treatments-xml/data/03/A7/E8/03A7E804897BDB57FF2D7783FAA1A193.xml
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<document id="C7ED04C66D3C7A7DABA0DC2182CECB3B" ID-CLB-Dataset="30796" ID-DOI="10.11646/zootaxa.4378.2.3" ID-GBIF-Dataset="db1a48bf-e5ec-4523-93bf-24af1ebb4a72" ID-ISSN="1175-5326" ID-Zenodo-Dep="1169497" ID-ZooBank="FD162D89-A6D6-44FA-AB5E-9DA724F7B62A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518134184692" checkinUser="plazi" docAuthor="Travouillon, Kenny J. &amp; Phillips, Matthew J." docDate="2018" docId="03A7E804897BDB57FF2D7783FAA1A193" docLanguage="en" docName="zootaxa.4378.2.3.pdf" docOrigin="Zootaxa 4378 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perameles papillon Travouillon &amp; Phillips 2018, sp. nov." docType="treatment" docUuid="453893E8-55DC-481A-B095-EAF67B434B8A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="241" masterDocId="FF9E907C8970DB46FFBA7321FFF1A27F" masterDocTitle="Total evidence analysis of the phylogenetic relationships of bandicoots and bilbies (Marsupialia: Peramelemorphia): reassessment of two species and description of a new species" masterLastPageNumber="256" masterPageNumber="224" pageNumber="235" updateTime="1698520503571" updateUser="ExternalLinkService">
<mods:mods id="54F945D66F55BB143076D5AAFFE853CB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2C6C8529AE17AC3F4C8974BBD5B87FAF">
<mods:title id="810BBE53DE420F69AE234CE893877C65">Total evidence analysis of the phylogenetic relationships of bandicoots and bilbies (Marsupialia: Peramelemorphia): reassessment of two species and description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="7157963353B617B86D4505540D6ABEA6" type="personal">
<mods:role id="0E051ACD231C0E61E8BECD8AB5BD6FD7">
<mods:roleTerm id="B4544DCBC7253E5036162A57D7D1E415">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF4B5A1EF31C88A0782EA5134731B2AB">Travouillon, Kenny J.</mods:namePart>
</mods:name>
<mods:name id="EE7977D6D4D02F64F579C43EC10A70BC" type="personal">
<mods:role id="55750E15976C01A1F2211F821DE1ACC6">
<mods:roleTerm id="603DD24B224521A0A9159B69E47DB705">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA8B214B6507ADDC3AB075BB680A50BD">Phillips, Matthew J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CE844E850791715CD9D96AAD14DB8D8F">text</mods:typeOfResource>
<mods:relatedItem id="457026CE32BDCA46580A67F0D5AE072E" type="host">
<mods:titleInfo id="7BD9328DACDCF4440704DC573C39AFCE">
<mods:title id="2D2F81D070566955EB5A80DDF8D0A185">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4156F97D906E6091D7E2EC660111BC3B">
<mods:date id="31C3822F02E9A4D86CC0FC9028E5672A">2018</mods:date>
<mods:detail id="1FE7C4EC44FDAFE99116E0D63691078F" type="pubDate">
<mods:number id="25DA1289A2470F2B4EB0DB834ABCFC1A">2018-02-08</mods:number>
</mods:detail>
<mods:detail id="D853EF99D537C50A2D632ACF72CCB9DF" type="volume">
<mods:number id="D00D7EF432855DB0B49B8F1F048DD338">4378</mods:number>
</mods:detail>
<mods:detail id="6410D39B14AEFAB1E47DC3D36FB6A001" type="issue">
<mods:number id="1C2A29A6C84389F632265A288CB23665">2</mods:number>
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<mods:start id="CFDF6109B48F2FC9041847B59A612227">224</mods:start>
<mods:end id="9CCAD5336B34218DC2850C9663198CA6">256</mods:end>
</mods:extent>
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<mods:classification id="510FF775434C0B70C7C061BB9CBCC774">journal article</mods:classification>
<mods:identifier id="A9C04DD579AB8FD9955A90AE14BD75D7" type="CLB-Dataset">30796</mods:identifier>
<mods:identifier id="3443AF46500B373B1AB41BB016404E11" type="DOI">10.11646/zootaxa.4378.2.3</mods:identifier>
<mods:identifier id="412EA92DB40C2E8C180A2FB152306947" type="GBIF-Dataset">db1a48bf-e5ec-4523-93bf-24af1ebb4a72</mods:identifier>
<mods:identifier id="AD12C39C0E2751502B608AED6AC083B8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3E426AC2429549B1ACD18FF583813508" type="Zenodo-Dep">1169497</mods:identifier>
<mods:identifier id="F4837652A3E794F1647857C7C2F28D28" type="ZooBank">FD162D89-A6D6-44FA-AB5E-9DA724F7B62A</mods:identifier>
</mods:mods>
<treatment id="03A7E804897BDB57FF2D7783FAA1A193" ID-DOI="http://doi.org/10.5281/zenodo.6485403" ID-GBIF-Taxon="140563207" ID-Zenodo-Dep="6485403" LSID="urn:lsid:zoobank.org:act:453893E8-55DC-481A-B095-EAF67B434B8A" httpUri="http://treatment.plazi.org/id/03A7E804897BDB57FF2D7783FAA1A193" lastPageId="17" lastPageNumber="241" pageId="11" pageNumber="235">
<subSubSection id="C3140A99897BDB4DFF2D7783FEF1A6A1" pageId="11" pageNumber="235" type="nomenclature">
<paragraph id="8BB15912897BDB4DFF2D7783FE15A6C2" blockId="11.[151,484,1186,1246]" box="[151,484,1186,1213]" pageId="11" pageNumber="235">
<heading id="D0F9EE7E897BDB4DFF2D7783FE15A6C2" bold="true" box="[151,484,1186,1213]" fontSize="11" level="1" pageId="11" pageNumber="235" reason="1">
<emphasis id="B97A8500897BDB4DFF2D7783FE15A6C2" bold="true" box="[151,484,1186,1213]" pageId="11" pageNumber="235">
<taxonomicName id="4C0E2291897BDB4DFF2D7783FE8EA6C3" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[151,383,1186,1212]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="11" pageNumber="235" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897BDB4DFF2D7783FE8EA6C3" bold="true" box="[151,383,1186,1212]" italics="true" pageId="11" pageNumber="235">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897BDB4DFE3C7782FE15A6C2" box="[390,484,1187,1213]" pageId="11" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB15912897BDB4DFF2D77E4FEF1A6A1" blockId="11.[151,484,1186,1246]" box="[151,256,1221,1246]" pageId="11" pageNumber="235">
<figureCitation id="13354597897BDB4DFF2D77E4FEF1A6A1" box="[151,256,1221,1246]" captionStart-0="FIGURE 5" captionStart-1="FIGURE6" captionStart-2="FIGURE 7" captionStartId-0="13.[151,250,1959,1981]" captionStartId-1="14.[151,250,1153,1175]" captionStartId-2="16.[151,250,1928,1950]" captionTargetBox-0="[177,1418,193,1949]" captionTargetBox-1="[151,1436,193,1131]" captionTargetBox-2="[151,1436,337,1906]" captionTargetId-0="figure@13.[177,1418,193,1949]" captionTargetId-1="figure@14.[151,1436,193,1131]" captionTargetId-2="figure@16.[151,1436,337,1906]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="16" captionText-0="FIGURE 5. Perameles papillon sp. nov., study skins. AC, holotype WAM M571; DF, paratype WAM M574; GI, paratype WAM M572.A, D, and G, dorsal view; B, E, and H, lateral view; C, F, and I, ventral view. Scale = 5cm." captionText-1="FIGURE6. Perameles papillonsp.nov., skulls.AC and GH, holotype WAMM571; DF and IJ, paratype WAMM577.A, D, G, I, dorsal view; B, E, ventral view; C, F, H, J, lateral view. Scale = 2cm." captionText-2="FIGURE 7. Perameles papillon sp. nov., upper and lower dentition of paratype WAM M577. AB, upper premolas; CD, upper molars; EF, lower premolars; GH, lower molars. A, C, E and G, occlusal view; B, D, F and H, lateral view. Scale = 2mm." httpUri-0="https://zenodo.org/record/1169507/files/figure.png" httpUri-1="https://zenodo.org/record/1169509/files/figure.png" httpUri-2="https://zenodo.org/record/1169511/files/figure.png" pageId="11" pageNumber="235">Figs. 57</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3140A99897BDB4AFF2D762CFDE0A1DB" lastPageId="12" lastPageNumber="236" pageId="11" pageNumber="235" type="materials_examined">
<paragraph id="8BB15912897BDB4DFF2D762CFD5DA759" blockId="11.[151,1437,1293,2038]" box="[151,684,1293,1318]" pageId="11" pageNumber="235">
<emphasis id="B97A8500897BDB4DFF2D762CFEF5A759" bold="true" box="[151,260,1293,1318]" pageId="11" pageNumber="235">Holotype</emphasis>
. WAM M571, skin and skull, female.
</paragraph>
<paragraph id="8BB15912897BDB4DFF7D7610FD4FA735" blockId="11.[151,1437,1293,2038]" box="[199,702,1329,1354]" pageId="11" pageNumber="235">
<materialsCitation id="3B66534F897BDB4DFF7D7610FD4FA735" ID-GBIF-Occurrence="1806521140" box="[199,702,1329,1354]" location="Ooldea" pageId="11" pageNumber="235" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<emphasis id="B97A8500897BDB4DFF7D7610FE94A735" bold="true" box="[199,357,1329,1354]" pageId="11" pageNumber="235">
<typeStatus id="54B5E7B0897BDB4DFF7D7610FEF3A735" box="[199,258,1329,1354]" pageId="11" pageNumber="235">Type</typeStatus>
locality.
</emphasis>
<location id="8ED10FC9897BDB4DFED67610FE31A735" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFED67610FE31A735" box="[364,448,1329,1354]" name="Ooldea" pageId="11" pageNumber="235" stateProvince="South Australia">Ooldea</location>
,
<collectingRegion id="49CA97F0897BDB4DFE717610FD8BA735" box="[459,634,1329,1354]" country="Australia" name="South Australia" pageId="11" pageNumber="235">South Australia</collectingRegion>
(SA).
</materialsCitation>
</paragraph>
<paragraph id="8BB15912897BDB4DFF7D7674FDEBA439" blockId="11.[151,1437,1293,2038]" pageId="11" pageNumber="235">
<materialsCitation id="3B66534F897BDB4DFF7D7674FEDDA781" ID-GBIF-Occurrence="1806521141" collectionCode="WAM" county="Rawlinna" location="Eucla Pass" municipality="Skin" pageId="11" pageNumber="235" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<typeStatus id="54B5E7B0897BDB4DFF7D7674FECEA711" box="[199,319,1365,1390]" pageId="11" pageNumber="235" type="paratype">
<emphasis id="B97A8500897BDB4DFF7D7674FECEA711" bold="true" box="[199,319,1365,1390]" pageId="11" pageNumber="235">Paratypes</emphasis>
</typeStatus>
.
<collectionCode id="ED1FC1D7897BDB4DFEEA7677FE65A711" box="[336,404,1366,1390]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="235">WAM</collectionCode>
M126, skull, sex unknown,
<location id="8ED10FC9897BDB4DFD5E7674FC92A712" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFD5E7674FC92A712" box="[740,867,1365,1390]" county="Rawlinna" municipality="Skin" name="Eucla Pass" pageId="11" pageNumber="235" stateProvince="Western Australia">Eucla Pass</location>
,
<collectingRegion id="49CA97F0897BDB4DFCCB7677FBCCA711" box="[881,1085,1365,1390]" country="Australia" name="Western Australia" pageId="11" pageNumber="235">
<collectingRegion id="49CA97F0897BDB4DFCCB7677FC3DA712" box="[881,972,1366,1389]" country="American Samoa" name="Western" pageId="11" pageNumber="235">Western</collectingRegion>
Australia
</collectingRegion>
(WA);
<collectionCode id="ED1FC1D7897BDB4DFB227677FB2CA711" box="[1176,1245,1366,1390]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="235">WAM</collectionCode>
M570, skin and skull, female,
<location id="8ED10FC9897BDB4DFE8E7658FE74A7ED" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFE8E7658FE74A7ED" box="[308,389,1401,1426]" county="Rawlinna" municipality="Skin" name="Ooldea" pageId="11" pageNumber="235" stateProvince="Western Australia">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897BDB4DFE6A765BFDE5A7ED" box="[464,532,1402,1426]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="235">WAM</collectionCode>
M572, skin and skull (juvenile), male,
<location id="8ED10FC9897BDB4DFC707658FBEAA7ED" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFC707658FBEAA7ED" box="[970,1051,1401,1426]" county="Rawlinna" municipality="Skin" name="Ooldea" pageId="11" pageNumber="235" stateProvince="Western Australia">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897BDB4DFBDD765BFB5AA7ED" box="[1127,1195,1402,1426]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="235">WAM</collectionCode>
M574, skin and skull (
<location id="8ED10FC9897BDB4DFF2576BCFEF3A7C9" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFF2576BCFEF3A7C9" box="[159,258,1437,1462]" county="Rawlinna" municipality="Skin" name="Juvenile" pageId="11" pageNumber="235" stateProvince="Western Australia">Juvenile</location>
), male,
<location id="8ED10FC9897BDB4DFEE476BCFE5EA7C9" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFEE476BCFE5EA7C9" box="[350,431,1437,1462]" county="Rawlinna" municipality="Skin" name="Ooldea" pageId="11" pageNumber="235" stateProvince="Western Australia">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897BDB4DFDBB76BFFDB7A7C9" box="[513,582,1438,1462]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="235">WAM</collectionCode>
M576, skin and skull, female,
<location id="8ED10FC9897BDB4DFC0A76BCFBF0A7C9" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFC0A76BCFBF0A7C9" box="[944,1025,1437,1462]" county="Rawlinna" municipality="Skin" name="Ooldea" pageId="11" pageNumber="235" stateProvince="Western Australia">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897BDB4DFBE976BFFB69A7C9" box="[1107,1176,1438,1462]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="235">WAM</collectionCode>
M577, skin and skull, female,
<location id="8ED10FC9897BDB4DFF4B76E0FEB3A7A5" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFF4B76E0FEB3A7A5" box="[241,322,1473,1498]" county="Rawlinna" municipality="Skin" name="Ooldea" pageId="11" pageNumber="235" stateProvince="Western Australia">Ooldea</location>
(SA); AM M4352,
<collectingMunicipality id="6BD5C368897BDB4DFD9876E0FDA4A7A5" box="[546,597,1473,1498]" pageId="11" pageNumber="235">Skin</collectingMunicipality>
and skull, female,
<collectingCounty id="62D0219E897BDB4DFC9676E0FC69A7A5" box="[812,920,1473,1498]" pageId="11" pageNumber="235">Rawlinna</collectingCounty>
(WA); AM M31547, alcohol specimen, male, Ooldea (SA)
</materialsCitation>
;
<materialsCitation id="3B66534F897BDB4DFE8076C7FCA6A781" ID-GBIF-Occurrence="1806521142" box="[314,855,1509,1534]" collectionCode="BMNH" location="Ooldea" pageId="11" pageNumber="235" specimenCode="BMNH 1925.10" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<specimenCode id="DBA8F169897BDB4DFE8076C7FE07A781" box="[314,502,1509,1534]" collectionCode="BMNH" pageId="11" pageNumber="235">BMNH 1925.10</specimenCode>
.8.26, skin, male,
<location id="8ED10FC9897BDB4DFD7876C4FCE2A781" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFD7876C4FCE2A781" box="[706,787,1509,1534]" name="Ooldea" pageId="11" pageNumber="235" stateProvince="Western Australia">Ooldea</location>
(SA)
</materialsCitation>
;
<materialsCitation id="3B66534F897BDB4DFCDF76C7FDEBA439" ID-GBIF-Occurrence="1806521144" collectionCode="SAM" location="Ooldea" municipality="Ooldea" pageId="11" pageNumber="235" specimenCode="BMNH 1925.10" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<specimenCode id="DBA8F169897BDB4DFCDF76C7FBD2A781" box="[869,1059,1509,1534]" collectionCode="BMNH" pageId="11" pageNumber="235">BMNH 1925.10</specimenCode>
.8.27, skin, female,
<location id="8ED10FC9897BDB4DFAB976C4FAA5A781" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFAB976C4FAA5A781" box="[1283,1364,1509,1534]" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235" stateProvince="Western Australia">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897BDB4DFF2D752BFF25A45D" box="[151,212,1546,1570]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="11" pageNumber="235">SAM</collectionCode>
M3973, alcohol specimen and skull, female, collected
<locationDeviation id="9A034030897BDB4DFCF77528FB5EA45D" box="[845,1199,1545,1570]" pageId="11" pageNumber="235">between Ooldea and Tallaringa</locationDeviation>
(SA);
<collectionCode id="ED1FC1D7897BDB4DFB45752BFACDA45D" box="[1279,1340,1546,1570]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="11" pageNumber="235">SAM</collectionCode>
M4640, skin and skull, male,
<collectingMunicipality id="6BD5C368897BDB4DFE3F750CFE27A439" box="[389,470,1581,1606]" pageId="11" pageNumber="235">Ooldea</collectingMunicipality>
(SA).
</materialsCitation>
</paragraph>
<paragraph id="8BB15912897BDB4AFF7D7570FDE0A1DB" blockId="11.[151,1437,1293,2038]" lastBlockId="12.[151,1437,151,2013]" lastPageId="12" lastPageNumber="236" pageId="11" pageNumber="235">
<emphasis id="B97A8500897BDB4DFF7D7570FE4DA415" bold="true" box="[199,444,1617,1642]" pageId="11" pageNumber="235">Referred specimens.</emphasis>
AMNH 196371, 2 left subfossil dentaries, Horseshoe Cave, Nullarbor (WA);
<materialsCitation id="3B66534F897BDB4AFAFD7573FB53A067" ID-GBIF-Occurrence="1806521145" collectionCode="MAGNT, NMV, SAM" county="Ooldea" lastPageId="12" lastPageNumber="236" location="south west Loongana" municipality="Ooldea" pageId="11" pageNumber="235" specimenCode="AMNH 220153, AMNH 220154" specimenCount="1">
<specimenCode id="DBA8F169897BDB4DFAFD7573FF1AA4F1" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="235">
AMNH
<date id="FFB07FD2897BDB4DFF2D7554FF1AA4F1" box="[151,235,1653,1678]" pageId="11" pageNumber="235" value="53-01-22">220153</date>
</specimenCode>
and
<specimenCode id="DBA8F169897BDB4DFE9F7557FE2AA4F1" box="[293,475,1653,1678]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="235">
AMNH
<date id="FFB07FD2897BDB4DFE3E7554FE2AA4F1" box="[388,475,1653,1678]" pageId="11" pageNumber="235" value="54-01-22">220154</date>
</specimenCode>
, juvenile subfossil skulls,
<location id="8ED10FC9897BDB4DFCB37554FC0EA4F1" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFCB37554FC0EA4F1" box="[777,1023,1653,1678]" county="Ooldea" municipality="Ooldea" name="south west Loongana" pageId="11" pageNumber="235">south west Loongana</location>
, Nullarbor (WA); AM S1821, skull, juvenile,
<location id="8ED10FC9897BDB4DFF4575B8FE9BA4CD" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFF4575B8FE9BA4CD" box="[255,362,1689,1714]" county="Ooldea" municipality="Ooldea" name="Rawlinna" pageId="11" pageNumber="235">Rawlinna</location>
(WA); AM M2986, skin and skull, female, Fisher, Nullarbor (SA); AM M4353, skin and skull, juvenile female,
<location id="8ED10FC9897BDB4DFEED759CFE33A4A9" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFEED759CFE33A4A9" box="[343,450,1725,1750]" county="Ooldea" municipality="Ooldea" name="Rawlinna" pageId="11" pageNumber="235">Rawlinna</location>
(WA); AM M4850, skin and skull, male,
<location id="8ED10FC9897BDB4DFC13759CFC0BA4A9" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFC13759CFC0BA4A9" box="[937,1018,1725,1750]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
(SA); AM M4851 and M4852, skin and skull, female,
<quantity id="4CF6F4F7897BDB4DFEDF75C3FE46A485" box="[357,439,1761,1786]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="11" pageNumber="235" unit="mi" value="9.0">9 miles</quantity>
west of
<location id="8ED10FC9897BDB4DFDAC75C0FD96A485" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFDAC75C0FD96A485" box="[534,615,1761,1786]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
(SA); AM M3049, alcohol specimen, juvenile female,
<location id="8ED10FC9897BDB4DFB6875C0FAD2A485" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFB6875C0FAD2A485" box="[1234,1315,1761,1786]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
(SA); AM M4368M4369, alcohol specimen, juvenile female,
<location id="8ED10FC9897BDB4DFCBB7424FC9DA561" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFCBB7424FC9DA561" box="[769,876,1797,1822]" county="Ooldea" municipality="Ooldea" name="Rawlinna" pageId="11" pageNumber="235">Rawlinna</location>
(WA); AM M4370M4371, alcohol specimen, juvenile male,
<location id="8ED10FC9897BDB4DFE847408FE58A53D" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFE847408FE58A53D" box="[318,425,1833,1858]" county="Ooldea" municipality="Ooldea" name="Rawlinna" pageId="11" pageNumber="235">Rawlinna</location>
(WA); AM M4939 and M4941, alcohol specimen, pouch young male,
<location id="8ED10FC9897BDB4DFB6A7408FAD0A53D" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFB6A7408FAD0A53D" box="[1232,1313,1833,1858]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
(SA); AM M4940, alcohol specimen, pouch young female,
<location id="8ED10FC9897BDB4DFD71746CFCEDA519" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFD71746CFCEDA519" box="[715,796,1869,1894]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
(SA); AM M49424943, alcohol specimen, male,
<location id="8ED10FC9897BDB4DFAD6746FFEFAA5F5" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFAD6746FFEFAA5F5" county="Ooldea" municipality="Ooldea" name="East West Line" pageId="11" pageNumber="235">East West Line</location>
(SA); AM M49444945, alcohol specimen, female,
<location id="8ED10FC9897BDB4DFCD77453FBEAA5F5" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFCD77453FBEAA5F5" box="[877,1051,1905,1930]" county="Ooldea" municipality="Ooldea" name="East West Line" pageId="11" pageNumber="235">East West Line</location>
(SA); AM M49784979, alcohol specimen, pouch young female of M4852, 9 miles west of
<location id="8ED10FC9897BDB4DFC9574B4FC71A5D1" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFC9574B4FC71A5D1" box="[815,896,1941,1966]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
(SA); AM M31545, alcohol specimen, juvenile male,
<location id="8ED10FC9897BDB4DFF617498FEDDA5AD" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFF617498FEDDA5AD" box="[219,300,1977,2002]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
(SA); AM M31548, alcohol specimen, male,
<location id="8ED10FC9897BDB4DFC917498FC8FA5AD" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFC917498FC8FA5AD" box="[811,894,1977,2002]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="11" pageNumber="235">Ooldea</location>
? (SA);
<collectionCode id="ED1FC1D7897BDB4DFC69749BFBCBA5AD" box="[979,1082,1978,2002]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34986" name="Museum and Art Gallery of the Northern Territory" pageId="11" pageNumber="235">MAGNT</collectionCode>
U7605, skull, specimen bred in
<location id="8ED10FC9897BDB4DFF2D74FCFF0DA589" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897BDB4DFF2D74FCFF0DA589" box="[151,252,2013,2038]" county="Ooldea" municipality="Ooldea" name="Adelaide" pageId="11" pageNumber="235">Adelaide</location>
in 1926 (original locality unknown);
<collectionCode id="ED1FC1D7897BDB4DFD1A74FFFD10A58A" box="[672,737,2014,2037]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="11" pageNumber="235">NMV</collectionCode>
C7173 and C7174, subfossil skulls, N11 Cave, Nullarbor East (SA);
<collectionCode id="ED1FC1D7897CDB4AFF5A73B9FED0A2D0" box="[224,289,152,175]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="236">NMV</collectionCode>
C7175 and C7176, subfossil skulls, Weebubbie Cave, Nullarbor (WA);
<collectionCode id="ED1FC1D7897CDB4AFBCE73B9FB44A2D0" box="[1140,1205,152,175]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="236">NMV</collectionCode>
C7177 and C7178, subfossil skulls, Weekes Cave, Nullarbor (SA);
<collectionCode id="ED1FC1D7897CDB4AFD7F739CFCF7A2AB" box="[709,774,189,212]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="12" pageNumber="236">NMV</collectionCode>
C7180, 2 subfossil dentaries, Koonalda Cave, Nullarbor (SA); QM J5028, alcohol specimen, female,
<location id="8ED10FC9897CDB4AFD2473FEFCFAA287" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFD2473FEFCFAA287" box="[670,779,223,248]" county="Ooldea" municipality="Ooldea" name="Nullarbor" pageId="12" pageNumber="236">Nullarbor</location>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFCE773C1FC6BA287" box="[861,922,224,248]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M846, alcohol specimen and skull, juvenile female,
<location id="8ED10FC9897CDB4AFF4B7225FEB3A362" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFF4B7225FEB3A362" box="[241,322,260,285]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="12" pageNumber="236">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFE357225FE3DA363" box="[399,460,260,284]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M1394, mount, male,
<location id="8ED10FC9897CDB4AFD767225FCECA362" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFD767225FCECA362" box="[716,797,260,285]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="12" pageNumber="236">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFCD07225FC56A363" box="[874,935,260,284]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M1395, mount,
<location id="8ED10FC9897CDB4AFBD97225FB45A362" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFBD97225FB45A362" box="[1123,1204,260,285]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="12" pageNumber="236">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFABA7225FACCA363" box="[1280,1341,260,284]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M1396, mount, female,
<location id="8ED10FC9897CDB4AFEF47206FE6EA33F" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFEF47206FE6EA33F" box="[334,415,295,320]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="12" pageNumber="236">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFE487209FDDEA33F" box="[498,559,296,320]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M1397, skin, male
<location id="8ED10FC9897CDB4AFCA77206FC9FA33F" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFCA77206FC9FA33F" box="[797,878,295,320]" county="Ooldea" municipality="Ooldea" name="Ooldea" pageId="12" pageNumber="236">Ooldea</location>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFC7B7209FC0FA33F" box="[961,1022,296,320]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M1398, skin, female
<collectingMunicipality id="6BD5C368897CDB4AFAB87206FAA2A33F" box="[1282,1363,295,320]" pageId="12" pageNumber="236">Ooldea</collectingMunicipality>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFF2D726DFF25A31B" box="[151,212,332,356]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M39263927, M3976, M39813982, M3984, M39873988, alcohol specimen and skull, female, no location data;
<collectionCode id="ED1FC1D7897CDB4AFF627251FEE4A3F7" box="[216,277,368,392]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M3977, M3983, M39853986, alcohol specimen, juvenile, no location data;
<collectionCode id="ED1FC1D7897CDB4AFB217251FB29A3F7" box="[1179,1240,368,392]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M3980, M3989, alcohol specimen and skull, male, no location data;
<collectionCode id="ED1FC1D7897CDB4AFD5772B5FCDBA3D3" box="[749,810,404,428]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M4639, skin and skull, juvenile female,
<collectingCounty id="62D0219E897CDB4AFABE72B5FAA4A3D2" box="[1284,1365,404,429]" pageId="12" pageNumber="236">Ooldea</collectingCounty>
(SA);
<collectionCode id="ED1FC1D7897CDB4AFF2D7299FF25A3AF" box="[151,212,440,464]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M5226, mount, no data;
<collectionCode id="ED1FC1D7897CDB4AFDBD7299FDB5A3AF" box="[519,580,440,464]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M6302, many subfossil skulls, Weebubbie Cave, Nullarbor (WA);
<collectionCode id="ED1FC1D7897CDB4AFADA7299FA6CA3AF" box="[1376,1437,440,464]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M6303, many subfossil skulls, Weekes Cave, Nullarbor (SA);
<collectionCode id="ED1FC1D7897CDB4AFCD572FDFC5AA38B" box="[879,939,476,500]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M6304, many subfossil skulls, N11 Cave, Nullarbor East (SA);
<collectionCode id="ED1FC1D7897CDB4AFE317121FE39A067" box="[395,456,512,536]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="12" pageNumber="236">SAM</collectionCode>
M6305, many subfossil skulls, Koonalda Cave, Nullarbor (SA)
</materialsCitation>
; WAM 76.4.21, skull, Abrakurrie Cave, Nullarbor (WA); WAM 67.4.114116, 67.10.305307, skulls, Homestead Cave, Nullarbor (WA); WAM 66.6.4553, 66.1.54, 68.3.6367, 69.5.17, skulls, Murra-el-elevyn Cave, Nullarbor (WA); WAM 69.7.557 559, skulls, Kjeldahl Cave, Cocklebiddy Roadhouse, Nullarbor (WA); WAM 73.1.107, 74.5.39, skulls, Webbs Cave, Nullarbor (WA); WAM 67.5.128, 67.5.130, skulls, Warbla Cave, Nullarbor (SA); WAM 67.10.420421, 68.3.118, skulls, Blowhole, Eucla, Nullarbor (WA); WAM 67.4.250, 72.1.8, 72.1.10881089, skulls, Horseshoe Cave, Nullarbor (WA); WAM 66.6.4553, skulls, Grahams Cave, Nullarbor (WA);
<materialsCitation id="3B66534F897CDB4AFBFE71F9FD1AA16B" ID-GBIF-Occurrence="1806521147" collectionCode="WAM" location="South of Cocklebiddy Cave" municipality="Nullarbor" pageId="12" pageNumber="236" specimenCode="WAM 68.3" specimenCount="1">
<specimenCode id="DBA8F169897CDB4AFBFE71F9FB37A08F" box="[1092,1222,727,752]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="236">WAM 68.3</specimenCode>
.51, skull, cave 3 3/
<quantity id="4CF6F4F7897CDB4AFF2D71DDFF2EA16A" box="[151,223,764,789]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="12" pageNumber="236" unit="mi" value="4.0">4 mile</quantity>
<location id="8ED10FC9897CDB4AFF5271DDFDD7A16B" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFF5271DDFDD7A16B" box="[232,550,764,789]" municipality="Nullarbor" name="South of Cocklebiddy Cave" pageId="12" pageNumber="236">South of Cocklebiddy Cave</location>
,
<collectingMunicipality id="6BD5C368897CDB4AFD8E71DDFD50A16A" box="[564,673,764,789]" pageId="12" pageNumber="236">Nullarbor</collectingMunicipality>
(WA)
</materialsCitation>
; WAM 68.3.33, skull, Mullamullang Cave, Nullarbor (WA); WAM 67.11.32, skull, Capstan Cave, Nullarbor (WA);
<materialsCitation id="3B66534F897CDB4AFC907001FEBFA123" ID-GBIF-Occurrence="1806521149" collectionCode="WAM" location="Eucla" municipality="Nullarbor" pageId="12" pageNumber="236" specimenCode="WAM 67.4" specimenCount="1">
<specimenCode id="DBA8F169897CDB4AFC907001FC40A147" box="[810,945,799,824]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="236">WAM 67.4</specimenCode>
.343, skull, cave
<quantity id="4CF6F4F7897CDB4AFBC77001FB15A147" box="[1149,1252,799,824]" metricMagnitude="4" metricUnit="m" metricValue="3.7014912" pageId="12" pageNumber="236" unit="mi" value="23.0">23 miles</quantity>
west of
<location id="8ED10FC9897CDB4AFAEF703EFA69A147" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFAEF703EFA69A147" box="[1365,1432,799,824]" municipality="Nullarbor" name="Eucla" pageId="12" pageNumber="236">Eucla</location>
,
<collectingMunicipality id="6BD5C368897CDB4AFF2D7065FEF5A122" box="[151,260,836,861]" pageId="12" pageNumber="236">Nullarbor</collectingMunicipality>
(WA)
</materialsCitation>
;
<materialsCitation id="3B66534F897CDB4AFEE07065FB96A123" ID-GBIF-Occurrence="1806521151" box="[346,1127,836,861]" collectionCode="WAM" location="Madura" municipality="Nullarbor" pageId="12" pageNumber="236" specimenCode="WAM 68.3" specimenCount="1">
<specimenCode id="DBA8F169897CDB4AFEE07065FE2AA122" box="[346,475,836,861]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="236">WAM 68.3</specimenCode>
.20, skull, cave
<quantity id="4CF6F4F7897CDB4AFD307065FD2FA122" box="[650,734,836,861]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="12" pageNumber="236" unit="mi" value="6.0">6 miles</quantity>
south of
<location id="8ED10FC9897CDB4AFCF07065FC55A122" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFCF07065FC55A122" box="[842,932,836,861]" municipality="Nullarbor" name="Madura" pageId="12" pageNumber="236">Madura</location>
,
<collectingMunicipality id="6BD5C368897CDB4AFC0A7065FBECA122" box="[944,1053,836,861]" pageId="12" pageNumber="236">Nullarbor</collectingMunicipality>
(WA)
</materialsCitation>
;
<materialsCitation id="3B66534F897CDB4AFBC97065FDE0A1DB" ID-GBIF-Occurrence="1806521153" collectionCode="WAM" location="Madura" municipality="Nullarbor" pageId="12" pageNumber="236" specimenCode="WAM 67.9" specimenCount="1">
<specimenCode id="DBA8F169897CDB4AFBC97065FB07A122" box="[1139,1270,836,861]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="236">WAM 67.9</specimenCode>
.134, cave near
<location id="8ED10FC9897CDB4AFF2D7046FF00A1FF" LSID="urn:lsid:plazi:treatment:03A7E804897BDB57FF2D7783FAA1A193:8ED10FC9897CDB4AFF2D7046FF00A1FF" box="[151,241,871,896]" municipality="Nullarbor" name="Madura" pageId="12" pageNumber="236">Madura</location>
, in doline, Dingo Douga, Nullarbor (WA);
<collectionCode id="ED1FC1D7897CDB4AFD407049FCCEA1FF" box="[762,831,872,896]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="236">WAM</collectionCode>
66.3.1416, 67.10.177182, skulls, Firestick Cave, Nullarbor (WA);
<collectingMunicipality id="6BD5C368897CDB4AFEE270ADFE34A1DA" box="[344,453,908,933]" pageId="12" pageNumber="236">Nullarbor</collectingMunicipality>
(WA).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3140A99897CDB4AFF7D708EFE01A6DF" pageId="12" pageNumber="236" type="diagnosis">
<paragraph id="8BB15912897CDB4AFF7D708EFE01A6DF" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="236">
<emphasis id="B97A8500897CDB4AFF7D708EFECBA1B7" bold="true" box="[199,314,943,968]" pageId="12" pageNumber="236">Diagnosis</emphasis>
.
<taxonomicName id="4C0E2291897CDB4AFEFD708EFDD0A1B7" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[327,545,943,968]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897CDB4AFEFD708EFDD0A1B7" box="[327,545,943,968]" italics="true" pageId="12" pageNumber="236">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897CDB4AFD92708EFD70A1B7" box="[552,641,943,968]" pageId="12" pageNumber="236" rank="species">
<emphasis id="B97A8500897CDB4AFD92708EFD70A1B7" bold="true" box="[552,641,943,968]" pageId="12" pageNumber="236">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from other species of
<taxonomicName id="4C0E2291897CDB4AFC73708EFBCEA1B7" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[969,1087,943,968]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897CDB4AFC73708EFBCEA1B7" box="[969,1087,943,968]" italics="true" pageId="12" pageNumber="236">Perameles</emphasis>
</taxonomicName>
in having a dark horizontal bar on its ears, a bracelet of light brown fur around its wrists and covering the heel of the foot and the posterior edges of the footpad, a dark butterfly pattern on its rump, shorter pes and tail (see results), enlarged lacrimal crest (except for
<emphasis id="B97A8500897CDB4AFF7A773CFEC4A64C" box="[192,309,1052,1076]" italics="true" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFF7A773CFED8A64C" authorityName="Gray" authorityYear="1838" box="[192,297,1052,1076]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="gunnii">P. gunnii</taxonomicName>
),
</emphasis>
enlarged bullae (largest of any
<taxonomicName id="4C0E2291897CDB4AFD18773DFCE9A64A" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[674,792,1052,1077]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897CDB4AFD18773DFCE9A64A" box="[674,792,1052,1077]" italics="true" pageId="12" pageNumber="236">Perameles</emphasis>
</taxonomicName>
), accessory fenestrae extending anteriorly to the canine, maxillopalatine fenestrae extending posteriorly to the M3, well-developed squamosal epitympanic sinus.
<taxonomicName id="4C0E2291897CDB4AFF2D7745FE80A602" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[151,369,1124,1149]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897CDB4AFF2D7745FE80A602" box="[151,369,1124,1149]" italics="true" pageId="12" pageNumber="236">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897CDB4AFEC37745FE23A602" box="[377,466,1124,1149]" pageId="12" pageNumber="236" rank="species">
<emphasis id="B97A8500897CDB4AFEC37745FE23A602" bold="true" box="[377,466,1124,1149]" pageId="12" pageNumber="236">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all members of
<taxonomicName id="4C0E2291897CDB4AFCAD7745FC2CA602" authorityName="Kirsch" authorityYear="1968" box="[791,989,1124,1149]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
(except
<taxonomicName id="4C0E2291897CDB4AFB877744FB32A603" baseAuthorityName="Wagner" baseAuthorityYear="1841" box="[1085,1219,1125,1148]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="myosuros">
<emphasis id="B97A8500897CDB4AFB877744FB32A603" box="[1085,1219,1125,1148]" italics="true" pageId="12" pageNumber="236">P. myosuros</emphasis>
</taxonomicName>
) in having females distinctively larger than males.
</paragraph>
</subSubSection>
<subSubSection id="C3140A99897CDB4AFF7D778DFAE3A6BB" box="[199,1298,1196,1221]" pageId="12" pageNumber="236" type="etymology">
<paragraph id="8BB15912897CDB4AFF7D778DFAE3A6BB" blockId="12.[151,1437,151,2013]" box="[199,1298,1196,1221]" pageId="12" pageNumber="236">
<emphasis id="B97A8500897CDB4AFF7D778DFEB7A6BA" bold="true" box="[199,326,1196,1221]" pageId="12" pageNumber="236">Etymology</emphasis>
. Papillon, French for butterfly, in reference to the butterfly shaped pattern on its rump.
</paragraph>
</subSubSection>
<subSubSection id="C3140A99897CDB4AFF7D77EEFD97A40B" pageId="12" pageNumber="236" type="discussion">
<paragraph id="8BB15912897CDB4AFF7D77EEFE01A79A" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="236">
<emphasis id="B97A8500897CDB4AFF7D77EEFEC5A697" bold="true" box="[199,308,1231,1256]" pageId="12" pageNumber="236">Remarks</emphasis>
.
<bibRefCitation id="EF9F24E3897CDB4AFEFE77F0FDA6A697" box="[324,599,1231,1256]" pageId="12" pageNumber="236" refId="ref20011" refString="Westerman, M., Kear, B. P., Aplin, K., Meredith, R. W., Emerling, C. &amp; Springer, M. S. (2012) Phylogenetic relationships of living and recently extinct bandicoots based on nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 62, 97 - 108. https: // doi. org / 10.1016 / j. ympev. 2011.09.009" type="journal article">
Westerman
<emphasis id="B97A8500897CDB4AFE7077F0FDF5A697" box="[458,516,1231,1256]" italics="true" pageId="12" pageNumber="236">et al.</emphasis>
(2012)
</bibRefCitation>
subsampled specimen AM M31545 from the Australian Museum, which was attributed to
<emphasis id="B97A8500897CDB4AFEE677D5FDB7A773" box="[348,582,1268,1293]" italics="true" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFEE677D5FDB0A773" box="[348,577,1268,1293]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="subSpecies" species="bougainville" subSpecies="notina">Perameles b. notina</taxonomicName>
,
</emphasis>
from Ooldea in
<collectingRegion id="49CA97F0897CDB4AFCBE77D5FC49A772" box="[772,952,1268,1293]" country="Australia" name="South Australia" pageId="12" pageNumber="236">
South
<collectingCountry id="F3191982897CDB4AFCF477D5FC49A772" box="[846,952,1268,1293]" name="Australia" pageId="12" pageNumber="236">Australia</collectingCountry>
</collectingRegion>
. After scoring specimens from this region, including the rest of the Nullarbor Plain in
<collectingRegion id="49CA97F0897CDB4AFDC37638FCB5A74F" box="[633,836,1303,1328]" country="Australia" name="Western Australia" pageId="12" pageNumber="236">
<collectingRegion id="49CA97F0897CDB4AFDC37638FD25A74F" box="[633,724,1305,1328]" country="American Samoa" name="Western" pageId="12" pageNumber="236">Western</collectingRegion>
<collectingCountry id="F3191982897CDB4AFD617636FCB5A74F" box="[731,836,1303,1328]" name="Australia" pageId="12" pageNumber="236">Australia</collectingCountry>
</collectingRegion>
, and comparing them to the
<typeStatus id="54B5E7B0897CDB4AFB397636FB14A74F" box="[1155,1253,1303,1328]" pageId="12" pageNumber="236" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0E2291897CDB4AFAB07636FF2EA72B" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="subSpecies" species="bougainville" subSpecies="notina">
<emphasis id="B97A8500897CDB4AFAB07636FF2EA72B" italics="true" pageId="12" pageNumber="236">Perameles b. notina</emphasis>
</taxonomicName>
(then named
<taxonomicName id="4C0E2291897CDB4AFE38761DFD47A72B" authorityName="Thomas" authorityYear="1922" box="[386,694,1340,1365]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897CDB4AFE38761DFD47A72B" box="[386,694,1340,1365]" italics="true" pageId="12" pageNumber="236">Perameles myosura notina</emphasis>
</taxonomicName>
) BMNH 43.8.12.21 from the head of St Vincent Gulf in
<collectingRegion id="49CA97F0897CDB4AFAE1761DFF0FA707" country="Australia" name="South Australia" pageId="12" pageNumber="236">
South
<collectingCountry id="F3191982897CDB4AFF2D767EFF0FA707" box="[151,254,1375,1400]" name="Australia" pageId="12" pageNumber="236">Australia</collectingCountry>
</collectingRegion>
, it became evident that they did not match in morphology, the
<typeStatus id="54B5E7B0897CDB4AFC0D7640FC16A706" box="[951,999,1377,1401]" pageId="12" pageNumber="236">type</typeStatus>
being much larger and several features on the skin and skull did not match any of the specimens recovered from the Nullarbor region. As a result, AM M31545 and all specimens found in the Nullarbor regions (present in collections in various museums) are considered here a new species.
</paragraph>
<paragraph id="8BB15912897CDB4AFF7D76CEFD97A40B" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="236">
The results of our analysis also show that
<taxonomicName id="4C0E2291897CDB4AFD7376CEFC58A477" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[713,937,1519,1544]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897CDB4AFD7376CEFC58A477" box="[713,937,1519,1544]" italics="true" pageId="12" pageNumber="236">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897CDB4AFC0C76CEFBE4A477" box="[950,1045,1519,1544]" pageId="12" pageNumber="236" rank="species">
<emphasis id="B97A8500897CDB4AFC0C76CEFBE4A477" bold="true" box="[950,1045,1519,1544]" pageId="12" pageNumber="236">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinct morphologically from
<taxonomicName id="4C0E2291897CDB4AFF2D7535FE50A452" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,417,1556,1581]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="bougainville">
<emphasis id="B97A8500897CDB4AFF2D7535FE50A452" box="[151,417,1556,1581]" italics="true" pageId="12" pageNumber="236">Perameles bougainville</emphasis>
</taxonomicName>
and
<emphasis id="B97A8500897CDB4AFE6D7534FD92A453" box="[471,611,1557,1580]" italics="true" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFE6D7534FDAEA453" authorityName="Spencer" authorityYear="1897" box="[471,607,1557,1580]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
,
</emphasis>
but also distinct to the currently unrecognised
<emphasis id="B97A8500897CDB4AFBCE7534FA9DA453" box="[1140,1388,1557,1580]" italics="true" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFBCE7534FB0CA453" baseAuthorityName="Wagner" baseAuthorityYear="1841" box="[1140,1277,1557,1580]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="myosuros">P. myosuros</taxonomicName>
,
<taxonomicName id="4C0E2291897CDB4AFAB27534FA9DA453" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1288,1388,1557,1580]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="notina">P. notina</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897CDB4AFF2D7518FEE2A42F" box="[151,275,1592,1616]" italics="true" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFF2D7518FEE1A42F" baseAuthorityName="Gray" baseAuthorityYear="1841" box="[151,272,1592,1616]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="fasciata">P. fasciata</taxonomicName>
.
</emphasis>
Our results further suggest that the latter three taxa are distinct taxa and should no longer be subsumed in
<taxonomicName id="4C0E2291897CDB4AFF0E757CFEAAA40A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[180,347,1628,1653]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="bougainville">
<emphasis id="B97A8500897CDB4AFF0E757CFEAAA40A" box="[180,347,1628,1653]" italics="true" pageId="12" pageNumber="236">P. bougainville</emphasis>
</taxonomicName>
(see discussion below).
</paragraph>
</subSubSection>
<subSubSection id="C3140A99897CDB4AFF7D755EFDB0A4C3" pageId="12" pageNumber="236" type="distribution">
<paragraph id="8BB15912897CDB4AFF7D755EFDB0A4C3" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="236">
<emphasis id="B97A8500897CDB4AFF7D755EFEA8A4E7" bold="true" box="[199,345,1663,1688]" pageId="12" pageNumber="236">Distribution</emphasis>
. Collected only in the Nullarbor Plain, on either side of the
<collectingRegion id="49CA97F0897CDB4AFB8775A0FB69A4E7" box="[1085,1176,1665,1688]" country="American Samoa" name="Western" pageId="12" pageNumber="236">Western</collectingRegion>
Australian and South Australian border (triangles in
<figureCitation id="13354597897CDB4AFE4A7585FDC2A4C3" box="[496,563,1700,1725]" captionStart="FIGURE 8" captionStartId="18.[151,250,1454,1476]" captionTargetBox="[166,1423,267,1424]" captionTargetId="figure@18.[151,1436,267,1433]" captionTargetPageId="18" captionText="FIGURE 8. Distribution of species in the Perameles bougainville complex, based on museum specimens (modern and some fossils). Circles, Perameles bougainville; squares, Perameles eremiana; crescents, Perameles myosuros; ovals, Perameles notina; star, Perameles fasciata; triangles,Perameles papillon sp. nov." httpUri="https://zenodo.org/record/1169513/files/figure.png" pageId="12" pageNumber="236">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3140A99897CDB57FF7D75E6FAA1A193" lastPageId="17" lastPageNumber="241" pageId="12" pageNumber="236" type="description">
<paragraph id="8BB15912897CDB4AFF7D75E6FC4CA50F" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="236">
<emphasis id="B97A8500897CDB4AFF7D75E6FEA0A49F" bold="true" box="[199,337,1735,1760]" pageId="12" pageNumber="236">Description</emphasis>
. The description of the skin is a comparison of
<taxonomicName id="4C0E2291897CDB4AFC2D75E6FB87A49F" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[919,1142,1735,1760]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897CDB4AFC2D75E6FB87A49F" box="[919,1142,1735,1760]" italics="true" pageId="12" pageNumber="236">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897CDB4AFB3875E6FB11A49F" box="[1154,1248,1735,1760]" pageId="12" pageNumber="236" rank="species">
<emphasis id="B97A8500897CDB4AFB3875E6FB11A49F" bold="true" box="[1154,1248,1735,1760]" pageId="12" pageNumber="236">sp. nov.</emphasis>
</taxonomicNameLabel>
, represented by
<typeStatus id="54B5E7B0897CDB4AFF2D75CDFF0EA57A" box="[151,255,1772,1797]" pageId="12" pageNumber="236" type="holotype">Holotype</typeStatus>
WAM M571 (
<figureCitation id="13354597897CDB4AFE1975CDFE17A57A" box="[419,486,1772,1797]" captionStart="FIGURE 5" captionStartId="13.[151,250,1959,1981]" captionTargetBox="[177,1418,193,1949]" captionTargetId="figure@13.[177,1418,193,1949]" captionTargetPageId="13" captionText="FIGURE 5. Perameles papillon sp. nov., study skins. AC, holotype WAM M571; DF, paratype WAM M574; GI, paratype WAM M572.A, D, and G, dorsal view; B, E, and H, lateral view; C, F, and I, ventral view. Scale = 5cm." httpUri="https://zenodo.org/record/1169507/files/figure.png" pageId="12" pageNumber="236">Fig. 5</figureCitation>
), to
<emphasis id="B97A8500897CDB4AFDA375CCFD13A57A" box="[537,738,1772,1797]" italics="true" pageId="12" pageNumber="236">
P. b.
<taxonomicName id="4C0E2291897CDB4AFDEA75CDFD2EA57A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[592,735,1772,1797]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
represented by WAM M3641 (Suppl.
<figureCitation id="13354597897CDB4AFB3575CDFB23A57B" box="[1167,1234,1772,1797]" captionStart="FIGURE 2" captionStartId="7.[151,250,1161,1183]" captionTargetBox="[163,1413,222,1123]" captionTargetId="figure@7.[151,1436,193,1140]" captionTargetPageId="7" captionText="FIGURE 2. Total evidence Bayesian inference timescale of peramelemorphian evolution. Node and tip calibrations are employed under a fossilized birth-death model, with molecular and morphological relaxed clock models unlinked. Blue bars show 95% highest posterior density intervals for node ages. The outgroup includes Didelphis and eight dasyuromorphians." httpUri="https://zenodo.org/record/1169501/files/figure.png" pageId="12" pageNumber="236">Fig. 2</figureCitation>
), and
<emphasis id="B97A8500897CDB4AFAAC75CCFA6DA57B" box="[1302,1436,1772,1797]" italics="true" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFAAC75CCFA66A57B" authorityName="Thomas" authorityYear="1922" box="[1302,1431,1772,1797]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
represented by the
<typeStatus id="54B5E7B0897CDB4AFED4742EFE21A557" box="[366,464,1807,1832]" pageId="12" pageNumber="236" type="holotype">holotype</typeStatus>
BMNH 43.8.12.21, unless specified otherwise (Suppl.
<figureCitation id="13354597897CDB4AFBFF742EFB78A557" box="[1093,1161,1807,1832]" captionStart="FIGURE3" captionStartId="9.[151,250,1769,1791]" captionTargetBox="[156,1394,719,1725]" captionTargetId="figure@9.[151,1436,697,1748]" captionTargetPageId="9" captionText="FIGURE3. Resultsof the morphometric analysisof thePerameles bougainville complex.A, Principal Component Analysis of cranial data; B, Canonical Variate Analysis of cranial data; C, Principal Component Analysis of dental data; D, Canonical Variate Analysis of dental data. Upright rectangles, Perameles bougainville; crosses, Perameles eremiana; diamonds, Perameles myosuros; circle, Perameles arenaria; stars, Perameles notina; triangles, Perameles fasciata; laying rectangles, Perameles papillon sp. nov." httpUri="https://zenodo.org/record/1169503/files/figure.png" pageId="12" pageNumber="236">Fig. 3</figureCitation>
).
<emphasis id="B97A8500897CDB4AFB257430FAC3A557" box="[1183,1330,1807,1832]" italics="true" pageId="12" pageNumber="236">
P. b.
<taxonomicName id="4C0E2291897CDB4AFB637431FAC3A557" baseAuthorityName="Gray" baseAuthorityYear="1841" box="[1241,1330,1808,1832]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="fasciata">fasciata</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897CDB4AFAD07430FEF0A533" italics="true" pageId="12" pageNumber="236">
P. b.
<taxonomicName id="4C0E2291897CDB4AFF2D7414FEF0A533" baseAuthorityName="Wagner" baseAuthorityYear="1841" box="[151,257,1845,1868]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="myosuros">myosuros</taxonomicName>
</emphasis>
are not used in the comparison here as they were not sampled by
<bibRefCitation id="EF9F24E3897CDB4AFC5C7414FB06A533" box="[998,1271,1844,1869]" pageId="12" pageNumber="236" refId="ref20011" refString="Westerman, M., Kear, B. P., Aplin, K., Meredith, R. W., Emerling, C. &amp; Springer, M. S. (2012) Phylogenetic relationships of living and recently extinct bandicoots based on nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 62, 97 - 108. https: // doi. org / 10.1016 / j. ympev. 2011.09.009" type="journal article">
Westerman
<emphasis id="B97A8500897CDB4AFBD17414FB52A532" box="[1131,1187,1844,1869]" italics="true" pageId="12" pageNumber="236">et al.</emphasis>
(2012)
</bibRefCitation>
, though we do suspect they are distinct taxa and will be investigated in future research.
</paragraph>
<paragraph id="8BB15912897CDB49FF6B745CFAC2A38B" blockId="12.[151,1437,151,2013]" lastBlockId="15.[151,1437,151,2013]" lastPageId="15" lastPageNumber="239" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFF6B745CFEBDA5EA" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[209,332,1916,1941]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897CDB4AFF6B745CFEBDA5EA" box="[209,332,1916,1941]" italics="true" pageId="12" pageNumber="236">P. papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897CDB4AFEEC745DFE43A5EA" box="[342,434,1916,1941]" pageId="12" pageNumber="236" rank="species">
<emphasis id="B97A8500897CDB4AFEEC745DFE43A5EA" bold="true" box="[342,434,1916,1941]" pageId="12" pageNumber="236">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar in size to
<emphasis id="B97A8500897CDB4AFD1E745CFC9CA5EA" box="[676,877,1916,1941]" italics="true" pageId="12" pageNumber="236">
P. b.
<taxonomicName id="4C0E2291897CDB4AFD5B745DFC9CA5EA" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[737,877,1916,1941]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
but much smaller than
<emphasis id="B97A8500897CDB4AFB38745CFAFCA5EB" box="[1154,1293,1916,1941]" italics="true" pageId="12" pageNumber="236">
<taxonomicName id="4C0E2291897CDB4AFB38745CFAF8A5EB" authorityName="Thomas" authorityYear="1922" box="[1154,1289,1916,1941]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="12" pageNumber="236" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
showing no marked sexual dimorphism between males and females, though females seem to be a little larger (see
<tableCitation id="C68C6CA9897CDB4AFAA77481FA8EA5C7" box="[1309,1407,1952,1977]" captionStart="TABLE 2" captionStartId="8.[151,239,1540,1563]" captionText="TABLE 2. Average skull length of males and females of Perameles bougainville complex taxon, including size ratios." pageId="12" pageNumber="236">Table S2</tableCitation>
3 for measurements,
<figureCitation id="13354597897CDB4AFED774E5FE5FA5A3" box="[365,430,1988,2013]" captionStart="FIGURE3" captionStartId="9.[151,250,1769,1791]" captionTargetBox="[156,1394,719,1725]" captionTargetId="figure@9.[151,1436,697,1748]" captionTargetPageId="9" captionText="FIGURE3. Resultsof the morphometric analysisof thePerameles bougainville complex.A, Principal Component Analysis of cranial data; B, Canonical Variate Analysis of cranial data; C, Principal Component Analysis of dental data; D, Canonical Variate Analysis of dental data. Upright rectangles, Perameles bougainville; crosses, Perameles eremiana; diamonds, Perameles myosuros; circle, Perameles arenaria; stars, Perameles notina; triangles, Perameles fasciata; laying rectangles, Perameles papillon sp. nov." httpUri="https://zenodo.org/record/1169503/files/figure.png" pageId="12" pageNumber="236">Fig. 3</figureCitation>
for morphometric analysis). The fur on the dorsal side of the head is a little darker and browner than
<emphasis id="B97A8500897EDB48FE8677FFFDFEA68A" box="[316,527,1244,1269]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FEC677FDFDFAA68A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[380,523,1244,1269]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
but lighter than
<emphasis id="B97A8500897EDB48FD6177FFFC9AA68A" box="[731,875,1244,1269]" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FD6177FFFC96A68A" authorityName="Thomas" authorityYear="1922" box="[731,871,1244,1269]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
.
</emphasis>
The ventral side is cream in colour as in
<emphasis id="B97A8500897EDB48FADD77FFFED2A766" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FF2D7621FED2A766" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,291,1280,1305]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
whereas it is greyer in
<taxonomicName id="4C0E2291897EDB48FD927623FD56A766" authorityName="Thomas" authorityYear="1922" box="[552,679,1280,1305]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897EDB48FD927623FD56A766" box="[552,679,1280,1305]" italics="true" pageId="14" pageNumber="238">P. b. notina</emphasis>
</taxonomicName>
. The ears have a noticeable dark horizontal bar across them, absent in both
<emphasis id="B97A8500897EDB48FF4F7607FE30A742" box="[245,449,1316,1341]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FE8F7605FE30A742" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[309,449,1316,1341]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897EDB48FDBA7607FD79A742" authorityName="Thomas" authorityYear="1922" box="[512,648,1316,1341]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897EDB48FDBA7607FD79A742" box="[512,648,1316,1341]" italics="true" pageId="14" pageNumber="238">P. b. notina</emphasis>
</taxonomicName>
or any other known peramelemorphian. Note that none of the wet specimens of
<taxonomicName id="4C0E2291897EDB48FE8E766BFE5EA71E" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[308,431,1352,1377]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897EDB48FE8E766BFE5EA71E" box="[308,431,1352,1377]" italics="true" pageId="14" pageNumber="238">P. papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897EDB48FE027669FDE2A71E" box="[440,531,1352,1377]" pageId="14" pageNumber="238" rank="species">
<emphasis id="B97A8500897EDB48FE027669FDE2A71E" bold="true" box="[440,531,1352,1377]" pageId="14" pageNumber="238">sp. nov.</emphasis>
</taxonomicNameLabel>
show this feature (dark bar on ear) but it is present in all dry skins, and this is suspected to be as a result of accelerated fading in alcohol specimens. The ears are long as is
<emphasis id="B97A8500897EDB48FB1D764FFA80A7FA" box="[1191,1393,1388,1413]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FB64764DFA9CA7FA" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1246,1389,1388,1413]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
but shorter than
<emphasis id="B97A8500897EDB48FE9F76B3FE5EA7D6" box="[293,431,1424,1449]" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FE9F76B3FE5AA7D6" authorityName="Thomas" authorityYear="1922" box="[293,427,1424,1449]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
.
</emphasis>
There are long dark vibrissae present at the front of the snout, above the eye and on the cheek below the eye. These are also present in
<emphasis id="B97A8500897EDB48FD6B7697FC6FA7B1" box="[721,926,1461,1486]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FCA87694FC6FA7B1" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[786,926,1461,1486]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897EDB48FC647697FB9FA7B2" box="[990,1134,1461,1486]" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FC647697FB9BA7B2" authorityName="Thomas" authorityYear="1922" box="[990,1130,1461,1486]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
but less obvious in
<emphasis id="B97A8500897EDB48FADC7697FED2A78E" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FF2D76F9FED2A78E" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,291,1496,1521]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
. The body is also darker and browner on the dorsal side than in
<emphasis id="B97A8500897EDB48FC4A76FBFB42A78E" box="[1008,1203,1496,1521]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FB9D76F9FB42A78E" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1063,1203,1496,1521]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
but lighter than
<emphasis id="B97A8500897EDB48FAD676FBFF17A46A" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FAD676FBFF13A46A" authorityName="Thomas" authorityYear="1922" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
.
</emphasis>
Both
<taxonomicName id="4C0E2291897EDB48FE9076DFFE53A469" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[298,418,1533,1558]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897EDB48FE9076DFFE53A469" box="[298,418,1533,1558]" italics="true" pageId="14" pageNumber="238">P. papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897EDB48FE1076DCFDF2A469" box="[426,515,1533,1558]" pageId="14" pageNumber="238" rank="species">
<emphasis id="B97A8500897EDB48FE1076DCFDF2A469" bold="true" box="[426,515,1533,1558]" pageId="14" pageNumber="238">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<emphasis id="B97A8500897EDB48FD8076DFFD0CA469" box="[570,765,1533,1558]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FDCB76DCFD0CA469" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[625,765,1533,1558]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
are cream coloured on the ventral side, while
<taxonomicName id="4C0E2291897EDB48FAB976DFFA72A46A" authorityName="Thomas" authorityYear="1922" box="[1283,1411,1533,1558]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897EDB48FAB976DFFA72A46A" box="[1283,1411,1533,1558]" italics="true" pageId="14" pageNumber="238">P. b. notina</emphasis>
</taxonomicName>
is mostly grey except for patches of cream. A complex pattern of different coloured bars is present on the rump (juveniles WAM M574 and M572 show the early stages of the development of this pattern on the rump), which is absent in
<emphasis id="B97A8500897EDB48FEB1754BFE29A4FE" box="[267,472,1640,1665]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FEF67549FE29A4FE" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[332,472,1640,1665]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
(
<emphasis id="B97A8500897EDB48FE56754BFD48A4FE" box="[492,697,1640,1665]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FD977549FD48A4FE" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[557,697,1640,1665]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
has a single very faint darkish bar across its rump that tapers laterally) but present in
<emphasis id="B97A8500897EDB48FE1175AFFDC7A4DA" box="[427,566,1677,1702]" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FE1175AFFDC0A4DA" authorityName="Thomas" authorityYear="1922" box="[427,561,1677,1702]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
though the
<typeStatus id="54B5E7B0897EDB48FD7875ACFCD5A4D9" box="[706,804,1677,1702]" pageId="14" pageNumber="238" type="holotype">holotype</typeStatus>
is faded, making this pattern less obvious. The pattern resembles that of
<taxonomicName id="4C0E2291897EDB48FEDA7593FE35A4B6" authorityName="Gray" authorityYear="1838" box="[352,452,1714,1737]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897EDB48FEDA7593FE35A4B6" box="[352,452,1714,1737]" italics="true" pageId="14" pageNumber="238">P. gunnii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E2291897EDB48FE467593FD8CA4B6" authorityName="Thomas" authorityYear="1922" box="[508,637,1712,1737]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897EDB48FE467593FD8CA4B6" box="[508,637,1712,1737]" italics="true" pageId="14" pageNumber="238">P. b. notina</emphasis>
</taxonomicName>
but it is less well defined. There isnt a central dark dorsal bar as in
<emphasis id="B97A8500897EDB48FA3D7593FF14A492" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FA3D7593FF10A492" authorityName="Gray" authorityYear="1838" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="gunnii">P. gunnii</taxonomicName>
,
</emphasis>
and most likely in
<emphasis id="B97A8500897EDB48FE7E75F7FDBDA492" box="[452,588,1749,1774]" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FE7E75F7FDB9A492" authorityName="Thomas" authorityYear="1922" box="[452,584,1749,1774]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
and the light bars are bicoloured, cream and golden brown (golden brown only in
<taxonomicName id="4C0E2291897EDB48FF4A75DBFEA4A56E" authorityName="Gray" authorityYear="1838" box="[240,341,1786,1809]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897EDB48FF4A75DBFEA4A56E" box="[240,341,1786,1809]" italics="true" pageId="14" pageNumber="238">P. gunnii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E2291897EDB48FE2A75DBFDE5A56E" authorityName="Thomas" authorityYear="1922" box="[400,532,1784,1809]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897EDB48FE2A75DBFDE5A56E" box="[400,532,1784,1809]" italics="true" pageId="14" pageNumber="238">P. b. notina</emphasis>
</taxonomicName>
). The anterior most two dark bars are almost completely black or at least dark brown (specimens are likely faded), and looks like butterfly wings in shape in dorsal view. The posterior darkest bars are much lighter in colour and less well defined, distinguishing
<taxonomicName id="4C0E2291897EDB48FC1C7463FBD0A526" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[934,1057,1856,1881]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897EDB48FC1C7463FBD0A526" box="[934,1057,1856,1881]" italics="true" pageId="14" pageNumber="238">P. papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897EDB48FB937461FB74A526" box="[1065,1157,1856,1881]" pageId="14" pageNumber="238" rank="species">
<emphasis id="B97A8500897EDB48FB937461FB74A526" bold="true" box="[1065,1157,1856,1881]" pageId="14" pageNumber="238">sp. nov.</emphasis>
</taxonomicNameLabel>
from
<taxonomicName id="4C0E2291897EDB48FB767463FAC1A526" authorityName="Gray" authorityYear="1838" box="[1228,1328,1858,1881]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897EDB48FB767463FAC1A526" box="[1228,1328,1858,1881]" italics="true" pageId="14" pageNumber="238">P. gunnii</emphasis>
</taxonomicName>
and
<emphasis id="B97A8500897EDB48FAD07463FF14A502" italics="true" pageId="14" pageNumber="238">
<taxonomicName id="4C0E2291897EDB48FAD07463FF10A502" authorityName="Thomas" authorityYear="1922" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
which have a well-defined posterior bar. The tail is short as in
<emphasis id="B97A8500897EDB48FC1E7447FB99A501" box="[932,1128,1893,1918]" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FC667444FB99A501" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[988,1128,1893,1918]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
(longer in
<taxonomicName id="4C0E2291897EDB48FB587447FA91A502" authorityName="Thomas" authorityYear="1922" box="[1250,1376,1893,1918]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897EDB48FB587447FA91A502" box="[1250,1376,1893,1918]" italics="true" pageId="14" pageNumber="238">P. b. notina</emphasis>
</taxonomicName>
), but it is tricoloured (dark brown dorsally, beige laterally and cream ventrally) instead of bicoloured as in
<emphasis id="B97A8500897EDB48FADF74ABFED2A5B9" italics="true" pageId="14" pageNumber="238">
P. b.
<taxonomicName id="4C0E2291897EDB48FF2D748CFED2A5B9" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,291,1965,1990]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
(the tail is dark brown dorsally and cream ventrally) or unicoloured as in
<taxonomicName id="4C0E2291897EDB48FBD3748FFB1BA5BA" authorityName="Thomas" authorityYear="1922" box="[1129,1258,1965,1990]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897EDB48FBD3748FFB1BA5BA" box="[1129,1258,1965,1990]" italics="true" pageId="14" pageNumber="238">P. b. notina</emphasis>
</taxonomicName>
(the tail is dark grey). The forelimbs are light brown in colour on the dorsal side up to the wrist, where it surrounds the wrist like a bracelet. The manus and the ventral side of the forelimbs are cream in colour. This pattern is different to
<emphasis id="B97A8500897FDB49FAD273B8FEDBA2AA" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FF2D739DFED7A2AA" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,294,188,213]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
which lacks the bracelet of brown fur around the wrist, and is lighter in colour dorsally. The forelimbs of
<taxonomicName id="4C0E2291897FDB49FF0273C0FECFA287" authorityName="Thomas" authorityYear="1922" box="[184,318,223,248]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FF0273C0FECFA287" box="[184,318,223,248]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
are completely grey. The manus has the three large toes (digit II, III and IV), and the typically reduced digits I and V as in all
<taxonomicName id="4C0E2291897FDB49FE437225FD76A362" authorityName="Gray" authorityYear="1825" box="[505,647,260,285]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
. Compared to
<emphasis id="B97A8500897FDB49FC917224FC1EA362" box="[811,1007,260,285]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FCD97225FC1EA362" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[867,1007,260,285]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897FDB49FB9C7224FB5CA363" box="[1062,1197,260,285]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FB9C7224FB58A363" authorityName="Thomas" authorityYear="1922" box="[1062,1193,260,285]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
the digits are shorter but have more pronounced toepads as in
<taxonomicName id="4C0E2291897FDB49FDE77208FD2AA33F" authorityName="Thomas" authorityYear="1922" box="[605,731,295,320]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FDE77208FD2AA33F" box="[605,731,295,320]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
(the toes of
<emphasis id="B97A8500897FDB49FCDC7208FBD9A33F" box="[870,1064,295,320]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FC267206FBD9A33F" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[924,1064,295,320]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
more slender). The hindlimbs, in the region of the tibia, are cream in colour anteriorly and greyish brown on the posterior side. The darker colour invades the heel of the cream coloured pes, and the posterior edges of the footpad, unlike in
<emphasis id="B97A8500897FDB49FB697250FA6DA3F7" box="[1235,1436,367,392]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FAAA724EFA6DA3F7" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1296,1436,367,392]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
where the pes is completely cream in colour and
<taxonomicName id="4C0E2291897FDB49FD6172B4FC93A3D3" authorityName="Thomas" authorityYear="1922" box="[731,866,404,429]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FD6172B4FC93A3D3" box="[731,866,404,429]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
where the pes is completely grey in colour. The footpad is hairless posterior to digit I to the heel as in
<taxonomicName id="4C0E2291897FDB49FD577298FC9AA3AF" authorityName="Thomas" authorityYear="1922" box="[749,875,439,464]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FD577298FC9AA3AF" box="[749,875,439,464]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
, while in
<emphasis id="B97A8500897FDB49FC637298FB53A3AF" box="[985,1186,439,464]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FBAA7296FB6EA3AF" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1040,1183,439,464]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
this area is covered in fur. As in the manus, the pes has shorter toes with thicker toepads than
<emphasis id="B97A8500897FDB49FC0972FCFB86A38A" box="[947,1143,476,501]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FC5172FDFB86A38A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1003,1143,476,501]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897FDB49FB1772FCFAC2A38B" box="[1197,1331,476,501]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FB1772FCFADEA38B" authorityName="Thomas" authorityYear="1922" box="[1197,1327,476,501]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="DF71099A897DDB4BFF2D7486FB47A5A4" httpUri="https://zenodo.org/record/1169507/files/figure.png" pageId="13" pageNumber="237" startId="13.[151,250,1959,1981]" targetBox="[177,1418,193,1949]" targetPageId="13">
<paragraph id="8BB15912897DDB4BFF2D7486FB47A5A4" blockId="13.[151,1436,1959,2011]" pageId="13" pageNumber="237">
<emphasis id="B97A8500897DDB4BFF2D7486FEE2A5C2" bold="true" box="[151,275,1959,1981]" pageId="13" pageNumber="237">FIGURE 5.</emphasis>
<taxonomicName id="4C0E2291897DDB4BFEA37486FE2EA5C3" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[281,479,1959,1980]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="13" pageNumber="237" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897DDB4BFEA37486FE2EA5C3" box="[281,479,1959,1980]" italics="true" pageId="13" pageNumber="237">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897DDB4BFE5C7486FDC7A5C2" box="[486,566,1959,1981]" pageId="13" pageNumber="237" rank="species">
<emphasis id="B97A8500897DDB4BFE5C7486FDC7A5C2" bold="true" box="[486,566,1959,1981]" pageId="13" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
, study skins. AC, holotype WAM M571; DF, paratype WAM M574; GI, paratype WAM M572. A, D, and G, dorsal view; B, E, and H, lateral view; C, F, and I, ventral view. Scale = 5cm.
</paragraph>
</caption>
<caption id="DF71099A897EDB48FF2D77A0FC51A6CA" httpUri="https://zenodo.org/record/1169509/files/figure.png" pageId="14" pageNumber="238" startId="14.[151,250,1153,1175]" targetBox="[151,1436,193,1131]" targetPageId="14">
<paragraph id="8BB15912897EDB48FF2D77A0FC51A6CA" blockId="14.[151,1436,1153,1205]" pageId="14" pageNumber="238">
<emphasis id="B97A8500897EDB48FF2D77A0FEE3A6E8" bold="true" box="[151,274,1153,1175]" pageId="14" pageNumber="238">FIGURE 6.</emphasis>
<taxonomicName id="4C0E2291897EDB48FEA277A0FE2FA6E9" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[280,478,1153,1174]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="14" pageNumber="238" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897EDB48FEA277A0FE2FA6E9" box="[280,478,1153,1174]" italics="true" pageId="14" pageNumber="238">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897EDB48FE5977A0FDC2A6E8" box="[483,563,1153,1175]" pageId="14" pageNumber="238" rank="species">
<emphasis id="B97A8500897EDB48FE5977A0FDC2A6E8" bold="true" box="[483,563,1153,1175]" pageId="14" pageNumber="238">sp. nov.</emphasis>
</taxonomicNameLabel>
, skulls. AC and GH, holotype WAM M571; DF and IJ, paratype WAM M577. A, D, G, I, dorsal view; B, E, ventral view; C, F, H, J, lateral view. Scale = 2cm.
</paragraph>
</caption>
<paragraph id="8BB15912897FDB49FF7D72DEFA74A74F" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="239">
The description of the skull and teeth is based on
<typeStatus id="54B5E7B0897FDB49FD407121FCAAA067" box="[762,859,512,536]" pageId="15" pageNumber="239" type="paratype">paratype</typeStatus>
WAM M577 for
<taxonomicName id="4C0E2291897FDB49FB9B72DEFB0DA067" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1057,1276,511,536]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897FDB49FB9B72DEFB0DA067" box="[1057,1276,511,536]" italics="true" pageId="15" pageNumber="239">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897FDB49FABE72DEFAAFA067" box="[1284,1374,511,536]" pageId="15" pageNumber="239" rank="species">
<emphasis id="B97A8500897FDB49FABE72DEFAAFA067" bold="true" box="[1284,1374,511,536]" pageId="15" pageNumber="239">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13354597897FDB49FAD472DEFF57A042" captionStart="FIGURE6" captionStartId="14.[151,250,1153,1175]" captionTargetBox="[151,1436,193,1131]" captionTargetId="figure@14.[151,1436,193,1131]" captionTargetPageId="14" captionText="FIGURE6. Perameles papillonsp.nov., skulls.AC and GH, holotype WAMM571; DF and IJ, paratype WAMM577.A, D, G, I, dorsal view; B, E, ventral view; C, F, H, J, lateral view. Scale = 2cm." httpUri="https://zenodo.org/record/1169509/files/figure.png" pageId="15" pageNumber="239">Fig. 6</figureCitation>
), and describes only features that differ from other
<emphasis id="B97A8500897FDB49FD5F7105FC93A042" box="[741,866,548,573]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FD5F7105FCAFA042" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[741,862,548,573]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">Perameles</taxonomicName>
,
</emphasis>
using WAM M6166 for
<emphasis id="B97A8500897FDB49FBCD7104FAB0A042" box="[1143,1345,548,573]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FB147105FACCA042" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1198,1341,548,573]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
BMNH 43.8.12.21 for
<emphasis id="B97A8500897FDB49FEFC7168FE24A01F" box="[326,469,583,608]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FEFC7168FE21A01F" authorityName="Thomas" authorityYear="1922" box="[326,464,583,608]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
WAM M2629 for
<emphasis id="B97A8500897FDB49FD007168FCB8A01F" box="[698,841,585,608]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FD007168FCB7A01F" authorityName="Spencer" authorityYear="1897" box="[698,838,585,608]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
,
</emphasis>
WAM M16590 for
<taxonomicName id="4C0E2291897FDB49FB867168FB55A01F" authorityName="Gray" authorityYear="1838" box="[1084,1188,585,608]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FB867168FB55A01F" box="[1084,1188,585,608]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
(Suppl.
<figureCitation id="13354597897FDB49FAB77166FAA5A01F" box="[1293,1364,583,608]" captionStart="FIGURE4" captionStartId="10.[151,250,1628,1650]" captionTargetBox="[166,1412,502,1590]" captionTargetId="figure@10.[151,1436,483,1607]" captionTargetPageId="10" captionText="FIGURE4. X-Ygraphsof the tooth length versustooth width, for the upper(AG) and lower(HN) premolars and molars,for Peroryctes raffrayana(squares), Peroryctes broadbenti (crosses), and Lemdubuoryctes aruensis (star)." httpUri="https://zenodo.org/record/1169505/files/figure.png" pageId="15" pageNumber="239">Fig. 4</figureCitation>
), QM JM18575 for
<taxonomicName id="4C0E2291897FDB49FE89714CFE34A0FA" authorityName="Thomas" authorityYear="1923" box="[307,453,620,645]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A8500897FDB49FE89714CFE34A0FA" box="[307,453,620,645]" italics="true" pageId="15" pageNumber="239">P. pallescens</emphasis>
</taxonomicName>
and WAM
<accessionNumber id="945DC4F1897FDB49FDF1714DFD5BA0FB" box="[587,682,620,644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M44421" pageId="15" pageNumber="239" type="EnaNcbi">M44421</accessionNumber>
for
<taxonomicName id="4C0E2291897FDB49FD66714CFCB5A0FB" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[732,836,621,644]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FD66714CFCB5A0FB" box="[732,836,621,644]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9F24E3897FDB49FC38714DFBA5A0FA" box="[898,1108,620,645]" pageId="15" pageNumber="239" refId="ref19545" refString="Travouillon, K. J. (2016 a) Investigating dental variation in Perameles nasuta Geoffroy, 1804, with morphological evidence to raise P. nasuta pallescens Thomas, 1923 to species rank. Zootaxa, 4114, 351 - 392." type="journal article">Travouillon 2016a</bibRefCitation>
), unless specified otherwise. The skull of
<taxonomicName id="4C0E2291897FDB49FE9071B0FE55A0D7" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[298,420,655,680]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897FDB49FE9071B0FE55A0D7" box="[298,420,655,680]" italics="true" pageId="15" pageNumber="239">P. papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897FDB49FE1471AEFDF8A0D7" box="[430,521,655,680]" pageId="15" pageNumber="239" rank="species">
<emphasis id="B97A8500897FDB49FE1471AEFDF8A0D7" bold="true" box="[430,521,655,680]" pageId="15" pageNumber="239">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar in size to
<emphasis id="B97A8500897FDB49FD4F71B0FC4DA0D7" box="[757,956,655,680]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FC8A71AEFC4DA0D7" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[816,956,655,680]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897FDB49FC4C71B0FB8DA0D7" authorityName="Spencer" authorityYear="1897" box="[1014,1148,657,680]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A8500897FDB49FC4C71B0FB8DA0D7" box="[1014,1148,657,680]" italics="true" pageId="15" pageNumber="239">P. eremiana</emphasis>
</taxonomicName>
(much smaller than
<emphasis id="B97A8500897FDB49FAD071B0FE1FA0B2" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FAD071B0FF10A0B3" authorityName="Thomas" authorityYear="1922" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FF567194FEA3A0B3" authorityName="Gray" authorityYear="1838" box="[236,338,693,716]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">P. gunnii</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FEE77194FE1FA0B2" authorityName="Thomas" authorityYear="1923" box="[349,494,692,717]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897FDB49FD997194FD78A0B3" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[547,649,693,716]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FD997194FD78A0B3" box="[547,649,693,716]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
) but it is shorter than
<taxonomicName id="4C0E2291897FDB49FC3A7194FBF5A0B3" authorityName="Spencer" authorityYear="1897" box="[896,1028,693,716]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A8500897FDB49FC3A7194FBF5A0B3" box="[896,1028,693,716]" italics="true" pageId="15" pageNumber="239">P. eremiana</emphasis>
</taxonomicName>
(nasals shorter in length), and much wider than both
<emphasis id="B97A8500897FDB49FEF671F8FDFEA08F" box="[332,527,727,752]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FE3971F6FDFEA08F" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[387,527,727,752]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897FDB49FDFE71F8FD38A08F" authorityName="Spencer" authorityYear="1897" box="[580,713,729,752]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A8500897FDB49FDFE71F8FD38A08F" box="[580,713,729,752]" italics="true" pageId="15" pageNumber="239">P. eremiana</emphasis>
</taxonomicName>
. In dorsal view, the lacrimal crest is highly developed, as seen in
<taxonomicName id="4C0E2291897FDB49FF2D71DCFF0BA16B" authorityName="Gray" authorityYear="1838" box="[151,250,765,788]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FF2D71DCFF0BA16B" box="[151,250,765,788]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
and
<emphasis id="B97A8500897FDB49FE9571DCFE44A16B" box="[303,437,764,789]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FE9571DCFE41A16B" authorityName="Thomas" authorityYear="1922" box="[303,432,764,789]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
and is visible as a distinct posteriorly orientated process in the orbit. The lacrimal crest is less developed in all other species of
<emphasis id="B97A8500897FDB49FD83703EFD46A147" box="[569,695,799,824]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FD83703EFD42A147" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[569,691,799,824]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">Perameles</taxonomicName>
.
</emphasis>
No sagittal crest is present, as in all
<taxonomicName id="4C0E2291897FDB49FBE8703EFB39A147" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1106,1224,799,824]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FBE8703EFB39A147" box="[1106,1224,799,824]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except in males of
<taxonomicName id="4C0E2291897FDB49FF2D7064FEF0A123" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[151,257,837,860]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FF2D7064FEF0A123" box="[151,257,837,860]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
and
<emphasis id="B97A8500897FDB49FE857064FE2AA122" box="[319,475,836,861]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FE857064FE27A122" authorityName="Thomas" authorityYear="1923" box="[319,470,836,861]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
.
</emphasis>
In ventral and lateral view, the alisphenoid tympanic process, or bullae, is highly distinctive in being the most developed (the second largest bullae) within the genus.
<emphasis id="B97A8500897FDB49FBCF7048FACEA1FF" box="[1141,1343,871,896]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FB097046FACEA1FF" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1203,1343,871,896]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
has the smallest bullae, followed by
<emphasis id="B97A8500897FDB49FE5A70ACFC2AA1DA" box="[480,987,908,933]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FE5A70ACFDBBA1DB" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[480,586,909,932]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FDED70ACFD4FA1DB" authorityName="Gray" authorityYear="1838" box="[599,702,909,932]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">P. gunnii</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FD7170ACFCA5A1DB" authorityName="Spencer" authorityYear="1897" box="[715,852,909,932]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FCDB70ACFC2AA1DA" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[865,987,908,933]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">P. papillon</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A249387B897FDB49FC5E70ADFBCEA1DA" box="[996,1087,908,933]" pageId="15" pageNumber="239" rank="species">
<emphasis id="B97A8500897FDB49FC5E70ADFBCEA1DA" bold="true" box="[996,1087,908,933]" pageId="15" pageNumber="239">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<emphasis id="B97A8500897FDB49FBC370ACFAF3A1DB" box="[1145,1282,908,933]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FBC370ACFB0CA1DB" authorityName="Thomas" authorityYear="1922" box="[1145,1277,908,933]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
in increasing order of size. On the palate, accessory fenestrae are present between the incisive fenestrae and the maxillopalatine fenestrae as in all
<taxonomicName id="4C0E2291897FDB49FED070F5FE11A192" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[362,480,980,1005]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FED070F5FE11A192" box="[362,480,980,1005]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except
<taxonomicName id="4C0E2291897FDB49FD8670F4FD55A193" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[572,676,981,1004]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FD8670F4FD55A193" box="[572,676,981,1004]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
and
<emphasis id="B97A8500897FDB49FD6570F4FC89A192" box="[735,888,980,1005]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FD6570F4FC85A192" authorityName="Thomas" authorityYear="1923" box="[735,884,980,1005]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
,
</emphasis>
but they extend anteriorly to the canine, unlike other species of
<taxonomicName id="4C0E2291897FDB49FEF670D6FE32A66F" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[332,451,1015,1040]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FEF670D6FE32A66F" box="[332,451,1015,1040]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except
<taxonomicName id="4C0E2291897FDB49FDA270D8FD67A66F" authorityName="Thomas" authorityYear="1922" box="[536,662,1015,1040]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FDA270D8FD67A66F" box="[536,662,1015,1040]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
where the anterior extension of the accessory fenestrae is posterior to the canines. The maxillopalatine fenestrae also extend further posteriorly than in other
<taxonomicName id="4C0E2291897FDB49FB28773DFAF9A64A" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1170,1288,1052,1077]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FB28773DFAF9A64A" box="[1170,1288,1052,1077]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except
<emphasis id="B97A8500897FDB49FADD773CFF14A627" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FADD773CFF10A627" authorityName="Thomas" authorityYear="1922" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
</emphasis>
from the anterior of the M3 to level with the middle of M3, instead of level with M2. The dividing septa of the maxillopalatine fenestrae are thick anteriorly and posteriorly, thinning centrally, as in
<taxonomicName id="4C0E2291897FDB49FB357745FAA8A603" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[1167,1369,1124,1149]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FB357745FAA8A603" box="[1167,1369,1124,1149]" italics="true" pageId="15" pageNumber="239">Perameles nasuta</emphasis>
</taxonomicName>
(very thin in all other
<taxonomicName id="4C0E2291897FDB49FEF277A6FE4FA6DF" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[328,446,1159,1184]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FEF277A6FE4FA6DF" box="[328,446,1159,1184]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
). Palatine fenestrae are present and large as in
<emphasis id="B97A8500897FDB49FC7077A8FB7CA6DF" box="[970,1165,1159,1184]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FBBB77A6FB7CA6DF" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1025,1165,1159,1184]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
(broken in
<taxonomicName id="4C0E2291897FDB49FAB777A8FA7DA6DF" authorityName="Thomas" authorityYear="1922" box="[1293,1420,1159,1184]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FAB777A8FA7DA6DF" box="[1293,1420,1159,1184]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
). In lateral view the orbitosphenoid is identified as a very reduced bone, as in
<emphasis id="B97A8500897FDB49FC43778CFAF1A6BA" box="[1017,1280,1196,1221]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FC43778CFB93A6BB" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[1017,1122,1197,1220]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FBD5778CFAF1A6BA" authorityName="Thomas" authorityYear="1923" box="[1135,1280,1196,1221]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897FDB49FA82778CFA6DA6BB" authorityName="Gray" authorityYear="1838" box="[1336,1436,1197,1220]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FA82778CFA6DA6BB" box="[1336,1436,1197,1220]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
(much larger in
<emphasis id="B97A8500897FDB49FEF277F0FDFBA697" box="[328,522,1231,1256]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FEC477EEFDFBA697" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[382,522,1231,1256]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897FDB49FDFA77F0FD3BA697" box="[576,714,1233,1256]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FDFA77F0FD36A697" authorityName="Spencer" authorityYear="1897" box="[576,711,1233,1256]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
,
</emphasis>
broken in
<taxonomicName id="4C0E2291897FDB49FCFB77F0FC4EA697" authorityName="Thomas" authorityYear="1922" box="[833,959,1231,1256]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FCFB77F0FC4EA697" box="[833,959,1231,1256]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
). The antorbital fossa is very deep as in all
<taxonomicName id="4C0E2291897FDB49FF2D77D5FEFCA772" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[151,269,1268,1293]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FF2D77D5FEFCA772" box="[151,269,1268,1293]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except in
<taxonomicName id="4C0E2291897FDB49FE3D77D4FE1EA773" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[391,495,1269,1292]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FE3D77D4FE1EA773" box="[391,495,1269,1292]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E2291897FDB49FD9077D4FD4AA772" authorityName="Thomas" authorityYear="1923" box="[554,699,1268,1293]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A8500897FDB49FD9077D4FD4AA772" box="[554,699,1268,1293]" italics="true" pageId="15" pageNumber="239">P. pallescens</emphasis>
</taxonomicName>
. The squamosal epitympanic sinus of
<taxonomicName id="4C0E2291897FDB49FBCF77D4FB01A772" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1141,1264,1268,1293]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A8500897FDB49FBCF77D4FB01A772" box="[1141,1264,1268,1293]" italics="true" pageId="15" pageNumber="239">P. papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B897FDB49FB4377D5FAA4A772" box="[1273,1365,1268,1293]" pageId="15" pageNumber="239" rank="species">
<emphasis id="B97A8500897FDB49FB4377D5FAA4A772" bold="true" box="[1273,1365,1268,1293]" pageId="15" pageNumber="239">sp. nov.</emphasis>
</taxonomicNameLabel>
is the most developed of any
<taxonomicName id="4C0E2291897FDB49FE267636FDE3A74F" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[412,530,1303,1328]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FE267636FDE3A74F" box="[412,530,1303,1328]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
(most visible in
<typeStatus id="54B5E7B0897FDB49FD747639FCDFA74F" box="[718,814,1304,1328]" pageId="15" pageNumber="239" type="paratype">paratype</typeStatus>
WAM M126), being wide with a high posterior wall.
</paragraph>
<paragraph id="8BB15912897FDB56FF7D761DFA61A363" blockId="15.[151,1437,151,2013]" lastBlockId="16.[151,1436,151,285]" lastPageId="16" lastPageNumber="240" pageId="15" pageNumber="239">
The
<date id="FFB07FD2897FDB49FF43761DFEE1A72A" box="[249,272,1340,1365]" pageId="15" pageNumber="239">I5</date>
is pointed and canine-like as in
<emphasis id="B97A8500897FDB49FDC1761CFBB0A72B" box="[635,1089,1340,1365]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FD09761DFCB3A72A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[691,834,1340,1365]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FCF7761CFC3EA72B" authorityName="Thomas" authorityYear="1922" box="[845,975,1340,1365]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FC60761CFBB0A72B" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[986,1089,1341,1364]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897FDB49FBCD761CFAFFA72A" box="[1143,1294,1340,1365]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FBCD761CFAFBA72A" authorityName="Thomas" authorityYear="1923" box="[1143,1290,1340,1365]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
.
</emphasis>
The canines are small with accessory cusps, in both males and females, as in
<emphasis id="B97A8500897FDB49FC3B7640FB2DA707" box="[897,1244,1375,1400]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FC06767EFBBAA707" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[956,1099,1375,1400]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FBE37640FB2DA707" authorityName="Thomas" authorityYear="1922" box="[1113,1244,1375,1400]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897FDB49FAAC7640FA6DA707" authorityName="Spencer" authorityYear="1897" box="[1302,1436,1377,1400]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A8500897FDB49FAAC7640FA6DA707" box="[1302,1436,1377,1400]" italics="true" pageId="15" pageNumber="239">P. eremiana</emphasis>
</taxonomicName>
(males of
<taxonomicName id="4C0E2291897FDB49FEB376A4FE9DA7E3" authorityName="Gray" authorityYear="1838" box="[265,364,1413,1436]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FEB376A4FE9DA7E3" box="[265,364,1413,1436]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FEC676A4FE12A7E3" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[380,483,1413,1436]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FEC676A4FE12A7E3" box="[380,483,1413,1436]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E2291897FDB49FDA176A4FD5DA7E2" authorityName="Thomas" authorityYear="1923" box="[539,684,1412,1437]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A8500897FDB49FDA176A4FD5DA7E2" box="[539,684,1412,1437]" italics="true" pageId="15" pageNumber="239">P. pallescens</emphasis>
</taxonomicName>
have an enlarged canine, and males of
<taxonomicName id="4C0E2291897FDB49FBD676A4FB22A7E3" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[1132,1235,1413,1436]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FBD676A4FB22A7E3" box="[1132,1235,1413,1436]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E2291897FDB49FAB176A4FA6DA7E2" authorityName="Thomas" authorityYear="1923" box="[1291,1436,1412,1437]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A8500897FDB49FAB176A4FA6DA7E2" box="[1291,1436,1412,1437]" italics="true" pageId="15" pageNumber="239">P. pallescens</emphasis>
</taxonomicName>
generally lack accessory cusps). The canines are entirely bordered by the maxilla as in all
<emphasis id="B97A8500897FDB49FB147686FADAA7BF" box="[1198,1323,1447,1472]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FB147686FAD6A7BF" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1198,1319,1447,1472]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">Perameles</taxonomicName>
,
</emphasis>
except
<taxonomicName id="4C0E2291897FDB49FA3D7688FEF1A79B" authorityName="Spencer" authorityYear="1897" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A8500897FDB49FA3D7688FEF1A79B" italics="true" pageId="15" pageNumber="239">P. eremiana</emphasis>
</taxonomicName>
(bordered by premaxilla anteriorly). The diastema between the canine and P1 is long (longer than P1) as in all
<taxonomicName id="4C0E2291897FDB49FF6076CEFEA1A477" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[218,336,1519,1544]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FF6076CEFEA1A477" box="[218,336,1519,1544]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except
<taxonomicName id="4C0E2291897FDB49FE1176D0FDE5A477" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[427,532,1521,1544]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FE1176D0FDE5A477" box="[427,532,1521,1544]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
and
<emphasis id="B97A8500897FDB49FDF776D0FD16A477" box="[589,743,1519,1544]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FDF776D0FD12A477" authorityName="Thomas" authorityYear="1923" box="[589,739,1519,1544]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
.
</emphasis>
The P1 is as long as the P2 as in
<taxonomicName id="4C0E2291897FDB49FBCB76D0FB27A477" authorityName="Gray" authorityYear="1838" box="[1137,1238,1521,1544]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FBCB76D0FB27A477" box="[1137,1238,1521,1544]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FB5F76D0FABFA477" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[1253,1358,1521,1544]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A8500897FDB49FB5F76D0FABFA477" box="[1253,1358,1521,1544]" italics="true" pageId="15" pageNumber="239">P. nasuta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E2291897FDB49FA3D76D0FEFDA452" authorityName="Thomas" authorityYear="1923" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A8500897FDB49FA3D76D0FEFDA452" italics="true" pageId="15" pageNumber="239">P. pallescens</emphasis>
</taxonomicName>
(shorter in
<emphasis id="B97A8500897FDB49FE287534FD91A452" box="[402,608,1556,1581]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FE777535FDADA452" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[461,604,1556,1581]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
longer in
<taxonomicName id="4C0E2291897FDB49FD627534FCAFA453" authorityName="Spencer" authorityYear="1897" box="[728,862,1557,1580]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A8500897FDB49FD627534FCAFA453" box="[728,862,1557,1580]" italics="true" pageId="15" pageNumber="239">P. eremiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E2291897FDB49FC2D7534FBEBA453" authorityName="Thomas" authorityYear="1922" box="[919,1050,1556,1581]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FC2D7534FBEBA453" box="[919,1050,1556,1581]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
), and the P2 is noticeably shorter than the P3, as in
<emphasis id="B97A8500897FDB49FECD7518FD8EA42F" box="[375,639,1591,1616]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FECD7518FDF2A42F" authorityName="Thomas" authorityYear="1922" box="[375,515,1591,1616]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">P. b. notina</taxonomicName>
,
<taxonomicName id="4C0E2291897FDB49FDAE7518FD8EA42F" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[532,639,1593,1616]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897FDB49FD7A7518FCA4A42F" authorityName="Thomas" authorityYear="1923" box="[704,853,1591,1616]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A8500897FDB49FD7A7518FCA4A42F" box="[704,853,1591,1616]" italics="true" pageId="15" pageNumber="239">P. pallescens</emphasis>
</taxonomicName>
(subequal in length in all other
<taxonomicName id="4C0E2291897FDB49FB647516FAA5A42F" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1246,1364,1591,1616]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FB647516FAA5A42F" box="[1246,1364,1591,1616]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
). The development of the lingual shelf of P3 is small as in
<emphasis id="B97A8500897FDB49FD4E757CFC4AA40A" box="[756,955,1628,1653]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FC95757DFC4AA40A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[815,955,1628,1653]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897FDB49FC4E757CFB73A40B" box="[1012,1154,1629,1652]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FC4E757CFB8CA40B" authorityName="Spencer" authorityYear="1897" box="[1012,1149,1629,1652]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
.
</emphasis>
The major cusp of P3 is similar in shape to all
<taxonomicName id="4C0E2291897FDB49FE28755EFDF9A4E7" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[402,520,1663,1688]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FE28755EFDF9A4E7" box="[402,520,1663,1688]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except
<taxonomicName id="4C0E2291897FDB49FDDB75A0FD34A4E7" authorityName="Gray" authorityYear="1838" box="[609,709,1665,1688]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FDDB75A0FD34A4E7" box="[609,709,1665,1688]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
(more conical). The upper molars of
<taxonomicName id="4C0E2291897FDB49FBD775A0FB1FA4E7" authorityName="Thomas" authorityYear="1922" box="[1133,1262,1663,1688]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="subSpecies" species="myosura" subSpecies="notina">
<emphasis id="B97A8500897FDB49FBD775A0FB1FA4E7" box="[1133,1262,1663,1688]" italics="true" pageId="15" pageNumber="239">P. b. notina</emphasis>
</taxonomicName>
are completely worn down and the
<typeStatus id="54B5E7B0897FDB49FEC27585FE2BA4C2" box="[376,474,1700,1725]" pageId="15" pageNumber="239" type="holotype">holotype</typeStatus>
lacks a dentary and therefore these are not used in the comparison following. On the M1, a short stylar crest is present anterior to the metastyle (best seen on juvenile WAM M572) with an associated StE, while StE is a conical cusp with no associated crest in
<emphasis id="B97A8500897FDB49FC9D75CCFC1BA57A" box="[807,1002,1772,1797]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FCE475CDFC1BA57A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[862,1002,1772,1797]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
(best seen in juvenile WAM M16086). StA is variable in size on M12. StB and StC are fused and indistinguishable as in
<emphasis id="B97A8500897FDB49FBC67430FAB9A557" box="[1148,1352,1807,1832]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FB06742EFAB9A557" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1212,1352,1807,1832]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A8500897FDB49FA3D7430FEF6A533" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FA3D7430FEF2A533" authorityName="Spencer" authorityYear="1897" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
.
</emphasis>
The preparacrista reconnects with the postparacrista posteriorly on the M1, as in
<emphasis id="B97A8500897FDB49FB6F7414FA94A533" box="[1237,1381,1845,1868]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FB6F7414FA90A533" authorityName="Spencer" authorityYear="1897" box="[1237,1377,1845,1868]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
.
</emphasis>
The postprotocrista on M1 terminates posterior to the metacone as in
<emphasis id="B97A8500897FDB49FCC47478FBB5A50F" box="[894,1092,1879,1904]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FC027476FBB5A50F" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[952,1092,1879,1904]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E2291897FDB49FBC77478FB12A50F" authorityName="Gray" authorityYear="1838" box="[1149,1251,1881,1904]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FBC77478FB12A50F" box="[1149,1251,1881,1904]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
with no distinct posterior cingulum present (terminates more buccally in all other
<emphasis id="B97A8500897FDB49FCC2745DFC04A5EA" box="[888,1013,1916,1941]" italics="true" pageId="15" pageNumber="239">
<taxonomicName id="4C0E2291897FDB49FCC2745DFC00A5EA" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[888,1009,1916,1941]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">Perameles</taxonomicName>
,
</emphasis>
as a posterior cingulum). On the M2, StB is conical as in all
<taxonomicName id="4C0E2291897FDB49FE2C7481FDFDA5C6" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[406,524,1952,1977]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="genus">
<emphasis id="B97A8500897FDB49FE2C7481FDFDA5C6" box="[406,524,1952,1977]" italics="true" pageId="15" pageNumber="239">Perameles</emphasis>
</taxonomicName>
except
<taxonomicName id="4C0E2291897FDB49FDD97480FD34A5C7" authorityName="Gray" authorityYear="1838" box="[611,709,1953,1976]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FDD97480FD34A5C7" box="[611,709,1953,1976]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
(oval in shape). No stylar crest is present posterior to the conical StD as seen in
<taxonomicName id="4C0E2291897FDB49FEF074E4FE40A5A3" authorityName="Gray" authorityYear="1838" box="[330,433,1989,2012]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A8500897FDB49FEF074E4FE40A5A3" box="[330,433,1989,2012]" italics="true" pageId="15" pageNumber="239">P. gunnii</emphasis>
</taxonomicName>
and
<emphasis id="B97A8500897FDB49FE5474E4FD49A5A2" box="[494,696,1988,2013]" italics="true" pageId="15" pageNumber="239">
P. b.
<taxonomicName id="4C0E2291897FDB49FD9674E5FD49A5A2" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[556,696,1988,2013]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="15" pageNumber="239" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
(small crest present in all other taxa). The postprotocrista ends posterior to the metacone as in all
<emphasis id="B97A85008960DB56FDAF73B6FD63A2CF" box="[533,658,151,176]" italics="true" pageId="16" pageNumber="240">
<taxonomicName id="4C0E22918960DB56FDAF73B6FD7FA2CF" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[533,654,151,176]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="genus">Perameles</taxonomicName>
,
</emphasis>
except
<emphasis id="B97A85008960DB56FD5373B8FCA7A2CF" box="[745,854,153,176]" italics="true" pageId="16" pageNumber="240">
<taxonomicName id="4C0E22918960DB56FD5373B8FCA3A2CF" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[745,850,153,176]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
.
</emphasis>
StE is present as a distinct large cusp as in
<taxonomicName id="4C0E22918960DB56FA8373B8FA6DA2CF" authorityName="Gray" authorityYear="1838" box="[1337,1436,153,176]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A85008960DB56FA8373B8FA6DA2CF" box="[1337,1436,153,176]" italics="true" pageId="16" pageNumber="240">P. gunnii</emphasis>
</taxonomicName>
(it is reduced in all other species). On the M3, the postprotocrista ends posterior to the metacone as in all
<emphasis id="B97A85008960DB56FF2D73FEFEE5A287" box="[151,276,223,248]" italics="true" pageId="16" pageNumber="240">
<taxonomicName id="4C0E22918960DB56FF2D73FEFEE1A287" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[151,272,223,248]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="genus">Perameles</taxonomicName>
,
</emphasis>
except
<emphasis id="B97A85008960DB56FED173C0FE29A287" box="[363,472,225,248]" italics="true" pageId="16" pageNumber="240">
<taxonomicName id="4C0E22918960DB56FED173C0FE25A287" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[363,468,225,248]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
.
</emphasis>
The metaconule is small with only a small shelf separating it from the lingual flank of the metacone, as in
<emphasis id="B97A85008960DB56FECE7224FDC9A362" box="[372,568,260,285]" italics="true" pageId="16" pageNumber="240">
P. b.
<taxonomicName id="4C0E22918960DB56FE167225FDC9A362" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[428,568,260,285]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E22918960DB56FDD47224FD0FA362" authorityName="Thomas" authorityYear="1923" box="[622,766,260,285]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A85008960DB56FDD47224FD0FA362" box="[622,766,260,285]" italics="true" pageId="16" pageNumber="240">P. pallescens</emphasis>
</taxonomicName>
(a larger metaconule and shelf is present in all other taxa).
</paragraph>
<caption id="DF71099A8960DB56FF2D74A9FF23A59F" httpUri="https://zenodo.org/record/1169511/files/figure.png" pageId="16" pageNumber="240" startId="16.[151,250,1928,1950]" targetBox="[151,1436,337,1906]" targetPageId="16">
<paragraph id="8BB159128960DB56FF2D74A9FF23A59F" blockId="16.[151,1436,1928,2016]" pageId="16" pageNumber="240">
<emphasis id="B97A85008960DB56FF2D74A9FEE4A5E1" bold="true" box="[151,277,1928,1950]" pageId="16" pageNumber="240">FIGURE 7.</emphasis>
<taxonomicName id="4C0E22918960DB56FEA474A9FE16A5E2" authority="Travouillon &amp; Phillips, 2018" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[286,487,1928,1949]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="240" phylum="Chordata" rank="species" species="papillon" status="sp. nov.">
<emphasis id="B97A85008960DB56FEA474A9FE16A5E2" box="[286,487,1928,1949]" italics="true" pageId="16" pageNumber="240">Perameles papillon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249387B8960DB56FE5574A9FDB3A5E1" box="[495,578,1928,1950]" pageId="16" pageNumber="240" rank="species">
<emphasis id="B97A85008960DB56FE5574A9FDB3A5E1" bold="true" box="[495,578,1928,1950]" pageId="16" pageNumber="240">sp. nov.</emphasis>
</taxonomicNameLabel>
, upper and lower dentition of paratype WAM M577. AB, upper premolas; CD, upper molars; EF, lower premolars; GH, lower molars. A, C, E and G, occlusal view; B, D, F and H, lateral view. Scale = 2mm.
</paragraph>
</caption>
<paragraph id="8BB159128961DB57FF2D73B9FE56A3F7" blockId="17.[151,1437,151,1005]" pageId="17" pageNumber="241">
StE is absent as in all
<taxonomicName id="4C0E22918961DB57FE3273B6FE0FA2CF" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[392,510,151,176]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FE3273B6FE0FA2CF" box="[392,510,151,176]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except
<emphasis id="B97A85008961DB57FDEE73B8FD4FA2CF" box="[596,702,153,176]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FDEE73B8FD4BA2CF" authorityName="Gray" authorityYear="1838" box="[596,698,153,176]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="gunnii">P. gunnii</taxonomicName>
.
</emphasis>
No anterior cingulum present as in all
<taxonomicName id="4C0E22918961DB57FBD473B6FB15A2CF" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1134,1252,151,176]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FBD473B6FB15A2CF" box="[1134,1252,151,176]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except for some specimens of
<taxonomicName id="4C0E22918961DB57FE8E739CFE36A2AA" authorityName="Thomas" authorityYear="1923" box="[308,455,188,213]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A85008961DB57FE8E739CFE36A2AA" box="[308,455,188,213]" italics="true" pageId="17" pageNumber="241">P. pallescens</emphasis>
</taxonomicName>
and
<emphasis id="B97A85008961DB57FDBA739CFD81A2AB" box="[512,624,189,212]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FDBA739CFD9AA2AB" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[512,619,189,212]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
.
</emphasis>
On the M4, StB is variable, sometimes absent, sometimes present as a small cusp. This feature is variable across
<emphasis id="B97A85008961DB57FDCA73FEFD1FA287" box="[624,750,223,248]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FDCA73FEFD1BA287" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[624,746,223,248]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">Perameles</taxonomicName>
.
</emphasis>
The postparacrista is curved, forming a small centrocrista, as in all
<emphasis id="B97A85008961DB57FF607225FEA6A362" box="[218,343,260,285]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FF607225FEA2A362" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[218,339,260,285]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">Perameles</taxonomicName>
,
</emphasis>
except
<taxonomicName id="4C0E22918961DB57FE0B7224FDE4A363" authorityName="Gray" authorityYear="1838" box="[433,533,261,284]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A85008961DB57FE0B7224FDE4A363" box="[433,533,261,284]" italics="true" pageId="17" pageNumber="241">P. gunnii</emphasis>
</taxonomicName>
and
<emphasis id="B97A85008961DB57FDF47224FCEDA362" box="[590,796,260,285]" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FD337225FCE9A362" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[649,792,260,285]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
,
</emphasis>
where it is straight. The postprotocrista ends anterior to the posterior end of the postparacrista as in all
<taxonomicName id="4C0E22918961DB57FD757206FCB4A33F" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[719,837,295,320]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FD757206FCB4A33F" box="[719,837,295,320]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except
<emphasis id="B97A85008961DB57FC1C7208FBE8A33F" box="[934,1049,297,320]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FC1C7208FBE4A33F" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[934,1045,297,320]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
.
</emphasis>
The metacone is small as in all
<taxonomicName id="4C0E22918961DB57FF2D726DFEFCA31A" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[151,269,332,357]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FF2D726DFEFCA31A" box="[151,269,332,357]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except
<taxonomicName id="4C0E22918961DB57FED9726CFE37A31B" authorityName="Gray" authorityYear="1838" box="[355,454,333,356]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A85008961DB57FED9726CFE37A31B" box="[355,454,333,356]" italics="true" pageId="17" pageNumber="241">P. gunnii</emphasis>
</taxonomicName>
and
<emphasis id="B97A85008961DB57FE41726CFD96A31B" box="[507,615,333,356]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FE41726CFD92A31B" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[507,611,333,356]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="nasuta">P. nasuta</taxonomicName>
,
</emphasis>
which have an enlarged metacone. StC/D is absent (sometimes present in
<taxonomicName id="4C0E22918961DB57FF2D7250FF0BA3F7" authorityName="Gray" authorityYear="1838" box="[151,250,369,392]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="B97A85008961DB57FF2D7250FF0BA3F7" box="[151,250,369,392]" italics="true" pageId="17" pageNumber="241">P. gunnii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E22918961DB57FE8A7250FE66A3F7" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[304,407,369,392]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A85008961DB57FE8A7250FE66A3F7" box="[304,407,369,392]" italics="true" pageId="17" pageNumber="241">P. nasuta</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB159128961DB57FF7D72B5FAA1A193" blockId="17.[151,1437,151,1005]" pageId="17" pageNumber="241">
The lower canine is small and has accessory cuspids as in all
<taxonomicName id="4C0E22918961DB57FC3672B5FBF3A3D2" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[908,1026,404,429]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FC3672B5FBF3A3D2" box="[908,1026,404,429]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except in males of
<taxonomicName id="4C0E22918961DB57FB5F72B4FABFA3D3" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[1253,1358,405,428]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A85008961DB57FB5F72B4FABFA3D3" box="[1253,1358,405,428]" italics="true" pageId="17" pageNumber="241">P. nasuta</emphasis>
</taxonomicName>
and
<emphasis id="B97A85008961DB57FA3D72B4FEE3A3AF" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FA3D72B4FEFEA3AF" authorityName="Thomas" authorityYear="1923" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
,
</emphasis>
which have enlarged unicuspid canines. The diastema between the canine and p1 is shorter than the length of p1, as in
<taxonomicName id="4C0E22918961DB57FECE72FCFDF6A38A" authorityName="Thomas" authorityYear="1923" box="[372,519,476,501]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="B97A85008961DB57FECE72FCFDF6A38A" box="[372,519,476,501]" italics="true" pageId="17" pageNumber="241">P. pallescens</emphasis>
</taxonomicName>
(longer that p1 length in all other taxa). The p1 is as long as the p2 as in
<emphasis id="B97A85008961DB57FAD372FCFED2A067" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FF2D72DEFED2A067" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,291,511,536]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E22918961DB57FEE27120FE2DA067" authorityName="Spencer" authorityYear="1897" box="[344,476,513,536]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A85008961DB57FEE27120FE2DA067" box="[344,476,513,536]" italics="true" pageId="17" pageNumber="241">P. eremiana</emphasis>
</taxonomicName>
(it is shorter in all other taxa). The p2 is longer than the p3 as in
<emphasis id="B97A85008961DB57FB117120FA9CA067" box="[1195,1389,511,536]" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FB5B72DEFA9CA067" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[1249,1389,511,536]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E22918961DB57FF2D7104FEEEA043" authorityName="Spencer" authorityYear="1897" box="[151,287,549,572]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A85008961DB57FF2D7104FEEEA043" box="[151,287,549,572]" italics="true" pageId="17" pageNumber="241">P. eremiana</emphasis>
</taxonomicName>
(subequal in all other taxa). No cuspid is present on the hypoflexid of the lower molars as in all
<taxonomicName id="4C0E22918961DB57FF2D7166FEFCA01F" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[151,269,583,608]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FF2D7166FEFCA01F" box="[151,269,583,608]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except
<taxonomicName id="4C0E22918961DB57FECB7168FE2FA01F" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[369,478,585,608]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A85008961DB57FECB7168FE2FA01F" box="[369,478,585,608]" italics="true" pageId="17" pageNumber="241">P. nasuta</emphasis>
</taxonomicName>
and
<emphasis id="B97A85008961DB57FD987168FD45A01F" box="[546,692,585,608]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FD987168FD41A01F" authorityName="Spencer" authorityYear="1897" box="[546,688,585,608]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
.
</emphasis>
On the m1, the paraconid-metaconid distance is equal to the metaconid-protoconid distance as in all
<taxonomicName id="4C0E22918961DB57FDE4714DFD25A0FA" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[606,724,620,645]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FDE4714DFD25A0FA" box="[606,724,620,645]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except
<emphasis id="B97A85008961DB57FC94714CFC36A0FA" box="[814,967,620,645]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FC94714CFC32A0FA" authorityName="Thomas" authorityYear="1923" box="[814,963,620,645]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
,
</emphasis>
which has a longer paraconid-metaconid distance. The posthypocristid is oblique to the tooth row and connects to the hypoconulid as in all
<taxonomicName id="4C0E22918961DB57FA9C71AEFA6DA0D7" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1318,1436,655,680]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FA9C71AEFA6DA0D7" box="[1318,1436,655,680]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
(sometimes perpendicular in
<emphasis id="B97A85008961DB57FE5A7194FD54A0B2" box="[480,677,692,717]" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FDA37195FD54A0B2" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[537,677,692,717]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and connects to entoconid). On the m2, the posthypocristid is also oblique and connects to a large hypoconulid (this feature varies across
<taxonomicName id="4C0E22918961DB57FC7F71F6FBCAA08F" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[965,1083,727,752]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FC7F71F6FBCAA08F" box="[965,1083,727,752]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
). On the m3, the hypoconid is buccal to the protoconid as in all
<taxonomicName id="4C0E22918961DB57FDB571DDFD74A16A" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[527,645,764,789]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FDB571DDFD74A16A" box="[527,645,764,789]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except
<emphasis id="B97A85008961DB57FD6471DCFC53A16A" box="[734,930,764,789]" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FCAC71DDFC53A16A" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[790,930,764,789]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and some specimens of
<emphasis id="B97A85008961DB57FB0271DCFAA1A16A" box="[1208,1360,764,789]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FB0271DCFABDA16A" authorityName="Thomas" authorityYear="1923" box="[1208,1356,764,789]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
,
</emphasis>
which have the two cuspid level with one another. The posthypocristid connects to the entoconid as in all
<taxonomicName id="4C0E22918961DB57FA9C703EFA6DA147" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1318,1436,799,824]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FA9C703EFA6DA147" box="[1318,1436,799,824]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except some specimens of
<taxonomicName id="4C0E22918961DB57FE767064FDC4A123" authorityName="É. Geoffroy Saint-Hilaire" authorityYear="1804" box="[460,565,837,860]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B97A85008961DB57FE767064FDC4A123" box="[460,565,837,860]" italics="true" pageId="17" pageNumber="241">P. nasuta</emphasis>
</taxonomicName>
and
<emphasis id="B97A85008961DB57FDCA7064FCFBA122" box="[624,778,836,861]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FDCA7064FCF7A122" authorityName="Thomas" authorityYear="1923" box="[624,774,836,861]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
,
</emphasis>
and the hypoconid is either absent or reduced as in
<emphasis id="B97A85008961DB57FAD37064FED2A1FF" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FF2D7046FED2A1FF" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,291,871,896]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A85008961DB57FED87048FE03A1FF" box="[354,498,873,896]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FED87048FE1FA1FF" authorityName="Spencer" authorityYear="1897" box="[354,494,873,896]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
.
</emphasis>
On the m4, the talonid is reduced but not as reduced as in
<emphasis id="B97A85008961DB57FB7A7048FED2A1DA" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FB7A7048FAA6A1FF" authorityName="Thomas" authorityYear="1923" box="[1216,1367,871,896]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="pallescens">P. pallescens</taxonomicName>
, P. b.
<taxonomicName id="4C0E22918961DB57FF2D70ADFED2A1DA" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[151,291,908,933]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A85008961DB57FEE670ACFE18A1DB" box="[348,489,909,932]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FEE670ACFE14A1DB" authorityName="Spencer" authorityYear="1897" box="[348,485,909,932]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
.
</emphasis>
The posthypocristid is perpendicular to the tooth row as in all
<taxonomicName id="4C0E22918961DB57FB0D70ADFADCA1DA" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[1207,1325,908,933]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">
<emphasis id="B97A85008961DB57FB0D70ADFADCA1DA" box="[1207,1325,908,933]" italics="true" pageId="17" pageNumber="241">Perameles</emphasis>
</taxonomicName>
except
<emphasis id="B97A85008961DB57FA3D70ACFF17A1B7" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FA3D70ACFF13A1B7" authorityName="Gray" authorityYear="1838" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="gunnii">P. gunnii</taxonomicName>
.
</emphasis>
The entoconid is large as in all
<emphasis id="B97A85008961DB57FDF0708EFD36A1B7" box="[586,711,943,968]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FDF0708EFD32A1B7" authorityName="Travouillon &amp; Phillips" authorityYear="2018" box="[586,707,943,968]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="genus">Perameles</taxonomicName>
,
</emphasis>
except
<emphasis id="B97A85008961DB57FCA77090FC10A1B7" box="[797,993,943,968]" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FCEF708EFC10A1B7" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[853,993,943,968]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<emphasis id="B97A85008961DB57FBAD7090FB52A1B7" box="[1047,1187,945,968]" italics="true" pageId="17" pageNumber="241">
<taxonomicName id="4C0E22918961DB57FBAD7090FB6FA1B7" authorityName="Spencer" authorityYear="1897" box="[1047,1182,945,968]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="eremiana">P. eremiana</taxonomicName>
.
</emphasis>
The buccal shelf ends at the buccal side of the hypoconid as in
<emphasis id="B97A85008961DB57FDE570F4FCD3A192" box="[607,802,980,1005]" italics="true" pageId="17" pageNumber="241">
P. b.
<taxonomicName id="4C0E22918961DB57FD2C70F5FCD3A192" baseAuthorityName="Quoy &amp; Gaymard" baseAuthorityYear="1824" box="[662,802,980,1005]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="bougainville">bougainville</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0E22918961DB57FCE370F4FC2CA193" authorityName="Spencer" authorityYear="1897" box="[857,989,981,1004]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="17" pageNumber="241" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B97A85008961DB57FCE370F4FC2CA193" box="[857,989,981,1004]" italics="true" pageId="17" pageNumber="241">P. eremiana</emphasis>
</taxonomicName>
(buccally reduced in other taxa).
</paragraph>
</subSubSection>
</treatment>
</document>