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<document id="FA74AB998BD9E53175C50AAA847FDE79" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit &amp; Katoh, Toru" docDate="2020" docId="03C86C16EB30574CFF6EDF2DFEC3FB9D" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) tanabai Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh 2020" docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="23" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="76C016A69263197F87DCE2D711E52D86">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="8556C9F6347F5975EA0A02DD4F1D28B0">Toda, Masanori J.</mods:namePart>
<mods:affiliation id="BE8BEEC98101EE1A482C326101D77559">Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="677670CBFA3B82DA083C18C243A14916">Bänziger, Hans</mods:namePart>
<mods:affiliation id="511BE2854228B2D6D791070AFC952C2A">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="53FD4EA0481B3256A7F1971467F28FCC">Sati, Pradeep C.</mods:namePart>
<mods:affiliation id="964DE3E64DA7B643E873B1FDCC488C60">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="4297E522E2147B1480A34BEA05D82E10">Fartyal, Rajendra S.</mods:namePart>
<mods:affiliation id="11C2064F38CA5DD8C752F7722EE0CF3D">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="5293D05A1EFEBA36D37FC6CA6D452311">Suwito, Awit</mods:namePart>
<mods:affiliation id="17375463D0EA13A3F9120D8C8FDEA988">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<mods:namePart id="929622B064DB42EB124D639650B1F019">Katoh, Toru</mods:namePart>
<mods:affiliation id="A4158A14CC6D103514F1C66979FCCDD5">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
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<mods:date id="8660338B7AE178D09F67801A2FDCFC97">2020</mods:date>
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<treatment id="03C86C16EB30574CFF6EDF2DFEC3FB9D" ID-DOI="http://doi.org/10.5281/zenodo.5221304" ID-GBIF-Taxon="164612792" ID-Zenodo-Dep="5221304" LSID="urn:lsid:plazi:treatment:03C86C16EB30574CFF6EDF2DFEC3FB9D" httpUri="http://treatment.plazi.org/id/03C86C16EB30574CFF6EDF2DFEC3FB9D" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="C37B8E8BEB30574CFF6EDF2DFCC1FD95" box="[151,810,549,576]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BDEDD00EB30574CFF6EDF2DFCC1FD95" blockId="22.[151,810,549,576]" box="[151,810,549,576]" pageId="22" pageNumber="23">
<heading id="D0966A6CEB30574CFF6EDF2DFCC1FD95" bold="true" box="[151,810,549,576]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B9150112EB30574CFF6EDF2DFCC1FD95" bold="true" box="[151,810,549,576]" pageId="22" pageNumber="23">
2.2.
<taxonomicName id="4C61A683EB30574CFF32DF2DFDE6FDEA" authorityName="Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh" authorityYear="2020" box="[203,525,549,576]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tanabai" subGenus="Ashima">
<emphasis id="B9150112EB30574CFF32DF2DFED8FDEA" bold="true" box="[203,307,549,576]" italics="true" pageId="22" pageNumber="23">Phortica</emphasis>
(
<emphasis id="B9150112EB30574CFEBDDF2DFE4AFDEA" bold="true" box="[324,417,549,576]" italics="true" pageId="22" pageNumber="23">Ashima</emphasis>
)
<emphasis id="B9150112EB30574CFE4BDF2DFDE6FDEA" bold="true" box="[434,525,549,576]" italics="true" pageId="22" pageNumber="23">tanabai</emphasis>
</taxonomicName>
species subgroup, new
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37B8E8BEB30574CFF6EDF64FEC3FB9D" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BDEDD00EB30574CFF6EDF64FBCFFD01" blockId="22.[151,1437,620,1079]" pageId="22" pageNumber="23">
<emphasis id="B9150112EB30574CFF6EDF64FEF9FD2D" bold="true" box="[151,274,620,647]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
Prementum with a pair of patches of pubescence (ch.12-1;
<figureCitation id="135AC185EB30574CFC47DF65FBFCFD22" box="[958,1047,621,648]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Fig. 6B</figureCitation>
); aedeagal, outer membrane ornamented with somewhat long, conical spines (ch.55-0, ch.56-2;
<figureCitation id="135AC185EB30574CFCAFDF99FC21FD01" box="[854,970,657,683]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6E,G</figureCitation>
,
<figureCitation id="135AC185EB30574CFC2FDF99FBFCFD01" box="[982,1047,657,683]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7G,H</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB30574CFF3EDFBCFD17FC61" blockId="22.[151,1437,620,1079]" pageId="22" pageNumber="23">
<emphasis id="B9150112EB30574CFF3EDFBCFE41FD65" bold="true" box="[199,426,692,719]" pageId="22" pageNumber="23">Shared characters.</emphasis>
Some interfrontal setulae as long as lowest setulae on ocellar triangle (ch.3-0;
<figureCitation id="135AC185EB30574CFACFDFBDFA7BFD65" box="[1334,1424,693,719]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">Fig. 7B</figureCitation>
). Arista apically with leaf-like expansion (ch.7-1;
<figureCitation id="135AC185EB30574CFD33DFD1FCC2FD59" box="[714,809,729,755]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6A</figureCitation>
,
<figureCitation id="135AC185EB30574CFCC1DFD1FCB7FD59" box="[824,860,729,755]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7C</figureCitation>
). Setulae just before prescutellar acrostichal setae somewhat longer than other acrostichal setulae (ch.15-1). Fore and mid tibiae entirely yellow (ch.17,18-0). Abdominal sternites III–V wider than long (ch.22,24,25-1;
<figureCitation id="135AC185EB30574CFD29DE29FCC1FC91" box="[720,810,801,827]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6C</figureCitation>
,
<figureCitation id="135AC185EB30574CFCCCDE29FCB0FC91" box="[821,859,801,827]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7D</figureCitation>
); III widest among sternites (ch.23-1;
<figureCitation id="135AC185EB30574CFAFDDE29FAB5FC91" box="[1284,1374,801,827]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6C</figureCitation>
,
<figureCitation id="135AC185EB30574CFA93DE29FA64FC91" box="[1386,1423,801,827]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7D</figureCitation>
), with dense setulae on medial surface (ch.26-1;
<figureCitation id="135AC185EB30574CFD48DE4DFCE7FCF5" box="[689,780,837,863]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6C</figureCitation>
,
<figureCitation id="135AC185EB30574CFCE0DE4DFCD4FCF5" box="[793,831,837,863]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7D</figureCitation>
). Pregonite not concave on dorsosubapical margin of apical knob (ch.48-0;
<figureCitation id="135AC185EB30574CFE69DE61FE01FC29" box="[400,490,873,899]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6E</figureCitation>
,
<figureCitation id="135AC185EB30574CFE0EDE61FDF6FC29" box="[503,541,873,899]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7G</figureCitation>
), without basal extension (ch.49-0;
<figureCitation id="135AC185EB30574CFC4ADE61FBE6FC29" box="[947,1037,873,899]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6E</figureCitation>
,
<figureCitation id="135AC185EB30574CFBE3DE61FBABFC29" box="[1050,1088,873,899]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7G</figureCitation>
). Posterior, rod-like process of phallapodeme not longer than aedeagal median rod (ch.62-0;
<figureCitation id="135AC185EB30574CFCC7DE85FC78FC02" box="[830,915,909,936]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Fig. 6E</figureCitation>
). Aedeagal sheath without lateral lobes bearing strongly sclerotized postgonites (ch.65-0;
<figureCitation id="135AC185EB30574CFD97DEB9FD28FC61" box="[622,707,945,971]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="22" pageNumber="23">Figs 6F</figureCitation>
,
<figureCitation id="135AC185EB30574CFD37DEB9FD04FC61" box="[718,751,945,971]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="22" pageNumber="23">7F</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB30574CFF3EDEDDFEC3FB9D" blockId="22.[151,1437,620,1079]" pageId="22" pageNumber="23">
<emphasis id="B9150112EB30574CFF3EDEDDFE7AFC45" bold="true" box="[199,401,980,1007]" pageId="22" pageNumber="23">Included species.</emphasis>
<taxonomicName id="4C61A683EB30574CFE61DEDDFC99FC45" authority="Cheng &amp; Chen" authorityName="Cheng &amp; Chen" authorityYear="2008" box="[408,882,981,1007]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="huiluoi" subGenus="Ashima">
<emphasis id="B9150112EB30574CFE61DEDDFE11FC45" box="[408,506,981,1007]" italics="true" pageId="22" pageNumber="23">Phortica</emphasis>
(
<emphasis id="B9150112EB30574CFDF3DEDDFDB5FC45" box="[522,606,981,1007]" italics="true" pageId="22" pageNumber="23">Ashima</emphasis>
)
<emphasis id="B9150112EB30574CFD97DEDDFD56FC45" box="[622,701,981,1007]" italics="true" pageId="22" pageNumber="23">huiluoi</emphasis>
Cheng &amp; Chen
</taxonomicName>
*;
<emphasis id="B9150112EB30574CFC71DEDEFC72FC45" box="[904,921,982,1007]" italics="true" pageId="22" pageNumber="23">P</emphasis>
.
<taxonomicName id="4C61A683EB30574CFC5EDEDDFBBAFC45" authorityName="Chen in Chen &amp; Maca" authorityYear="2012" box="[935,1105,981,1007]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mediarista" subGenus="Ashima">
(
<emphasis id="B9150112EB30574CFC48DEDDFC29FC45" box="[945,962,981,1007]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.)
<emphasis id="B9150112EB30574CFC20DEDDFBBAFC45" box="[985,1105,981,1007]" italics="true" pageId="22" pageNumber="23">mediarista</emphasis>
</taxonomicName>
;
<emphasis id="B9150112EB30574CFBA6DEDEFB9BFC45" box="[1119,1136,982,1007]" italics="true" pageId="22" pageNumber="23">P</emphasis>
.
<taxonomicName id="4C61A683EB30574CFB87DEDDFAEFFC45" box="[1150,1284,981,1007]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tanabei" subGenus="Ashima">
(
<emphasis id="B9150112EB30574CFB7EDEDDFB73FC45" box="[1159,1176,981,1007]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.)
<emphasis id="B9150112EB30574CFB49DEDDFAEFFC45" box="[1200,1284,981,1007]" italics="true" pageId="22" pageNumber="23">tanabei</emphasis>
</taxonomicName>
;
<emphasis id="B9150112EB30574CFAEBDEDEFAC8FC45" box="[1298,1315,982,1007]" italics="true" pageId="22" pageNumber="23">P</emphasis>
.
<taxonomicName id="4C61A683EB30574CFAC8DEDDFF53FBB9" authority="Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh, 2020" authorityName="Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh" authorityYear="2020" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="akutsui" status="sp. nov." subGenus="Ashima">
(
<emphasis id="B9150112EB30574CFAC3DEDDFAA0FC45" box="[1338,1355,981,1007]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.)
<emphasis id="B9150112EB30574CFA9ADEDDFF53FBB9" italics="true" pageId="22" pageNumber="23">akutsui</emphasis>
</taxonomicName>
<emphasis id="B9150112EB30574CFF47DEF1FEFCFBB9" bold="true" box="[190,279,1017,1043]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A226BC69EB30574CFF47DEF1FEFCFBB9" box="[190,279,1017,1043]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; and
<emphasis id="B9150112EB30574CFEADDEF2FE8EFBB9" box="[340,357,1018,1043]" italics="true" pageId="22" pageNumber="23">P</emphasis>
.
<taxonomicName id="4C61A683EB30574CFE8BDEF1FDDCFBB9" authority="Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh, 2020" authorityName="Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh" authorityYear="2020" box="[370,567,1017,1043]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="halimunensis" status="sp. nov." subGenus="Ashima">
(
<emphasis id="B9150112EB30574CFE85DEF1FE66FBB9" box="[380,397,1017,1043]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.)
<emphasis id="B9150112EB30574CFE5ADEF1FDDCFBB9" box="[419,567,1017,1043]" italics="true" pageId="22" pageNumber="23">halimunensis</emphasis>
</taxonomicName>
<emphasis id="B9150112EB30574CFDC7DEF1FD7CFBB9" bold="true" box="[574,663,1017,1043]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A226BC69EB30574CFDC7DEF1FD7CFBB9" box="[574,663,1017,1043]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
* The assignment was judged from the original description by
<bibRefCitation id="EFF0A0F1EB30574CFAAADEF1FECFFB9D" author="Cheng" etAl="et al." firstAuthor="Cheng" pageId="22" pageNumber="23" pagination="614 - 632" refId="ref34455" refString="Cheng, Y., Gao, J. J., Watabe, H. &amp; Chen, H. W. (2008) Revision of the genus Phortica Schiner 1862 in China (Diptera: Drosophilidae). Zoological Studies, 47, 614 - 632." type="journal article" year="2008">
Cheng
<emphasis id="B9150112EB30574CFF6ED916FF23FB9D" box="[151,200,1053,1079]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (2008)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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