" box="[545,599,1469,1491]" pageId="28" pageNumber="83">(15e)</emphasis>
|
||
posterior and
|
||
<emphasis id="B915EAD7FFE5FFABFD459C9BFCEBA8C2" bold="true" box="[742,793,1469,1491]" pageId="28" pageNumber="83">(15f)</emphasis>
|
||
right lateral views.
|
||
<emphasis id="B915EAD7FFE5FFABFC429C9BFBEAA8C2" bold="true" box="[993,1048,1469,1491]" pageId="28" pageNumber="83">(15g)</emphasis>
|
||
Female from the Rodopi Mts, Greece: left vulva, antero-mesal view (setae omitted). Abbreviations:
|
||
<emphasis id="B915EAD7FFE5FFABFCAD9CFAFCC3A8E0" bold="true" box="[782,817,1500,1521]" italics="true" pageId="28" pageNumber="83">asp</emphasis>
|
||
: anterior process of the solenomere,
|
||
<emphasis id="B915EAD7FFE5FFABFB139CFAFB3AA8E0" bold="true" box="[1200,1224,1500,1521]" italics="true" pageId="28" pageNumber="83">ca</emphasis>
|
||
: central ampulla,
|
||
<emphasis id="B915EAD7FFE5FFABFA209CFAFA67A8E0" bold="true" box="[1411,1429,1500,1521]" italics="true" pageId="28" pageNumber="83">ct</emphasis>
|
||
: connecting tube;
|
||
<emphasis id="B915EAD7FFE5FFABFEF79CDCFE90AB00" bold="true" box="[340,354,1530,1553]" italics="true" pageId="28" pageNumber="83">h</emphasis>
|
||
: hump,
|
||
<emphasis id="B915EAD7FFE5FFABFE679CDDFE2FAB01" bold="true" box="[452,477,1531,1552]" italics="true" pageId="28" pageNumber="83">op</emphasis>
|
||
: operculum,
|
||
<emphasis id="B915EAD7FFE5FFABFDD19CDDFD79AB01" bold="true" box="[626,651,1531,1552]" italics="true" pageId="28" pageNumber="83">pa</emphasis>
|
||
: posterior ampulla,
|
||
<emphasis id="B915EAD7FFE5FFABFCCF9CDDFC77AB01" bold="true" box="[876,901,1531,1552]" italics="true" pageId="28" pageNumber="83">pp</emphasis>
|
||
: posterior process,
|
||
<emphasis id="B915EAD7FFE5FFABFBC19CDDFB77AB01" bold="true" box="[1122,1157,1531,1552]" italics="true" pageId="28" pageNumber="83">psp</emphasis>
|
||
: posterior process of the solenomere,
|
||
<emphasis id="B915EAD7FFE5FFABFEBB9F3CFED5AB3E" bold="true" box="[280,295,1562,1583]" italics="true" pageId="28" pageNumber="83">P</emphasis>
|
||
: promere. Scale bars: 0.2mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37B654EFFE5FFABFF649F70FE12ABA6" pageId="28" pageNumber="83" type="distribution">
|
||
<paragraph id="8BDE36C5FFE5FFABFF649F70FE12ABA6" blockId="28.[151,1437,1622,2007]" pageId="28" pageNumber="83">
|
||
<emphasis id="B915EAD7FFE5FFABFF649F70FE34AB7E" bold="true" box="[199,454,1622,1647]" pageId="28" pageNumber="83">
|
||
Distribution.
|
||
<collectingCountry id="F3767655FFE5FFABFEC49F70FE34AB7E" box="[359,454,1622,1647]" name="Albania" pageId="28" pageNumber="83">Albania</collectingCountry>
|
||
</emphasis>
|
||
(Mauriès
|
||
<emphasis id="B915EAD7FFE5FFABFD9B9F71FD83AB7F" box="[568,625,1622,1646]" italics="true" pageId="28" pageNumber="83">et al.</emphasis>
|
||
1997);
|
||
<emphasis id="B915EAD7FFE5FFABFD649F70FCDDAB7E" bold="true" box="[711,815,1622,1647]" pageId="28" pageNumber="83">
|
||
<collectingCountry id="F3767655FFE5FFABFD649F70FCDDAB7E" box="[711,815,1622,1647]" name="Bulgaria" pageId="28" pageNumber="83">Bulgaria</collectingCountry>
|
||
</emphasis>
|
||
(Vagalinski & Stoev 2007);
|
||
<emphasis id="B915EAD7FFE5FFABFBCC9F70FB3DAB7E" bold="true" box="[1135,1231,1622,1647]" pageId="28" pageNumber="83">
|
||
<collectingCountry id="F3767655FFE5FFABFBCC9F70FB3DAB7E" box="[1135,1231,1622,1647]" name="Georgia" pageId="28" pageNumber="83">Georgia</collectingCountry>
|
||
</emphasis>
|
||
: Picunda (Issajew 1911; Kobakhidze 1965);
|
||
<emphasis id="B915EAD7FFE5FFABFE1D9F5CFDE3AB82" bold="true" box="[446,529,1658,1683]" pageId="28" pageNumber="83">
|
||
<collectingCountry id="F3767655FFE5FFABFE1D9F5CFDE3AB82" box="[446,529,1658,1683]" name="Greece" pageId="28" pageNumber="83">Greece</collectingCountry>
|
||
</emphasis>
|
||
(Strasser 1974, 1976);
|
||
<emphasis id="B915EAD7FFE5FFABFCBF9F5CFBC1AB82" bold="true" box="[796,1075,1658,1683]" pageId="28" pageNumber="83">
|
||
<collectingCountry id="F3767655FFE5FFABFCBF9F5CFBC1AB82" box="[796,1075,1658,1683]" name="Macedonia" pageId="28" pageNumber="83">Republic of Macedonia</collectingCountry>
|
||
</emphasis>
|
||
(Makarov
|
||
<emphasis id="B915EAD7FFE5FFABFB119F5DFB19AB82" box="[1202,1259,1659,1683]" italics="true" pageId="28" pageNumber="83">et al.</emphasis>
|
||
2004);
|
||
<emphasis id="B915EAD7FFE5FFABFAE69F5CFA66AB82" bold="true" box="[1349,1428,1658,1683]" pageId="28" pageNumber="83">
|
||
<collectingCountry id="F3767655FFE5FFABFAE69F5CFA66AB82" box="[1349,1428,1658,1683]" name="Russia" pageId="28" pageNumber="83">Russia</collectingCountry>
|
||
</emphasis>
|
||
: Novorossiysk (Lignau 1903).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B654EFFE5FFABFF649FE4FC53AAC6" pageId="28" pageNumber="83" type="diagnosis">
|
||
<paragraph id="8BDE36C5FFE5FFABFF649FE4FE07AA32" blockId="28.[151,1437,1622,2007]" pageId="28" pageNumber="83">
|
||
<emphasis id="B915EAD7FFE5FFABFF649FE4FEB3ABCA" bold="true" box="[199,321,1730,1755]" pageId="28" pageNumber="83">Diagnosis.</emphasis>
|
||
Differs from all consubgeners by the presence of a large basal hump (
|
||
<emphasis id="B915EAD7FFE5FFABFBC89FE4FB88ABCA" bold="true" box="[1131,1146,1730,1755]" italics="true" pageId="28" pageNumber="83">h</emphasis>
|
||
) mesally on opisthomere (
|
||
<figureCitation id="135A2A40FFE5FFABFF3C9FC0FEA7ABEE" box="[159,341,1766,1791]" captionStart="FIGURES 15 a – g" captionStartId="28.[151,264,1408,1430]" captionTargetBox="[198,1393,271,1384]" captionTargetId="figure@28.[190,1397,265,1386]" captionTargetPageId="28" captionText="FIGURES 15 a – g. Megaphyllum hercules. Male from Kechrokambos, Lekanis Mts, Greece: (15 a) gonopods, posterior view; (15 b) right gonopods, mesal view; (15 c) left promere, mesal view; (15 d) left opisthomere, mesal view. Male from Demir Kapija, Republic of Macedonia: penis, (15 e) posterior and (15 f) right lateral views. (15 g) Female from the Rodopi Mts, Greece: left vulva, antero-mesal view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223787/files/figure.png" pageId="28" pageNumber="83">Figs 15a–b, 15d</figureCitation>
|
||
) and from other members of the species group by the very broad, rounded posterior opisthomere process (
|
||
<emphasis id="B915EAD7FFE5FFABFF589E2DFEE5AA33" bold="true" box="[251,279,1803,1826]" italics="true" pageId="28" pageNumber="83">pp</emphasis>
|
||
) (
|
||
<figureCitation id="135A2A40FFE5FFABFE8C9E2CFE15AA32" box="[303,487,1802,1827]" captionStart="FIGURES 15 a – g" captionStartId="28.[151,264,1408,1430]" captionTargetBox="[198,1393,271,1384]" captionTargetId="figure@28.[190,1397,265,1386]" captionTargetPageId="28" captionText="FIGURES 15 a – g. Megaphyllum hercules. Male from Kechrokambos, Lekanis Mts, Greece: (15 a) gonopods, posterior view; (15 b) right gonopods, mesal view; (15 c) left promere, mesal view; (15 d) left opisthomere, mesal view. Male from Demir Kapija, Republic of Macedonia: penis, (15 e) posterior and (15 f) right lateral views. (15 g) Female from the Rodopi Mts, Greece: left vulva, antero-mesal view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223787/files/figure.png" pageId="28" pageNumber="83">Figs 15a–b, 15d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFE5FFABFF649E08FC34AA9E" blockId="28.[151,1437,1622,2007]" pageId="28" pageNumber="83">
|
||
Opisthomere’s (
|
||
<figureCitation id="135A2A40FFE5FFABFEDD9E08FDCDAA56" box="[382,575,1838,1863]" captionStart="FIGURES 15 a – g" captionStartId="28.[151,264,1408,1430]" captionTargetBox="[198,1393,271,1384]" captionTargetId="figure@28.[190,1397,265,1386]" captionTargetPageId="28" captionText="FIGURES 15 a – g. Megaphyllum hercules. Male from Kechrokambos, Lekanis Mts, Greece: (15 a) gonopods, posterior view; (15 b) right gonopods, mesal view; (15 c) left promere, mesal view; (15 d) left opisthomere, mesal view. Male from Demir Kapija, Republic of Macedonia: penis, (15 e) posterior and (15 f) right lateral views. (15 g) Female from the Rodopi Mts, Greece: left vulva, antero-mesal view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223787/files/figure.png" pageId="28" pageNumber="83">Figs 15a–b, 15d</figureCitation>
|
||
) posterior process (
|
||
<emphasis id="B915EAD7FFE5FFABFC8E9E09FCBBAA57" bold="true" box="[813,841,1839,1862]" italics="true" pageId="28" pageNumber="83">pp</emphasis>
|
||
) broad, rounded and shorter than the solenomere. Anterior process (
|
||
<emphasis id="B915EAD7FFE5FFABFEC49E75FE71AA7B" bold="true" box="[359,387,1875,1898]" italics="true" pageId="28" pageNumber="83">ap</emphasis>
|
||
) and anterior hump (
|
||
<emphasis id="B915EAD7FFE5FFABFDDF9E74FD6BAA7A" bold="true" box="[636,665,1874,1899]" italics="true" pageId="28" pageNumber="83">ah</emphasis>
|
||
) hardly detectable. Medially the opisthomeres reaching each other with a large basal hump (
|
||
<emphasis id="B915EAD7FFE5FFABFE119E50FE33AA9E" bold="true" box="[434,449,1910,1935]" italics="true" pageId="28" pageNumber="83">h</emphasis>
|
||
). Vulva robust, bursa skull-shaped (
|
||
<figureCitation id="135A2A40FFE5FFABFCF49E50FC4BAA9E" box="[855,953,1910,1935]" captionStart="FIGURES 15 a – g" captionStartId="28.[151,264,1408,1430]" captionTargetBox="[198,1393,271,1384]" captionTargetId="figure@28.[190,1397,265,1386]" captionTargetPageId="28" captionText="FIGURES 15 a – g. Megaphyllum hercules. Male from Kechrokambos, Lekanis Mts, Greece: (15 a) gonopods, posterior view; (15 b) right gonopods, mesal view; (15 c) left promere, mesal view; (15 d) left opisthomere, mesal view. Male from Demir Kapija, Republic of Macedonia: penis, (15 e) posterior and (15 f) right lateral views. (15 g) Female from the Rodopi Mts, Greece: left vulva, antero-mesal view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223787/files/figure.png" pageId="28" pageNumber="83">
|
||
Fig.
|
||
<quantity id="4C999B20FFE5FFABFC2F9E50FC4BAA9E" box="[908,953,1910,1935]" metricMagnitude="-2" metricUnit="kg" metricValue="1.5" pageId="28" pageNumber="83" unit="g" value="15.0">15g</quantity>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFE5FFABFF649EBCFC53AAC6" blockId="28.[151,1437,1622,2007]" pageId="28" pageNumber="83">
|
||
Both males and females dark grey with a bright or reddish dorsal longitudinal band. Body length and height: males:
|
||
<quantity id="4C999B20FFE5FFABFF449E98FE77AAC6" box="[231,389,1982,2007]" metricMagnitude="-2" metricUnit="m" metricValue="4.1450000000000005" metricValueMax="4.720000000000001" metricValueMin="3.5700000000000003" pageId="28" pageNumber="83" unit="mm" value="41.45" valueMax="47.2" valueMin="35.7">35.7–47.2mm</quantity>
|
||
,
|
||
<quantity id="4C999B20FFE5FFABFE339E98FDE1AAC6" box="[400,531,1982,2007]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="28" pageNumber="83" unit="mm" value="2.95" valueMax="3.5" valueMin="2.4">2.4–3.5mm</quantity>
|
||
; females:
|
||
<quantity id="4C999B20FFE5FFABFD279E98FCD6AAC6" box="[644,804,1982,2007]" metricMagnitude="-2" metricUnit="m" metricValue="4.555" metricValueMax="5.46" metricValueMin="3.65" pageId="28" pageNumber="83" unit="mm" value="45.55" valueMax="54.6" valueMin="36.5">36.5–54.6mm</quantity>
|
||
;
|
||
<quantity id="4C999B20FFE5FFABFC8C9E99FC6FAAC6" box="[815,925,1983,2007]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.7" metricValueMin="3.0" pageId="28" pageNumber="83" unit="mm" value="3.85" valueMax="4.7" valueMin="3.0">3–4.7mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B654EFFE4FFAAFF6499B1FB4CACBD" pageId="29" pageNumber="84" type="discussion">
|
||
<paragraph id="8BDE36C5FFE4FFAAFF6499B1FABCAC74" blockId="29.[151,1436,151,429]" pageId="29" pageNumber="84">
|
||
<emphasis id="B915EAD7FFE4FFAAFF6499B1FEC9ADA1" bold="true" box="[199,315,151,176]" pageId="29" pageNumber="84">Remarks.</emphasis>
|
||
Apart from the Balkan Peninsula this species was reported from the western coasts of the Black Sea:
|
||
<collectingCountry id="F3767655FFE4FFAAFF34999AFF03ADC4" box="[151,241,188,213]" name="Georgia" pageId="29" pageNumber="84">Georgia</collectingCountry>
|
||
(Issajew 1911) and
|
||
<collectingCountry id="F3767655FFE4FFAAFE40999AFDDCADC4" box="[483,558,188,213]" name="Russia" pageId="29" pageNumber="84">Russia</collectingCountry>
|
||
(Lignau 1903). Though Lignau referred to a single female, Issajew (1911) found numerous individuals. Thus, Lohmander (1936) was sure about the existence of the species in the westernmost border of the Caucasus and stated that as
|
||
<taxonomicName id="4C614D46FFE4FFAAFCB99823FC53AC0D" box="[794,929,260,284]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="29" pageNumber="84" phylum="Arthropoda" rank="species" species="hercules">
|
||
<emphasis id="B915EAD7FFE4FFAAFCB99823FC53AC0D" box="[794,929,260,284]" italics="true" pageId="29" pageNumber="84">M. hercules</emphasis>
|
||
</taxonomicName>
|
||
differed considerably from
|
||
<taxonomicName id="4C614D46FFE4FFAAFB4A9823FE8AAC51" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="29" pageNumber="84" phylum="Arthropoda" rank="species" species="spathulatum">
|
||
<emphasis id="B915EAD7FFE4FFAAFB4A9823FA6EAC0D" box="[1257,1436,260,284]" italics="true" pageId="29" pageNumber="84">M. spathulatum</emphasis>
|
||
(Lohmander, 1936)
|
||
</taxonomicName>
|
||
, mistaking them would have been unlikely. Without revision of Lignau’s, Issajew’s and Kobakhidze’s material we can neither confirm nor deny the species’ presence outside the Balkan Peninsula.
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFE4FFAAFF649857FB4CACBD" blockId="29.[151,1436,151,429]" pageId="29" pageNumber="84">
|
||
<taxonomicName id="4C614D46FFE4FFAAFF649857FEB8AC99" box="[199,330,368,392]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="29" pageNumber="84" phylum="Arthropoda" rank="species" species="hercules">
|
||
<emphasis id="B915EAD7FFE4FFAAFF649857FEB8AC99" box="[199,330,368,392]" italics="true" pageId="29" pageNumber="84">M. hercules</emphasis>
|
||
</taxonomicName>
|
||
was originally described as subspecies of
|
||
<taxonomicName id="4C614D46FFE4FFAAFC849857FB69AC99" authority="(Verhoeff 1901a)" baseAuthorityName="Verhoeff" baseAuthorityYear="1901" box="[807,1179,367,392]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="29" pageNumber="84" phylum="Arthropoda" rank="species" species="unilineatum">
|
||
<emphasis id="B915EAD7FFE4FFAAFC849857FC21AC99" box="[807,979,368,392]" italics="true" pageId="29" pageNumber="84">M. unilineatum</emphasis>
|
||
(Verhoeff 1901a)
|
||
</taxonomicName>
|
||
, but was later elevated to species level by Strasser (1967). This was supported with new data by Lazányi
|
||
<emphasis id="B915EAD7FFE4FFAAFB8F98B3FB97ACBD" box="[1068,1125,404,428]" italics="true" pageId="29" pageNumber="84">et al.</emphasis>
|
||
(2012).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |