201 lines
21 KiB
XML
201 lines
21 KiB
XML
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<mods:title id="6725ABB1065FD43546209609827667B7">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="78D336BEC8C872299537254385B0A854">Toda, Masanori J.</mods:namePart>
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<mods:affiliation id="154C7282E6A133572B0964A0E207BB91">Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="13CCFB665C3F28EF726BC4BD870F1258">Bänziger, Hans</mods:namePart>
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<mods:affiliation id="6DE58CD5B6B64FFB3309AE888CD64F82">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="850115933B0427783C31DCB01A19C6AE">Sati, Pradeep C.</mods:namePart>
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<mods:affiliation id="F04A99AA5F5ADDD0FB65D51AB0DFC338">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="20494BD2635444B552935EC167A9C729">Fartyal, Rajendra S.</mods:namePart>
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<mods:affiliation id="9A3BFF94E3D0A76866036E4CEBC65B58">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="EC446A4217407EA05039569EB919D8F3">Suwito, Awit</mods:namePart>
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<mods:affiliation id="F56FA444426E6BA29A9DB1E4849EB92F">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<mods:namePart id="4B6E6938875F7A53F6E8D4C4DB7B5485">Katoh, Toru</mods:namePart>
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<mods:affiliation id="90AB252DF74917395D0853594149DD92">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
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<mods:title id="F19E47C56EB604507871DD31382C1D4F">Zootaxa</mods:title>
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<mods:number id="7493A21D4115D904969FDC20C0740161">4789</mods:number>
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<treatment id="03C86C16EB285755FF6EDB69FE80FEE9" ID-DOI="http://doi.org/10.5281/zenodo.5221280" ID-GBIF-Taxon="164612806" ID-Zenodo-Dep="5221280" LSID="urn:lsid:plazi:treatment:03C86C16EB285755FF6EDB69FE80FEE9" httpUri="http://treatment.plazi.org/id/03C86C16EB285755FF6EDB69FE80FEE9" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<emphasis id="B9150112EB285754FF6EDB69FC80F9D1" bold="true" box="[151,875,1633,1660]" pageId="14" pageNumber="15">
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6)
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<taxonomicName id="4C61A683EB285754FF4EDB69FC80F9D1" authority="(Tsacas & Okada, 1983)" baseAuthorityName="Tsacas & Okada" baseAuthorityYear="1983" box="[183,875,1633,1660]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phyllochaeta" subGenus="Ashima">
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<emphasis id="B9150112EB285754FF4EDB69FEF4F9D6" bold="true" box="[183,287,1633,1660]" italics="true" pageId="14" pageNumber="15">Phortica</emphasis>
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(
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<emphasis id="B9150112EB285754FEC9DB69FE66F9D6" bold="true" box="[304,397,1633,1660]" italics="true" pageId="14" pageNumber="15">Ashima</emphasis>
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)
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<emphasis id="B9150112EB285754FE67DB69FDD2F9D6" bold="true" box="[414,569,1633,1660]" italics="true" pageId="14" pageNumber="15">phyllochaeta</emphasis>
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(
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<bibRefCitation id="EFF0A0F1EB285754FDB1DB69FC88F9D1" author="Tsacas & Okada" box="[584,867,1633,1659]" firstAuthor="Tsacas" pageId="14" pageNumber="15" pagination="228 - 237" refId="ref37303" refString="Tsacas, L. & Okada, T. (1983) On the Oriental and New Guinean species of the genus Amiota Loew originally described by Duda as Phortica foliiseta, with descriptions of three new species (Diptera, Drosophilidae). Kontyu, 51, 228 - 237." type="journal article" year="1983">Tsacas & Okada, 1983</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDEDD00EB285754FF6EDBAEFD14F914" blockId="14.[151,767,1701,1757]" box="[151,767,1701,1726]" pageId="14" pageNumber="15">
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<treatmentCitation id="0AC0FB11EB285754FF6EDBAEFD10F914" author="Tsacas, L. & Okada, T." box="[151,763,1701,1726]" page="231" pageId="14" pageNumber="15" year="1983">
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<taxonomicName id="4C61A683EB285754FF6EDBAEFD10F914" ID-CoL="CSK8" authority="Tsacas & Okada, 1983: 231" authorityName="Tsacas & Okada" authorityPageNumber="231" authorityYear="1983" box="[151,763,1701,1726]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phyllochaeta" subGenus="Phortica">
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<emphasis id="B9150112EB285754FF6EDBAEFF0BF917" box="[151,224,1702,1725]" italics="true" pageId="14" pageNumber="15">Amiota</emphasis>
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(
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<emphasis id="B9150112EB285754FF16DBAEFEA3F914" box="[239,328,1702,1726]" italics="true" pageId="14" pageNumber="15">Phortica</emphasis>
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)
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<emphasis id="B9150112EB285754FEAEDBAEFE31F914" box="[343,474,1702,1726]" italics="true" pageId="14" pageNumber="15">phyllochaeta</emphasis>
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<bibRefCitation id="EFF0A0F1EB285754FE19DBAEFD10F914" author="Tsacas & Okada" box="[480,763,1701,1726]" firstAuthor="Tsacas" pageId="14" pageNumber="15" pagination="228 - 237" refId="ref37303" refString="Tsacas, L. & Okada, T. (1983) On the Oriental and New Guinean species of the genus Amiota Loew originally described by Duda as Phortica foliiseta, with descriptions of three new species (Diptera, Drosophilidae). Kontyu, 51, 228 - 237." type="journal article" year="1983">Tsacas & Okada, 1983: 231</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BDEDD00EB285754FF6EDBCCFD19F977" blockId="14.[151,767,1701,1757]" box="[151,754,1732,1757]" pageId="14" pageNumber="15">
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<treatmentCitationGroup id="AB71FA2EEB285754FF6EDBCCFD19F977" box="[151,754,1732,1757]" pageId="14" pageNumber="15">
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<treatmentCitation id="0AC0FB11EB285754FF6EDBCCFD06F977" author="Chen, H. W. & Maca, J." box="[151,749,1732,1757]" page="507" pageId="14" pageNumber="15" year="2012">
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<taxonomicName id="4C61A683EB285754FF6EDBCCFD06F977" authority=": Chen & Maca, 2012: 507" authorityName=": Chen & Maca" authorityPageNumber="507" authorityYear="2012" box="[151,749,1732,1757]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phyllochaeta" subGenus="Ashima">
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<emphasis id="B9150112EB285754FF6EDBCCFF1BF976" box="[151,240,1732,1756]" italics="true" pageId="14" pageNumber="15">Phortica</emphasis>
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(
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<emphasis id="B9150112EB285754FF06DBCCFEA0F976" box="[255,331,1732,1756]" italics="true" pageId="14" pageNumber="15">Ashima</emphasis>
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)
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<emphasis id="B9150112EB285754FEA0DBCCFE37F976" box="[345,476,1732,1756]" italics="true" pageId="14" pageNumber="15">phyllochaeta</emphasis>
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:
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<bibRefCitation id="EFF0A0F1EB285754FE13DBCCFD06F977" author="Chen & Maca" box="[490,749,1732,1757]" firstAuthor="Chen" pageId="14" pageNumber="15" pagination="493 - 509" refId="ref33941" refString="Chen, H. W. & Maca, J. (2012) Ten new species of the genus Phortica from the Afrotropical and Oriental regions (Diptera: Drosophilidae). Zootaxa, 3478 (1), 493 - 509. https: // doi. org / 10.11646 / zootaxa. 3478.1.42" type="journal article" year="2012">Chen & Máca, 2012: 507</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B8E8BEB285754FF6EDA02FE94F857" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="8BDEDD00EB285754FF6EDA02FDB7F8E3" blockId="14.[151,1437,1802,2045]" pageId="14" pageNumber="15">
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<emphasis id="B9150112EB285754FF6EDA02FEF9F88F" bold="true" box="[151,274,1802,1829]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
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Aedeagal sheath with two sclerotized, marginally serrated postgonites on anterior portion of left lateral lobe (“
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<figureCitation id="135AC185EB285754FF1BDA27FECCF8E3" box="[226,295,1839,1865]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
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” in
|
||
<bibRefCitation id="EFF0A0F1EB285754FEACDA27FDA4F8E3" author="Tsacas & Okada" box="[341,591,1839,1865]" firstAuthor="Tsacas" pageId="14" pageNumber="15" pagination="228 - 237" refId="ref37303" refString="Tsacas, L. & Okada, T. (1983) On the Oriental and New Guinean species of the genus Amiota Loew originally described by Duda as Phortica foliiseta, with descriptions of three new species (Diptera, Drosophilidae). Kontyu, 51, 228 - 237." type="journal article" year="1983">Tsacas & Okada 1983</bibRefCitation>
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).
|
||
</paragraph>
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<paragraph id="8BDEDD00EB285754FF3EDA5AFC26F81F" blockId="14.[151,1437,1802,2045]" pageId="14" pageNumber="15">
|
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<emphasis id="B9150112EB285754FF3EDA5AFDE5F8C7" bold="true" box="[199,526,1874,1901]" pageId="14" pageNumber="15">Supplementary description</emphasis>
|
||
(not repeating characters common to
|
||
<taxonomicName id="4C61A683EB285754FC3FDA5CFBBBF8C7" authorityName="Chen in Chen & Maca" authorityYear="2012" box="[966,1104,1875,1901]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="biclavata">
|
||
<emphasis id="B9150112EB285754FC3FDA5CFC3CF8C7" box="[966,983,1876,1901]" italics="true" pageId="14" pageNumber="15">P</emphasis>
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||
.
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||
<emphasis id="B9150112EB285754FC11DA5BFBBBF8C7" box="[1000,1104,1875,1901]" italics="true" pageId="14" pageNumber="15">biclavata</emphasis>
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</taxonomicName>
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||
). Supracervical setae 12–14. Dorsomedial, tentorial apodeme 2/5 as long as basal, parallel portion of dorsolateral, tentorial apodeme. Cibarial, medial sensilla 9–11 per side; posterior sensilla approximately 6 per side.
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||
</paragraph>
|
||
<paragraph id="8BDEDD00EB285754FF3EDAB6FE94F857" blockId="14.[151,1437,1802,2045]" pageId="14" pageNumber="15">
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<emphasis id="B9150112EB285754FF3EDAB6FE55F873" bold="true" box="[199,446,1982,2009]" pageId="14" pageNumber="15">Specimen examined.</emphasis>
|
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<materialsCitation id="3B09D75DEB285754FE3EDAB7FE97F857" ID-GBIF-Occurrence="2636268371" collectingDate="1977-03-18" collectionCode="T, NSMT" collectorName="Okada" country="Papua New Guinea" elevation="1200" location="Wau" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Morobe" typeStatus="paratype">
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<collectingCountry id="F3769D90EB285754FE3EDAB7FD4CF873" box="[455,679,1983,2009]" name="Papua New Guinea" pageId="14" pageNumber="15">Papua New Guinea</collectingCountry>
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||
:
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<specimenCount id="9D671689EB285754FD4DDAB7FD3CF873" box="[692,727,1983,2009]" pageId="14" pageNumber="15" type="male">1♂</specimenCount>
|
||
<typeStatus id="54DA63A2EB285754FD19DAB7FCA8F873" box="[736,835,1983,2009]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
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,
|
||
<location id="8EBE8BDBEB285754FCA9DAB7FC6DF873" LSID="urn:lsid:plazi:treatment:03C86C16EB285755FF6EDB69FE80FEE9:8EBE8BDBEB285754FCA9DAB7FC6DF873" box="[848,902,1983,2009]" country="Papua New Guinea" name="Wau" pageId="14" pageNumber="15" stateProvince="Morobe">Wau</location>
|
||
,
|
||
<collectingRegion id="49A513E2EB285754FC6ADAB7FBAEF873" box="[915,1093,1983,2009]" country="Papua New Guinea" name="Morobe" pageId="14" pageNumber="15">Morobe district</collectingRegion>
|
||
,
|
||
<elevation id="004C3A33EB285754FBABDAB7FB04F873" box="[1106,1263,1983,2009]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="15" unit="m" value="1200.0">
|
||
<quantity id="4C9970E5EB285754FBABDAB7FB5BF873" box="[1106,1200,1983,2009]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="15" unit="m" value="1200.0">1,200 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFDFFBC0EB285754FB04DAB7FA92F873" box="[1277,1401,1983,2009]" pageId="14" pageNumber="15" value="1977-03-18">
|
||
<collectingDate id="EF9B0228EB285754FB04DAB7FA92F873" box="[1277,1401,1983,2009]" pageId="14" pageNumber="15" value="1977-03-18">18.iii.1977</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED7045C5EB285754FA7FDAB7FA73F873" box="[1414,1432,1983,2009]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="15">T</collectionCode>
|
||
.
|
||
<collectorName id="2694B8D6EB285754FF6EDAEBFF0BF857" box="[151,224,2019,2045]" pageId="14" pageNumber="15">Okada</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED7045C5EB285754FEE6DAEBFE98F857" box="[287,371,2019,2045]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="14" pageNumber="15">NSMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB295755FF3EDD90FDACFF19" box="[199,583,152,179]" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="8BDEDD00EB295755FF3EDD90FDACFF19" blockId="15.[151,1437,152,323]" box="[199,583,152,179]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9150112EB295755FF3EDD90FE8AFF19" bold="true" box="[199,353,152,179]" pageId="15" pageNumber="16">Distribution.</emphasis>
|
||
<collectingCountry id="F3769D90EB295755FE9EDD91FDAFFF19" box="[359,580,153,179]" name="Papua New Guinea" pageId="15" pageNumber="16">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB295755FF3EDDB5FE80FEE9" pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph id="8BDEDD00EB295755FF3EDDB5FE80FEE9" blockId="15.[151,1437,152,323]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9150112EB295755FF3EDDB5FED0FF7D" bold="true" box="[199,315,189,215]" pageId="15" pageNumber="16">Remarks.</emphasis>
|
||
This species closely resembles the foregoing species,
|
||
<taxonomicName id="4C61A683EB295755FC56DDB6FBD3FF7D" authorityName="Chen in Chen & Maca" authorityYear="2012" box="[943,1080,189,215]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="biclavata">
|
||
<emphasis id="B9150112EB295755FC56DDB6FC2BFF7D" box="[943,960,190,215]" italics="true" pageId="15" pageNumber="16">P</emphasis>
|
||
.
|
||
<emphasis id="B9150112EB295755FC29DDB5FBD3FF7D" box="[976,1080,189,215]" italics="true" pageId="15" pageNumber="16">biclavata</emphasis>
|
||
</taxonomicName>
|
||
, in having the aedeagal sheath with two marginally serrated postgonites on left lateral lobe and one anterior, large, marginally serrated postgonite and posterior, small, not serrated one on right lobe. However, these two species can be distinguished from each other by their diagnoses.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |