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<mods:title id="D97714A66751F9B651E09901B62B4A55">Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J / K boundary</mods:title>
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<treatment id="03C887B9FF9DFFC676D4A7CAFAFEFAFC" ID-DOI="http://doi.org/10.5281/zenodo.6929056" ID-GBIF-Taxon="197866766" ID-Zenodo-Dep="6929056" LSID="urn:lsid:plazi:treatment:03C887B9FF9DFFC676D4A7CAFAFEFAFC" httpUri="http://treatment.plazi.org/id/03C887B9FF9DFFC676D4A7CAFAFEFAFC" lastPageId="45" lastPageNumber="725" pageId="43" pageNumber="724">
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<taxonomicName id="4C614D2CFF9DFFC076D4A7CAFB06F9BC" baseAuthorityName="Sigogneau-Russell &amp; Ensom" baseAuthorityYear="1998" box="[1006,1261,1522,1549]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="43" pageNumber="724" phylum="Chordata" rank="species" species="taraktes">
<emphasis id="B915EABDFF9DFFC076D4A7CAFB9BF9BC" bold="true" box="[1006,1136,1522,1549]" italics="true" pageId="43" pageNumber="724">Thereuodon</emphasis>
cf. 
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<paragraph id="8BDE36AFFF9DFFC07111A42BFB5AF99C" blockId="43.[1067,1201,1554,1581]" box="[1067,1201,1554,1581]" pageId="43" pageNumber="724">
(
<figureCitation id="135A2A2AFF9DFFC0710EA42AFB6BF99C" box="[1076,1152,1554,1581]" captionStart="FIG" captionStartId="46.[132,143,1963,1980]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-21@46.[132,1455,215,1878]" captionTargetPageId="46" captionText="FIG. 32. — Mammal teeth from Angeac-Charente: A, B, premolariform or molariform tooth of Gobiconodon? sp. (ANG M-21) in occlusal (A) and labial (B) views; C, D, premolariform tooth of Mammalia indet. (ANG M-34) in lingual (C) and labial (D) views; E, F, left lower molar of Triconodon sp. (ANG M-02) in lingual (E) and occlusal (F) views; G-I, left upper molar of Thereuodon cf. taraktes (ANG M-23) in labial (G), lingual (H) and occlusal (I) views. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902113" httpUri="https://zenodo.org/record/6902113/files/figure.png" pageId="43" pageNumber="724">Fig. 32</figureCitation>
G-I)
</paragraph>
<paragraph id="8BDE36AFFF9DFFC07617A46BFC53F9DC" blockId="43.[811,1457,1619,2028]" box="[813,952,1619,1645]" pageId="43" pageNumber="724">DESCRIPTION</paragraph>
<paragraph id="8BDE36AFFF9DFFC77617A44AFEBBF89D" blockId="43.[811,1457,1619,2028]" lastBlockId="44.[132,775,1714,2028]" lastPageId="44" lastPageNumber="725" pageId="43" pageNumber="724">
An isolated tooth crown (
<figureCitation id="135A2A2AFF9DFFC07101A44AFB6EF93D" box="[1083,1157,1650,1676]" captionStart="FIG" captionStartId="46.[132,143,1963,1980]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-21@46.[132,1455,215,1878]" captionTargetPageId="46" captionText="FIG. 32. — Mammal teeth from Angeac-Charente: A, B, premolariform or molariform tooth of Gobiconodon? sp. (ANG M-21) in occlusal (A) and labial (B) views; C, D, premolariform tooth of Mammalia indet. (ANG M-34) in lingual (C) and labial (D) views; E, F, left lower molar of Triconodon sp. (ANG M-02) in lingual (E) and occlusal (F) views; G-I, left upper molar of Thereuodon cf. taraktes (ANG M-23) in labial (G), lingual (H) and occlusal (I) views. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902113" httpUri="https://zenodo.org/record/6902113/files/figure.png" pageId="43" pageNumber="724">Fig. 32</figureCitation>
G-I) is assigned to the genus 
<taxonomicName id="4C614D2CFF9DFFC07611A4AAFC4CF91D" authorityName="Sigogneau-Russell" authorityYear="1989" box="[811,935,1682,1708]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="724" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9DFFC07611A4AAFC4CF91D" box="[811,935,1682,1708]" italics="true" pageId="43" pageNumber="724">Thereuodon</emphasis>
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of the monogeneric family 
<taxonomicName id="4C614D2CFF9DFFC071D3A4AAFA41F91D" authorityName="Sigogneau-Russell &amp; Ensom" authorityYear="1998" box="[1257,1450,1682,1708]" class="Mammalia" family="Thereuodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="724" phylum="Chordata" rank="family">Thereuodontidae</taxonomicName>
, on the basis of several characters (Sigogneau-Russell 1989; Sigogneau-Russell &amp; Ensom 1998; Kielan-Jaworowska 
<emphasis id="B915EABDFF9DFFC07047A4EBFA5BF95D" box="[1405,1456,1746,1772]" italics="true" pageId="43" pageNumber="724">et al.</emphasis>
2004): it is labio-lingually narrow, with a great development of the parastylar lobe, forming almost a basin (
<figureCitation id="135A2A2AFF9DFFC0700EA52AFA67F89D" box="[1332,1420,1810,1836]" captionStart="FIG" captionStartId="46.[132,143,1963,1980]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-21@46.[132,1455,215,1878]" captionTargetPageId="46" captionText="FIG. 32. — Mammal teeth from Angeac-Charente: A, B, premolariform or molariform tooth of Gobiconodon? sp. (ANG M-21) in occlusal (A) and labial (B) views; C, D, premolariform tooth of Mammalia indet. (ANG M-34) in lingual (C) and labial (D) views; E, F, left lower molar of Triconodon sp. (ANG M-02) in lingual (E) and occlusal (F) views; G-I, left upper molar of Thereuodon cf. taraktes (ANG M-23) in labial (G), lingual (H) and occlusal (I) views. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902113" httpUri="https://zenodo.org/record/6902113/files/figure.png" pageId="43" pageNumber="724">Fig. 32I</figureCitation>
); a small vertical concavity can be seen on the anterior face of the paracone; the stylocone is well-developed (
<figureCitation id="135A2A2AFF9DFFC071D5A569FABBF8DD" box="[1263,1360,1873,1900]" captionStart="FIG" captionStartId="46.[132,143,1963,1980]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-21@46.[132,1455,215,1878]" captionTargetPageId="46" captionText="FIG. 32. — Mammal teeth from Angeac-Charente: A, B, premolariform or molariform tooth of Gobiconodon? sp. (ANG M-21) in occlusal (A) and labial (B) views; C, D, premolariform tooth of Mammalia indet. (ANG M-34) in lingual (C) and labial (D) views; E, F, left lower molar of Triconodon sp. (ANG M-02) in lingual (E) and occlusal (F) views; G-I, left upper molar of Thereuodon cf. taraktes (ANG M-23) in labial (G), lingual (H) and occlusal (I) views. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902113" httpUri="https://zenodo.org/record/6902113/files/figure.png" pageId="43" pageNumber="724">Fig. 32G</figureCitation>
); cusp D occurs just behind the stylocone, although it is heavily abraded on this specimen (
<figureCitation id="135A2A2AFF9DFFC076D6A5A9FBA6F81A" box="[1004,1101,1937,1963]" captionStart="FIG" captionStartId="46.[132,143,1963,1980]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-21@46.[132,1455,215,1878]" captionTargetPageId="46" captionText="FIG. 32. — Mammal teeth from Angeac-Charente: A, B, premolariform or molariform tooth of Gobiconodon? sp. (ANG M-21) in occlusal (A) and labial (B) views; C, D, premolariform tooth of Mammalia indet. (ANG M-34) in lingual (C) and labial (D) views; E, F, left lower molar of Triconodon sp. (ANG M-02) in lingual (E) and occlusal (F) views; G-I, left upper molar of Thereuodon cf. taraktes (ANG M-23) in labial (G), lingual (H) and occlusal (I) views. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902113" httpUri="https://zenodo.org/record/6902113/files/figure.png" pageId="43" pageNumber="724">Fig. 32G</figureCitation>
); a vertical ridge, the mediocrista, links the paracone to the cusp D, although it is weakly pronounced (
<figureCitation id="135A2A2AFF9DFFC076A7A5E9FC1DF85D" box="[925,1014,2001,2028]" captionStart="FIG" captionStartId="46.[132,143,1963,1980]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-21@46.[132,1455,215,1878]" captionTargetPageId="46" captionText="FIG. 32. — Mammal teeth from Angeac-Charente: A, B, premolariform or molariform tooth of Gobiconodon? sp. (ANG M-21) in occlusal (A) and labial (B) views; C, D, premolariform tooth of Mammalia indet. (ANG M-34) in lingual (C) and labial (D) views; E, F, left lower molar of Triconodon sp. (ANG M-02) in lingual (E) and occlusal (F) views; G-I, left upper molar of Thereuodon cf. taraktes (ANG M-23) in labial (G), lingual (H) and occlusal (I) views. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902113" httpUri="https://zenodo.org/record/6902113/files/figure.png" pageId="43" pageNumber="724">Fig. 32I</figureCitation>
). Thus, three transversely narrow basins are present in occlusal view. Moreover, the paracone is high compared to the labial wall and labially but not posteriorly recurved (
<figureCitation id="135A2A2AFF9AFFC775CAA4CAFE82F8BD" box="[240,361,1778,1804]" captionStart="FIG" captionStartId="46.[132,143,1963,1980]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-21@46.[132,1455,215,1878]" captionTargetPageId="46" captionText="FIG. 32. — Mammal teeth from Angeac-Charente: A, B, premolariform or molariform tooth of Gobiconodon? sp. (ANG M-21) in occlusal (A) and labial (B) views; C, D, premolariform tooth of Mammalia indet. (ANG M-34) in lingual (C) and labial (D) views; E, F, left lower molar of Triconodon sp. (ANG M-02) in lingual (E) and occlusal (F) views; G-I, left upper molar of Thereuodon cf. taraktes (ANG M-23) in labial (G), lingual (H) and occlusal (I) views. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902113" httpUri="https://zenodo.org/record/6902113/files/figure.png" pageId="44" pageNumber="725">Fig. 32G, I</figureCitation>
). This crown is well-preserved, lacking only the metastyle.
</paragraph>
</subSubSection>
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<paragraph id="8BDE36AFFF9AFFC775BEA401FC8BF9C6" blockId="44.[132,1457,1593,1655]" pageId="44" pageNumber="725">
FIG. 30. — Ornithomimosaur remains from Angeac-Charente: 
<emphasis id="B915EABDFF9AFFC777B1A401FD46F9FB" bold="true" box="[651,685,1593,1610]" pageId="44" pageNumber="725">A -C</emphasis>
, proximal end of right tibia (ANG12-1893) in posterior (
<emphasis id="B915EABDFF9AFFC77145A401FB67F9FB" bold="true" box="[1151,1164,1593,1610]" pageId="44" pageNumber="725">A</emphasis>
), anterior (
<emphasis id="B915EABDFF9AFFC771DDA401FB1FF9FB" bold="true" box="[1255,1268,1593,1610]" pageId="44" pageNumber="725">B</emphasis>
) and lateral (
<emphasis id="B915EABDFF9AFFC77058A401FA9BF9FB" bold="true" box="[1378,1392,1593,1610]" pageId="44" pageNumber="725">C</emphasis>
) views; 
<emphasis id="B915EABDFF9AFFC775BEA477FF79F9D1" bold="true" box="[132,146,1615,1632]" pageId="44" pageNumber="725">D</emphasis>
, distal end of right tibia (ANG10-56) in anterior view; 
<emphasis id="B915EABDFF9AFFC77771A477FDBCF9D1" bold="true" box="[587,599,1615,1632]" pageId="44" pageNumber="725">E</emphasis>
, proximal end of right fibula (ANG11-696) in medial view; 
<emphasis id="B915EABDFF9AFFC7710DA477FBBCF9D1" bold="true" box="[1079,1111,1615,1632]" pageId="44" pageNumber="725">F -H</emphasis>
, Left astragalus and calcaneum (ANG12- 1803) in anterior (
<emphasis id="B915EABDFF9AFFC7742FA45EFECBF9C6" bold="true" box="[277,288,1638,1655]" pageId="44" pageNumber="725">F</emphasis>
), posterior (
<emphasis id="B915EABDFF9AFFC774BEA45EFE79F9C6" bold="true" box="[388,402,1638,1655]" pageId="44" pageNumber="725">G</emphasis>
) and dorsal (
<emphasis id="B915EABDFF9AFFC774C4A45EFDE7F9C6" bold="true" box="[510,524,1638,1655]" pageId="44" pageNumber="725">H</emphasis>
) views. Scale bar: A-E, 4 cm; F-H, 2 cm.
</paragraph>
</caption>
<subSubSection id="C37B6524FF9AFFC675A1A50AFAFEFAFC" lastPageId="45" lastPageNumber="726" pageId="44" pageNumber="725" type="discussion">
<paragraph id="8BDE36AFFF9AFFC775A1A50AFAD9F81D" blockId="44.[132,775,1714,2028]" lastBlockId="44.[811,1458,1714,2028]" pageId="44" pageNumber="725">
Two species are known for the genus 
<taxonomicName id="4C614D2CFF9AFFC77769A509FD3AF8FA" authorityName="Sigogneau-Russell" authorityYear="1989" box="[595,721,1841,1867]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9AFFC77769A509FD3AF8FA" box="[595,721,1841,1867]" italics="true" pageId="44" pageNumber="725">Thereuodon</emphasis>
</taxonomicName>
: the type-species 
<taxonomicName id="4C614D2CFF9AFFC77429A56AFD2CF8DD" authority="(Sigogneau-Russell 1989)" baseAuthorityName="Sigogneau-Russell" baseAuthorityYear="1989" box="[275,711,1873,1900]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="dahmanii">
<emphasis id="B915EABDFF9AFFC77429A56AFE77F8DA" box="[275,412,1873,1899]" italics="true" pageId="44" pageNumber="725">T. dahmanii</emphasis>
(Sigogneau-Russell 1989)
</taxonomicName>
from the Jurassic/Cretaceous transition of the Ksar Metlili Formation, 
<collectingCountry id="F376763FFF9AFFC775DDA5AAFEBBF81D" box="[231,336,1938,1964]" name="Morocco" pageId="44" pageNumber="725">Morocco</collectingCountry>
, and 
<taxonomicName id="4C614D2CFF9AFFC774AEA5AAFEF7F87D" authority="(Sigogneau-Russell &amp; Ensom 1998)" baseAuthorityName="Sigogneau-Russell &amp; Ensom" baseAuthorityYear="1998" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="taraktes">
<emphasis id="B915EABDFF9AFFC774AEA5AAFDE3F81A" box="[404,520,1937,1963]" italics="true" pageId="44" pageNumber="725">T. taraktes</emphasis>
(Sigogneau-Russell &amp; Ensom 1998)
</taxonomicName>
, from the Early Cretaceous (Berriasian) Purbeck Group, 
<collectingCountry id="F376763FFF9AFFC77422A5E9FE92F85A" box="[280,377,2001,2027]" name="United Kingdom" pageId="44" pageNumber="725">England</collectingCountry>
. In the Angeac-Charente specimen, the great protrusion of the parastylar cusp and the weakly marked mediacrista are reminiscent of 
<taxonomicName id="4C614D2CFF9AFFC771DAA4EBFAA4F95D" baseAuthorityName="Sigogneau-Russell &amp; Ensom" baseAuthorityYear="1998" box="[1248,1359,1746,1772]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="taraktes">
<emphasis id="B915EABDFF9AFFC771DAA4EBFAA4F95D" box="[1248,1359,1746,1772]" italics="true" pageId="44" pageNumber="725">T. taraktes</emphasis>
</taxonomicName>
, but the well-developed metacone is close to what is observed in 
<taxonomicName id="4C614D2CFF9AFFC770A3A4CBFC7EF89D" baseAuthorityName="Sigogneau-Russell" baseAuthorityYear="1989" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="dahmanii">
<emphasis id="B915EABDFF9AFFC770A3A4CBFC7EF89D" italics="true" pageId="44" pageNumber="725">T. dahmanii</emphasis>
</taxonomicName>
. The stylocone is closer to cusp D and less sharp than in 
<taxonomicName id="4C614D2CFF9AFFC77644A50AFBEBF8FA" baseAuthorityName="Sigogneau-Russell" baseAuthorityYear="1989" box="[894,1024,1841,1867]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="dahmanii">
<emphasis id="B915EABDFF9AFFC77644A50AFBEBF8FA" box="[894,1024,1841,1867]" italics="true" pageId="44" pageNumber="725">T. dahmanii</emphasis>
</taxonomicName>
, but this conformation is similar to that of 
<taxonomicName id="4C614D2CFF9AFFC77673A56AFC5CF8DA" baseAuthorityName="Sigogneau-Russell &amp; Ensom" baseAuthorityYear="1998" box="[841,951,1873,1899]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="taraktes">
<emphasis id="B915EABDFF9AFFC77673A56AFC5CF8DA" box="[841,951,1873,1899]" italics="true" pageId="44" pageNumber="725">T. taraktes</emphasis>
</taxonomicName>
. Finally, as in 
<taxonomicName id="4C614D2CFF9AFFC77169A56AFB2AF8DA" baseAuthorityName="Sigogneau-Russell &amp; Ensom" baseAuthorityYear="1998" box="[1107,1217,1873,1899]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="taraktes">
<emphasis id="B915EABDFF9AFFC77169A56AFB2AF8DA" box="[1107,1217,1873,1899]" italics="true" pageId="44" pageNumber="725">T. taraktes</emphasis>
</taxonomicName>
but unlike 
<taxonomicName id="4C614D2CFF9AFFC77005A56AFCA0F83A" baseAuthorityName="Sigogneau-Russell" baseAuthorityYear="1989" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="dahmanii">
<emphasis id="B915EABDFF9AFFC77005A56AFCA0F83A" italics="true" pageId="44" pageNumber="725">T. dahmanii</emphasis>
</taxonomicName>
, there is no cuspule “c”. In conclusion, we tentatively assign this specimen to 
<taxonomicName id="4C614D2CFF9AFFC7710EA5A9FAC0F81A" baseAuthorityName="Sigogneau-Russell &amp; Ensom" baseAuthorityYear="1998" box="[1076,1323,1937,1963]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="44" pageNumber="725" phylum="Chordata" rank="species" species="taraktes">
<emphasis id="B915EABDFF9AFFC7710EA5A9FB44F81A" box="[1076,1199,1937,1963]" italics="true" pageId="44" pageNumber="725">Thereuodon</emphasis>
cf. 
<emphasis id="B915EABDFF9AFFC771E3A5A9FAC0F81A" box="[1241,1323,1937,1963]" italics="true" pageId="44" pageNumber="725">taraktes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDE36AFFF9AFFC67679A589FAFEFAFC" blockId="44.[811,1458,1714,2028]" lastBlockId="45.[811,1455,1267,1357]" lastPageId="45" lastPageNumber="726" pageId="44" pageNumber="725">
Sigogneau-Russell (1989) and Sigogneau-Russell &amp; Ensom (1998) considered the teeth they referred to the two species of 
<taxonomicName id="4C614D2CFF9BFFC67415A6CBFE45FABC" authorityName="Sigogneau-Russell" authorityYear="1989" box="[303,430,1267,1293]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC67415A6CBFE45FABC" box="[303,430,1267,1293]" italics="true" pageId="45" pageNumber="726">Thereuodon</emphasis>
</taxonomicName>
as “symmetrodontan” permanent molars. However, the upper deciduous premolars of 
<taxonomicName id="4C614D2CFF9BFFC675B9A70BFE8EFAFC" authorityName="Martin" authorityYear="2002" box="[131,357,1331,1357]" class="Mammalia" genus="Nanolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="species" species="drescherae">
<emphasis id="B915EABDFF9BFFC675B9A70BFE8EFAFC" box="[131,357,1331,1357]" italics="true" pageId="45" pageNumber="726">Nanolestes drescherae</emphasis>
</taxonomicName>
, a stem-Zatheria from the Late Jurassic of Guimarota, 
<collectingCountry id="F376763FFF9BFFC67461A76BFE50FADC" box="[347,443,1363,1389]" name="Portugal" pageId="45" pageNumber="726">Portugal</collectingCountry>
(
<bibRefCitation id="EFF04B5EFF9BFFC674F5A76BFD8EFADC" author="Martin" box="[463,613,1363,1389]" firstAuthor="Martin" pageId="45" pageNumber="726" pagination="1 - 119" refId="ref43478" refString="MARTIN T. 1999. - Dryolestidae (Dryolestoidea, Mammalia) aus dem Oberen Jura-von Portugal. Abhandlungen der senckenbergischen naturforschenden Gesellschaft 550: 1 - 119." type="journal article" year="1999">Martin 1999</bibRefCitation>
, 
<bibRefCitation id="EFF04B5EFF9BFFC6774DA76BFD59FADC" author="Martin" box="[631,690,1363,1389]" firstAuthor="Martin" pageId="45" pageNumber="726" pagination="322 - 348" refId="ref43507" refString="MARTIN T. 2002. - New stem lineage representatives of Zatheria (Mammalia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 22: 322 - 348. https: // doi. org / cb 2 w 55" type="journal article" year="2002">2002</bibRefCitation>
), show some similarities with the teeth of 
<taxonomicName id="4C614D2CFF9BFFC67734A74BFD66FA3C" authorityName="Sigogneau-Russell" authorityYear="1989" box="[526,653,1395,1421]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC67734A74BFD66FA3C" box="[526,653,1395,1421]" italics="true" pageId="45" pageNumber="726">Thereuodon</emphasis>
</taxonomicName>
. Thus, the DP?3-5 of 
<taxonomicName id="4C614D2CFF9BFFC675D5A7ACFE94FA1C" authorityName="Martin" authorityYear="2002" box="[239,383,1427,1453]" class="Mammalia" genus="Nanolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="species" species="drescherae">
<emphasis id="B915EABDFF9BFFC675D5A7ACFE94FA1C" box="[239,383,1427,1453]" italics="true" pageId="45" pageNumber="726">N. drescherae</emphasis>
</taxonomicName>
share with the teeth of 
<taxonomicName id="4C614D2CFF9BFFC677BDA7ABFCEEFA1C" authorityName="Sigogneau-Russell" authorityYear="1989" box="[647,773,1427,1453]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC677BDA7ABFCEEFA1C" box="[647,773,1427,1453]" italics="true" pageId="45" pageNumber="726">Thereuodon</emphasis>
</taxonomicName>
an obtuse-angled trigone, a low and recurved paracone, and a trigone basin divided by a ridge connecting stylocone and paracone. These characters are diagnostic of Holotheria, a clade that include the last common ancestor to 
<taxonomicName id="4C614D2CFF9BFFC67599A40AFEB2F9FD" authorityName="Kermack, Kermack &amp; Mussett" authorityYear="1968" box="[163,345,1586,1612]" class="Mammalia" family="Kuehneotheriidae" genus="Kuehneotherium" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC67599A40AFEB2F9FD" box="[163,345,1586,1612]" italics="true" pageId="45" pageNumber="726">Kuehneotherium</emphasis>
</taxonomicName>
and Theria, and all of its descendants (
<bibRefCitation id="EFF04B5EFF9BFFC675B4A46BFEC5F9DD" author="Hopson" box="[142,302,1618,1645]" firstAuthor="Hopson" pageId="45" pageNumber="726" pagination="109 - 219" refId="ref41437" refString="HOPSON J. A. 1994. - Synapsid Evolution and the Radiation of Non-Eutherian Mammals, in SPENCER R. S. (ed.), Major Features of Vertebrate Evolution. Short Courses in Paleontology, Cambridge University Press, Cambridge: 109 - 219." type="book chapter" year="1994">Hopson 1994</bibRefCitation>
; 
<bibRefCitation id="EFF04B5EFF9BFFC67405A46AFDE1F9DC" author="WIBLE J. R. &amp; ROUGIER G. W. &amp; NOVACEK M. J. &amp; MCKENNA M. C. &amp; DASHZEVEG C." box="[319,522,1618,1645]" journalOrPublisher="American Museum Novitates" pageId="45" pageNumber="726" pagination="1 - 20" part="3149" publicationUrl="http://hdl.handle.net/2246/3672" refId="ref50741" refString="WIBLE J. R., ROUGIER G. W., NOVACEK M. J., MCKENNA M. C. &amp; DASHZEVEG C. 1995. - A mammalian petrosal from the Early Cretaceous of Mongolia: implications for the evolution of the ear region and mammaliamorph interrelationships. American Museum Novitates 3149: 1 - 20. http: // hdl. handle. net / 2246 / 3672" title="A mammalian petrosal from the Early Cretaceous of Mongolia: implications for the evolution of the ear region and mammaliamorph interrelationships" type="journal article" year="1995">
Wible 
<emphasis id="B915EABDFF9BFFC674B7A46BFE2DF9DD" box="[397,454,1618,1644]" italics="true" pageId="45" pageNumber="726">et al.</emphasis>
1995
</bibRefCitation>
). 
<bibRefCitation id="EFF04B5EFF9BFFC6771EA46AFD27F9DC" author="Martin" box="[548,716,1618,1645]" firstAuthor="Martin" pageId="45" pageNumber="726" pagination="322 - 348" refId="ref43507" refString="MARTIN T. 2002. - New stem lineage representatives of Zatheria (Mammalia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 22: 322 - 348. https: // doi. org / cb 2 w 55" type="journal article" year="2002">Martin (2002)</bibRefCitation>
thus reinterpreted the teeth of 
<taxonomicName id="4C614D2CFF9BFFC674FEA44AFDA3F93D" authorityName="Sigogneau-Russell" authorityYear="1989" box="[452,584,1650,1676]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC674FEA44AFDA3F93D" box="[452,584,1650,1676]" italics="true" pageId="45" pageNumber="726">Thereuodon</emphasis>
</taxonomicName>
as holotherian, probably zatherian upper deciduous premolars. Moreover, 
<bibRefCitation id="EFF04B5EFF9BFFC6758FA48BFE69F97C" author="Bonaparte" box="[181,386,1714,1741]" firstAuthor="Bonaparte" pageId="45" pageNumber="726" pagination="63 - 83" refId="ref36548" refString="BONAPARTE J. F. 1990. - New Late Cretaceous mammals from the Los Alamitos Formation, northern Patagonia. National Geographic Research 6 (1): 63 - 83." type="journal article" year="1990">Bonaparte (1990)</bibRefCitation>
and Sigogneau-Russell &amp; Ensom (1998) observed a gross resemblance between the teeth of 
<taxonomicName id="4C614D2CFF9BFFC6759EA4CAFEC8F8BD" authorityName="Sigogneau-Russell" authorityYear="1989" box="[164,291,1778,1804]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC6759EA4CAFEC8F8BD" box="[164,291,1778,1804]" italics="true" pageId="45" pageNumber="726">Thereuodon</emphasis>
</taxonomicName>
and the upper cheek teeth of 
<taxonomicName id="4C614D2CFF9BFFC677B2A4CAFCEBF8BD" authorityName="Bonaparte" authorityYear="1990" box="[648,768,1778,1804]" class="Mammalia" family="Barbereniidae" genus="Barberenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC677B2A4CAFCEBF8BD" box="[648,768,1778,1804]" italics="true" pageId="45" pageNumber="726">Barberenia</emphasis>
</taxonomicName>
, from the Late Cretaceous of 
<collectingCountry id="F376763FFF9BFFC674EAA52AFDADF89D" box="[464,582,1810,1836]" name="Argentina" pageId="45" pageNumber="726">Argentina</collectingCountry>
. These were subsequently identified by 
<bibRefCitation id="EFF04B5EFF9BFFC674BDA509FDC0F8FD" author="Martin" box="[391,555,1841,1868]" firstAuthor="Martin" pageId="45" pageNumber="726" pagination="1 - 119" refId="ref43478" refString="MARTIN T. 1999. - Dryolestidae (Dryolestoidea, Mammalia) aus dem Oberen Jura-von Portugal. Abhandlungen der senckenbergischen naturforschenden Gesellschaft 550: 1 - 119." type="journal article" year="1999">Martin (1999)</bibRefCitation>
as upper deciduous premolars, probably belonging to the “dryolestoidean” 
<taxonomicName id="4C614D2CFF9BFFC675BEA549FF17F83A" box="[132,252,1905,1931]" class="Mammalia" family="Brandoniidae" genus="Brandonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC675BEA549FF17F83A" box="[132,252,1905,1931]" italics="true" pageId="45" pageNumber="726">Brandonia</emphasis>
</taxonomicName>
from the same formation, an opinion then followed by 
<bibRefCitation id="EFF04B5EFF9BFFC67434A5AAFE36F81D" author="Bonaparte" box="[270,477,1937,1964]" firstAuthor="Bonaparte" pageId="45" pageNumber="726" pagination="339 - 371" refId="ref36581" refString="BONAPARTE J. F. 2002. - New Dryolestida (Theria) from the Late Cretaceous of Los Alamitos, Argentina, and paleogeographical comments. Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 224: 339 - 371. https: // doi. org / 10.1127 / njgpa / 224 / 2002 / 339" type="journal article" year="2002">Bonaparte (2002)</bibRefCitation>
. Thus, it seems that 
<taxonomicName id="4C614D2CFF9BFFC677FBA5A9FF2CF87A" authorityName="Sigogneau-Russell" authorityYear="1989" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC677FBA5A9FF2CF87A" italics="true" pageId="45" pageNumber="726">Thereuodon</emphasis>
</taxonomicName>
should be considered as a stem-Zatheria (
<bibRefCitation id="EFF04B5EFF9BFFC6778CA589FF55F85A" author="Martin" firstAuthor="Martin" pageId="45" pageNumber="726" pagination="322 - 348" refId="ref43507" refString="MARTIN T. 2002. - New stem lineage representatives of Zatheria (Mammalia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 22: 322 - 348. https: // doi. org / cb 2 w 55" type="journal article" year="2002">Martin 2002</bibRefCitation>
). However, from a nomenclatural point of view, the fact that 
<taxonomicName id="4C614D2CFF9BFFC676A3A6CBFBF2FABC" authorityName="Sigogneau-Russell" authorityYear="1989" box="[921,1049,1267,1293]" class="Mammalia" family="Woutersiidae" genus="Thereuodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="726" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF9BFFC676A3A6CBFBF2FABC" box="[921,1049,1267,1293]" italics="true" pageId="45" pageNumber="726">Thereuodon</emphasis>
</taxonomicName>
is known only by milk teeth does not invalidate the genus, which is why we use it here to precise the identification of this specimen.
</paragraph>
</subSubSection>
</treatment>
</document>