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<document id="84862F65E045E6AFD19032CC3D8A13BF" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFCDE53E50C686BB67EDFEF5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) morulus Assing 2017, spec. nov." docType="treatment" docUuid="6F52A4A8-EA4C-472C-AEA3-C3BF6A2C1795" docUuidSource="ZooBank" docVersion="14" lastPageNumber="129" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="129" updateTime="1699471407760" updateUser="ExternalLinkService">
<mods:mods id="231CFE62C1525A95F4F320BBD9E15E30" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CEF1A04A62BC355929D74DE8B0253D7A">
<mods:title id="8399C80A960714331AF115457F172164">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
<mods:name id="6447BDBE01AE9DB292E2FFB0C1738DD5" type="personal">
<mods:role id="4F460CF0A8A3EC81D0826B6B2C9ED293">
<mods:roleTerm id="58A46A3400B82AF67E88C52F2BA2AC19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F08C3F247BFE7CD9FC16A53F15A327C">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="0BF653812F7A8FAF032A00D6691052BC">text</mods:typeOfResource>
<mods:relatedItem id="77134CEB28A182891D9A2F1C40FB8DFA" type="host">
<mods:titleInfo id="D20B7CEB8D9F6EBB78D722CB8647E16D">
<mods:title id="9939BE0FADC9F7A8423D4317CEA4890D">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="0347AF0C422B3042E553C04DB786C653">
<mods:date id="70477D9D253D1D7408F86F95978EF38B">2017</mods:date>
<mods:detail id="E704EA913F0A47ABB80187229ACAF380" type="pubDate">
<mods:number id="D59AE4DB77A25DD0F8745C4626BA7C94">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="28883ABE29E60F7396B53A2B4F5AD592" type="volume">
<mods:number id="31C8E97201098EF97457C52C5A9759AD">67</mods:number>
</mods:detail>
<mods:detail id="C7AC84D8633FA00E3C6323FBCBDB8096" type="issue">
<mods:number id="797C0A198C92B1B3F946552335BE3696">1</mods:number>
</mods:detail>
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<mods:start id="4766FB91ABC6D9EAA45371FD83A702D0">117</mods:start>
<mods:end id="9260CBDAF00280D82F99FEFB6E173737">192</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="A45AD2F8E2BCC6CEA6C387335A6271E0">journal article</mods:classification>
<mods:identifier id="3125F737462FA276FF79EC791BBF1E82" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="4E93AA7261FEEF5B87489C27C25B4338" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
<mods:identifier id="E6BA51F6E26A8A1565A4792012F52B39" type="GBIF-Dataset">39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43</mods:identifier>
<mods:identifier id="9114FD244732085A518F42FB0D1AA6DE" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="C0BA28EF2F1D09B7CB62C0014C3EBE61" type="Zenodo-Dep">5742363</mods:identifier>
<mods:identifier id="FC31377F9D8E48D1153B2D02A40C53C5" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03A787BAFFCDE53E50C686BB67EDFEF5" ID-DOI="http://doi.org/10.5281/zenodo.5888457" ID-GBIF-Taxon="192498160" ID-Zenodo-Dep="5888457" LSID="urn:lsid:zoobank.org:act:6F52A4A8-EA4C-472C-AEA3-C3BF6A2C1795" httpUri="http://treatment.plazi.org/id/03A787BAFFCDE53E50C686BB67EDFEF5" lastPageNumber="129" pageId="12" pageNumber="129">
<subSubSection id="C3146527FFCDE53E50C686BB6126FE7A" box="[227,557,435,462]" pageId="12" pageNumber="129" type="nomenclature">
<paragraph id="8BB136ACFFCDE53E50C686BB6126FE7A" blockId="12.[227,753,435,527]" box="[227,557,435,462]" pageId="12" pageNumber="129">
<heading id="D0F981C0FFCDE53E50C686BB6126FE7A" box="[227,557,435,462]" fontSize="12" level="2" pageId="12" pageNumber="129" reason="2">
<taxonomicName id="4C0E4D2FFFCDE53E50C686BB62B6FE7A" ID-CoL="7GHYG" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[227,445,435,462]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="morulus" status="sp. nov." subGenus="Zyras">
<emphasis id="B97AEABEFFCDE53E50C686BB6212FE7A" box="[227,281,435,462]" italics="true" pageId="12" pageNumber="129">Zyras</emphasis>
(
<emphasis id="B97AEABEFFCDE53E510286BB6256FE7A" box="[295,349,435,462]" italics="true" pageId="12" pageNumber="129">Zyras</emphasis>
)
<emphasis id="B97AEABEFFCDE53E514F86BB62B6FE7A" box="[362,445,435,462]" italics="true" pageId="12" pageNumber="129">morulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFCDE53E51E186BB6126FE7A" box="[452,557,435,462]" pageId="12" pageNumber="129" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFCDE53E50C686D361FAFE5B" box="[227,753,475,495]" pageId="12" pageNumber="129" type="description">
<paragraph id="8BB136ACFFCDE53E50C686D361FAFE5B" blockId="12.[227,753,435,527]" box="[227,753,475,495]" pageId="12" pageNumber="129">
<uri id="FF9F3AAEFFCDE53E50C686D361FAFE5B" box="[227,753,475,495]" pageId="12" pageNumber="129">
urn: lsid:zoobank.org:act:
<uuid id="FFA80C79FFCDE53E51B786D361FAFE5B" box="[402,753,475,495]" pageId="12" pageNumber="129">6F52A4A8-EA4C-472C-AEA3-C3BF6A2C1795</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFCDE53E50C686F36286FDBB" box="[227,397,507,527]" pageId="12" pageNumber="129" type="description">
<paragraph id="8BB136ACFFCDE53E50C686F36286FDBB" blockId="12.[227,753,435,527]" box="[227,397,507,527]" pageId="12" pageNumber="129">
(
<figureCitation id="13352A29FFCDE53E50C286F3625BFDBB" box="[231,336,507,527]" captionStart="Figs 260–277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="12" pageNumber="129">Figs 260–265</figureCitation>
,
<figureCitation id="13352A29FFCDE53E517286F36282FDBB" box="[343,393,507,527]" captionStart="Map 4" captionStartId="65.[154,199,794,815]" captionTargetBox="[151,1398,230,785]" captionTargetId="figure-63@65.[151,1398,230,785]" captionTargetPageId="65" captionText="Map 4: Revised distributions of Zyras condignus (black circles) and Z. morulus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742401" httpUri="https://zenodo.org/record/5742401/files/figure.png" pageId="12" pageNumber="129">Map 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFCDE53E5098852161F1FC75" pageId="12" pageNumber="129" type="materials_examined">
<paragraph id="8BB136ACFFCDE53E5098852161F1FC75" blockId="12.[189,794,553,962]" pageId="12" pageNumber="129">
<emphasis id="B97AEABEFFCDE53E50988521625FFDF5" bold="true" box="[189,340,553,577]" pageId="12" pageNumber="129">Type material</emphasis>
:
<materialsCitation id="3B663CF1FFCDE53E514485216167FD15" ID-GBIF-Occurrence="3450162306" collectingDate="1998-07-13" collectionCode="SMTD" collectorName="Jager" country="Nepal" county="Marpha" determinerName="V. Assing" elevation="2321" location="Dhaulagiri-Himal" municipality="Yak Kharka" pageId="12" pageNumber="129" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B5880EFFCDE53E5144852162CFFDF5" box="[353,452,553,577]" pageId="12" pageNumber="129" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFCDE53E51EE852262EDFDF5" box="[459,486,554,577]" pageId="12" pageNumber="129" type="generic">♂:</specimenCount>
“
<collectingCountry id="F319763CFFCDE53E51D385216141FDF5" box="[502,586,553,577]" name="Nepal" pageId="12" pageNumber="129">NEPAL</collectingCountry>
,
<location id="8ED16077FFCDE53E52718521601DFDF5" LSID="urn:lsid:plazi:treatment:03A787BAFFCDE53E50C686BB67EDFEF5:8ED16077FFCDE53E52718521601DFDF5" box="[596,790,553,577]" country="Nepal" county="Marpha" municipality="Yak Kharka" name="Dhaulagiri-Himal" pageId="12" pageNumber="129">Dhaulagiri-Himal</location>
,
<location id="8ED16077FFCDE53E50988541626DFDD5" LSID="urn:lsid:plazi:treatment:03A787BAFFCDE53E50C686BB67EDFEF5:8ED16077FFCDE53E50988541626DFDD5" box="[189,358,585,609]" country="Nepal" county="Marpha" municipality="Yak Kharka" name="Kali Gand. vall." pageId="12" pageNumber="129">Kali Gand. vall.</location>
,
<location id="8ED16077FFCDE53E5155854162E6FDD5" LSID="urn:lsid:plazi:treatment:03A787BAFFCDE53E50C686BB67EDFEF5:8ED16077FFCDE53E5155854162E6FDD5" box="[368,493,585,609]" country="Nepal" county="Marpha" municipality="Yak Kharka" name="Yak Kharka" pageId="12" pageNumber="129">Yak Kharka</location>
(
<location id="8ED16077FFCDE53E51DE85416180FDD5" LSID="urn:lsid:plazi:treatment:03A787BAFFCDE53E50C686BB67EDFEF5:8ED16077FFCDE53E51DE85416180FDD5" box="[507,651,585,609]" country="Nepal" county="Marpha" municipality="Yak Kharka" name="upp. Marpha" pageId="12" pageNumber="129">upp. Marpha</location>
),
<quantity id="4CF69B49FFCDE53E52B88541601DFDD5" box="[669,790,585,609]" metricMagnitude="3" metricUnit="m" metricValue="2.3205" metricValueMax="4.6" metricValueMin="0.040999999999999995" pageId="12" pageNumber="129" unit="m" value="2320.5" valueMax="4600.0" valueMin="41.0">
<elevation id="0023D19FFFCDE53E52B88541601DFDD5" box="[669,790,585,609]" metricMagnitude="3" metricUnit="m" metricValue="2.3205" metricValueMax="4.6" metricValueMin="0.040999999999999995" pageId="12" pageNumber="129" unit="m" value="2320.5" valueMax="4600.0" valueMin="41.0">41–4600 m</elevation>
</quantity>
, 12./
<date id="FFB0106CFFCDE53E50C18561626EFD35" box="[228,357,617,641]" pageId="12" pageNumber="129" value="1998-07-13">
<collectingDate id="EFF4E984FFCDE53E50C18561626EFD35" box="[228,357,617,641]" pageId="12" pageNumber="129" value="1998-07-13">13.VII.1998</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFCDE53E5181856262D2FD36" box="[420,473,618,642]" pageId="12" pageNumber="129">Jäger</collectorName>
/ Holotypus ♂
<taxonomicName id="4C0E4D2FFFCDE53E52B6856263D0FD15" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="morulus" status="sp. nov.">
<emphasis id="B97AEABEFFCDE53E52B6856263D0FD15" italics="true" pageId="12" pageNumber="129">Zyras morulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFCDE53E50C185826217FD15" box="[228,284,650,673]" pageId="12" pageNumber="129" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E6DDA68FFCDE53E517E858262B6FD15" box="[347,445,649,673]" pageId="12" pageNumber="129">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFCDE53E52338581616FFD15" box="[534,612,649,673]" collectionName="Germany, Dresden, Museum fuer Tierkunde" pageId="12" pageNumber="129">SMTD</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFCDE53E525C85826117FD75" ID-GBIF-Occurrence="3455141330" collectingDate="1997-06-10" collectionCode="SMTD" collectorName="Jager" country="Nepal" county="Marpha" determinerName="V. Assing" elevation="2321" location="Annapurna mts." municipality="Telbrung Danda" pageId="12" pageNumber="129" specimenCount="1" typeStatus="paratype">
<typeStatus id="54B5880EFFCDE53E525C858261E8FD16" box="[633,739,650,674]" pageId="12" pageNumber="129" type="paratype">Paratypes</typeStatus>
: 1 ♂: same data as holotype (cAss);
</materialsCitation>
<materialsCitation id="3B663CF1FFCDE53E520985A16212FC95" ID-GBIF-Occurrence="3455141319" collectingDate="1997-06-10" collectionCode="SMTD" collectorName="Jager" country="Nepal" county="Marpha" determinerName="V. Assing" elevation="2321" location="Annapurna mts." municipality="Telbrung Danda" pageId="12" pageNumber="129" specimenCount="1" typeStatus="paratype">
1 ♂ [slightly teneral] “
<collectingCountry id="F319763CFFCDE53E50E385C16213FD55" box="[198,280,713,737]" name="Nepal" pageId="12" pageNumber="129">NEPAL</collectingCountry>
HIMALAYA, SE
<location id="8ED16077FFCDE53E51C885C161ACFD55" LSID="urn:lsid:plazi:treatment:03A787BAFFCDE53E50C686BB67EDFEF5:8ED16077FFCDE53E51C885C161ACFD55" box="[493,679,713,737]" country="Nepal" county="Marpha" municipality="Telbrung Danda" name="Annapurna mts." pageId="12" pageNumber="129">Annapurna mts.</location>
,
<collectingMunicipality id="6BD5ACD6FFCDE53E529F85C1620EFCB5" pageId="12" pageNumber="129">Telbrung Danda</collectingMunicipality>
/ Abies-Rhododendr.-forest,
<date id="FFB0106CFFCDE53E521E85E161BAFCB5" box="[571,689,745,769]" pageId="12" pageNumber="129" value="1997-06-10">
<collectingDate id="EFF4E984FFCDE53E521E85E161BAFCB5" box="[571,689,745,769]" pageId="12" pageNumber="129" value="1997-06-10">10.VI.1997</collectingDate>
</date>
, lg. Jäger” (
<collectionCode id="ED1FAE69FFCDE53E50E184016219FC95" box="[196,274,777,801]" collectionName="Germany, Dresden, Museum fuer Tierkunde" pageId="12" pageNumber="129">SMTD</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFCDE53E510184016212FC35" ID-GBIF-Occurrence="3450162402" collectingDate="1998-06-24" collectionCode="SMTD" collectorName="Ankauf A. Dobbertin" country="Nepal" elevation="1716" location="Myagdi distr." municipality="Ruyachaur Duri" pageId="12" pageNumber="129" specimenCount="1" typeStatus="paratype">
1 ♀: “
<collectingCountry id="F319763CFFCDE53E517B840162B9FC95" box="[350,434,777,801]" name="Nepal" pageId="12" pageNumber="129">NEPAL</collectingCountry>
,
<location id="8ED16077FFCDE53E519984016141FC95" LSID="urn:lsid:plazi:treatment:03A787BAFFCDE53E50C686BB67EDFEF5:8ED16077FFCDE53E519984016141FC95" box="[444,586,777,801]" country="Nepal" municipality="Ruyachaur Duri" name="Myagdi distr." pageId="12" pageNumber="129">Myagdi distr.</location>
, S-slope
<collectingMunicipality id="6BD5ACD6FFCDE53E5282840163F9FCF5" pageId="12" pageNumber="129">Ruyachaur Duri</collectingMunicipality>
,
<quantity id="4CF69B49FFCDE53E50DB8421627CFCF5" box="[254,375,809,833]" metricMagnitude="3" metricUnit="m" metricValue="1.7165" metricValueMax="3.4" metricValueMin="0.032999999999999995" pageId="12" pageNumber="129" unit="m" value="1716.5" valueMax="3400.0" valueMin="33.0">
<elevation id="0023D19FFFCDE53E50DB8421627CFCF5" box="[254,375,809,833]" metricMagnitude="3" metricUnit="m" metricValue="1.7165" metricValueMax="3.4" metricValueMin="0.032999999999999995" pageId="12" pageNumber="129" unit="m" value="1716.5" valueMax="3400.0" valueMin="33.0">33–3400 m</elevation>
</quantity>
,
<date id="FFB0106CFFCDE53E51A6842162F2FCF5" box="[387,505,809,833]" pageId="12" pageNumber="129" value="1998-06-24">
<collectingDate id="EFF4E984FFCDE53E51A6842162F2FCF5" box="[387,505,809,833]" pageId="12" pageNumber="129" value="1998-06-24">24.VI.1998</collectingDate>
</date>
, Berndt/Schmidt /
<collectorName id="26FB537AFFCDE53E52EF84216246FCD5" pageId="12" pageNumber="129">Ankauf A. Dobbertin</collectorName>
, Rostock, 2001, Museum Dresden” (
<collectionCode id="ED1FAE69FFCDE53E50E184616219FC35" box="[196,274,873,897]" collectionName="Germany, Dresden, Museum fuer Tierkunde" pageId="12" pageNumber="129">SMTD</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFCDE53E5103846161FDFC75" ID-GBIF-Occurrence="3450162381" collectionCode="DEI, SDEI" collectorName="Frabrizi &amp; Ankauf J. Schmidt" country="Nepal" elevation="3900" location="Taunja Danda" municipality="Annapurna Mts." pageId="12" pageNumber="129" specimenCount="1" typeStatus="paratype">
1 ♀: “S Lamjun Himal, E-slope
<location id="8ED16077FFCDE53E52598461601DFC35" LSID="urn:lsid:plazi:treatment:03A787BAFFCDE53E50C686BB67EDFEF5:8ED16077FFCDE53E52598461601DFC35" box="[636,790,873,897]" country="Nepal" municipality="Annapurna Mts." name="Taunja Danda" pageId="12" pageNumber="129">Taunja Danda</location>
,
<quantity id="4CF69B49FFCDE53E509884816205FC15" box="[189,270,905,929]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="12" pageNumber="129" unit="m" value="3900.0">
<elevation id="0023D19FFFCDE53E509884816205FC15" box="[189,270,905,929]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="12" pageNumber="129" unit="m" value="3900.0">3900 m</elevation>
</quantity>
,
<date id="FFB0106CFFCDE53E513F8481626DFC15" box="[282,358,905,929]" pageId="12" pageNumber="129" value="95-11-08">11.8.95</date>
/
<collectingCountry id="F319763CFFCDE53E5158848162C6FC15" box="[381,461,905,929]" name="Nepal" pageId="12" pageNumber="129">NEPAL</collectingCountry>
<collectingMunicipality id="6BD5ACD6FFCDE53E51F084816182FC15" box="[469,649,905,929]" pageId="12" pageNumber="129">Annapurna Mts.</collectingMunicipality>
, leg.
<collectorName id="26FB537AFFCDE53E52E58481601DFC15" box="[704,790,905,929]" pageId="12" pageNumber="129">Frabrizi</collectorName>
, Jäger, Schmidt /
<collectionCode id="ED1FAE69FFCDE53E514E84A2629CFC75" box="[363,407,938,961]" collectionName="Germany, Muencheberg, Deutsches Entomologisches Institut im ZALF" pageId="12" pageNumber="129">DEI</collectionCode>
<collectorName id="26FB537AFFCDE53E51B984A16154FC75" box="[412,607,937,961]" pageId="12" pageNumber="129">Ankauf J. Schmidt</collectorName>
1999” (
<collectionCode id="ED1FAE69FFCDE53E529584A161E5FC75" box="[688,750,937,961]" collectionName="SDEI" pageId="12" pageNumber="129">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFCDE53E509884E16235FBF5" pageId="12" pageNumber="129" type="etymology">
<paragraph id="8BB136ACFFCDE53E509884E16235FBF5" blockId="12.[189,794,1001,1089]" pageId="12" pageNumber="129">
<emphasis id="B97AEABEFFCDE53E509884E1623FFBB5" bold="true" box="[189,308,1001,1025]" pageId="12" pageNumber="129">Etymology</emphasis>
: The specific epithet (Latin, adjective: black) alludes to the nearly completely black coloration of the whole body.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFCDE53E50988361617EF815" pageId="12" pageNumber="129" type="description">
<paragraph id="8BB136ACFFCDE53E50988361611DFAB5" blockId="12.[189,794,1129,1665]" pageId="12" pageNumber="129">
<emphasis id="B97AEABEFFCDE53E509883616234FB35" bold="true" box="[189,319,1129,1153]" pageId="12" pageNumber="129">Description</emphasis>
: Body length 7.0–
<quantity id="4CF69B49FFCDE53E52368361616DFB35" box="[531,614,1129,1153]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="129" unit="mm" value="8.5">8.5 mm</quantity>
; length of forebody
<quantity id="4CF69B49FFCDE53E51258381628AFB15" box="[256,385,1161,1185]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="12" pageNumber="129" unit="mm" value="3.45" valueMax="3.6" valueMin="3.3">3.3–3.6 mm</quantity>
. Coloration: body completely black, except for the reddish to dark-brown tarsi, the reddish apical maxillary palpomere, and sometimes the dark reddish-brown antennomere XI.
</paragraph>
<paragraph id="8BB136ACFFCDE53E50988201625CFA75" blockId="12.[189,794,1129,1665]" pageId="12" pageNumber="129">
Elytra with punctation not subject to sexual dimorphism, dense, but near scutellum not extremely dense (similar to the condition in female
<taxonomicName id="4C0E4D2FFFCDE53E519382426124FAD5" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[438,559,1353,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFCDE53E519382426124FAD5" box="[438,559,1353,1377]" italics="true" pageId="12" pageNumber="129">Z. pindarae</emphasis>
</taxonomicName>
). Abdominal segment VII on whole surface with sparse non-setiferous micropunctation. Other external characters (
<figureCitation id="13352A29FFCDE53E5241828161FEFA15" box="[612,757,1417,1441]" captionStart="Figs 260–277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="12" pageNumber="129">Figs 260–262</figureCitation>
) as in
<taxonomicName id="4C0E4D2FFFCDE53E50FC82A26259FA75" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[217,338,1449,1473]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFCDE53E50FC82A26259FA75" box="[217,338,1449,1473]" italics="true" pageId="12" pageNumber="129">Z. pindarae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB136ACFFCDE53E509882C26139F9D5" blockId="12.[189,794,1129,1665]" pageId="12" pageNumber="129">
♂: shapes and chaetotaxy of tergite and sternite VIII as in
<taxonomicName id="4C0E4D2FFFCDE53E50C482E26251F9B5" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[225,346,1513,1537]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFCDE53E50C482E26251F9B5" box="[225,346,1513,1537]" italics="true" pageId="12" pageNumber="129">Z. pindarae</emphasis>
</taxonomicName>
; median lobe of aedeagus 0.9–1.0 mm long and shaped as in
<figureCitation id="13352A29FFCDE53E518881016137F995" box="[429,572,1545,1569]" captionStart="Figs 260–277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="12" pageNumber="129">Figs 263–264</figureCitation>
; paramere (
<figureCitation id="13352A29FFCDE53E529E81016019F995" box="[699,786,1545,1569]" captionStart="Figs 260–277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="12" pageNumber="129">Fig. 265</figureCitation>
)
<quantity id="4CF69B49FFCDE53E509881216204F9F5" box="[189,271,1577,1601]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="129" unit="mm" value="1.3">1.3 mm</quantity>
long, much longer than median lobe, and with rather long and slender apical lobe.
</paragraph>
<paragraph id="8BB136ACFFCDE53E50988162619EF935" blockId="12.[189,794,1129,1665]" box="[189,661,1641,1665]" pageId="12" pageNumber="129">
♀: tergite and sternite VIII as in
<taxonomicName id="4C0E4D2FFFCDE53E52328162619BF935" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[535,656,1641,1665]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFCDE53E52328162619BF935" box="[535,656,1641,1665]" italics="true" pageId="12" pageNumber="129">Z. pindarae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB136ACFFCDE53E509881A1617EF815" blockId="12.[189,794,1705,1953]" pageId="12" pageNumber="129">
<emphasis id="B97AEABEFFCDE53E509881A1629DF975" bold="true" box="[189,406,1705,1729]" pageId="12" pageNumber="129">Comparative notes</emphasis>
:
<taxonomicName id="4C0E4D2FFFCDE53E518E81A26140F975" authorityName="Assing" authorityYear="2017" box="[427,587,1705,1729]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="morulus">
<emphasis id="B97AEABEFFCDE53E518E81A26140F975" box="[427,587,1705,1729]" italics="true" pageId="12" pageNumber="129">Zyras morulus</emphasis>
</taxonomicName>
is readily distinguished from the highly similar and evidently closely related
<taxonomicName id="4C0E4D2FFFCDE53E512B81E2628CF8B5" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[270,391,1769,1793]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFCDE53E512B81E2628CF8B5" box="[270,391,1769,1793]" italics="true" pageId="12" pageNumber="129">Z. pindarae</emphasis>
</taxonomicName>
particularly by black tibiae, different punctation of the male elytra and of the abdominal tergite VII, by a basally much broader ventral process of the aedeagus (ventral view), and (both absolutely and relatively) distinctly longer parameres with a longer and more slender apical lobe of the paramere.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFCDE53E509880C167EDFEF5" pageId="12" pageNumber="129" type="distribution">
<paragraph id="8BB136ACFFCDE53E509880C167EDFEF5" blockId="12.[189,794,1993,2082]" lastBlockId="12.[831,1436,233,321]" pageId="12" pageNumber="129">
<emphasis id="B97AEABEFFCDE53E509880C16126F855" bold="true" box="[189,557,1993,2017]" pageId="12" pageNumber="129">Distribution and natural history</emphasis>
: The
<typeStatus id="54B5880EFFCDE53E525780C261ABF856" box="[626,672,1994,2018]" pageId="12" pageNumber="129">type</typeStatus>
specimens were collected in four localities in the
<collectingRegion id="49CAF84EFFCDE53E525180E161ECF7B5" box="[628,743,2025,2049]" country="Nepal" name="Dhawalagiri" pageId="12" pageNumber="129">Dhaulagiri</collectingRegion>
and Annapurna ranges (
<figureCitation id="13352A29FFCDE53E51BF8F0162E9F795" box="[410,482,2057,2081]" captionStart="Map 4" captionStartId="65.[154,199,794,815]" captionTargetBox="[151,1398,230,785]" captionTargetId="figure-63@65.[151,1398,230,785]" captionTargetPageId="65" captionText="Map 4: Revised distributions of Zyras condignus (black circles) and Z. morulus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742401" httpUri="https://zenodo.org/record/5742401/files/figure.png" pageId="12" pageNumber="129">Map 4</figureCitation>
). The species appears to be confined to high-altitude habitats; the elevations range from approximately 3300 to above
<quantity id="4CF69B49FFCDE53E54E38601661CFE95" box="[1222,1303,265,289]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="12" pageNumber="129" unit="m" value="4100.0">4100 m</quantity>
. One specimen collected in June is slightly teneral.
</paragraph>
</subSubSection>
</treatment>
</document>
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