332 lines
40 KiB
XML
332 lines
40 KiB
XML
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<mods:title id="4C7967065E939A71F12E4A07F6F12458">Two new species of the genus Sitana Cuvier, 1829 (Reptilia: Agamidae) from Sri Lanka, including a taxonomic revision of the Indian Sitana species</mods:title>
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<mods:namePart id="4888E0B38B8B0379E6265B465B5B9ABC">Thasun Amarasinghe, A. A.</mods:namePart>
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<mods:namePart id="0901E21EE122D77CB14BA020446BC3C4">Ineich, Ivan</mods:namePart>
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<mods:namePart id="57B0AE578A754A64A3173CBA5716D28B">Suranjan Karunarathna, D. M. S.</mods:namePart>
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<mods:namePart id="AAE1E5C1BED7C760FD7D18237C3E36E5">Botejue, W. Madhava S.</mods:namePart>
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<mods:namePart id="7FA03AA2A6BE9C526E6B31D2978DDF2A">Campbell, Patrick D.</mods:namePart>
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<mods:title id="3A0D43010B9E62720412BE10AC60DA2C">Zootaxa</mods:title>
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<mods:date id="8E184EADD73AAD68F3D0A5DB79BB54AA">2015</mods:date>
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<mods:number id="025E3560099E6B19952F4749C0B75E85">3915</mods:number>
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<treatment id="DA632C027E54E51146A6FEA5FDF1ED7B" ID-DOI="http://doi.org/10.5281/zenodo.6113353" ID-GBIF-Taxon="127671454" ID-Zenodo-Dep="6113353" LSID="urn:lsid:plazi:treatment:DA632C027E54E51146A6FEA5FDF1ED7B" httpUri="http://treatment.plazi.org/id/DA632C027E54E51146A6FEA5FDF1ED7B" lastPageId="17" lastPageNumber="84" pageId="14" pageNumber="81">
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<subSubSection id="1AD0CE9F7E54E50E46A6FEA5FD50E8D6" pageId="14" pageNumber="81" type="nomenclature">
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<paragraph id="52759D147E54E50E46A6FEA5FDA7E8AF" blockId="14.[151,550,373,433]" box="[151,550,373,399]" pageId="14" pageNumber="81">
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<heading id="093D2A787E54E50E46A6FEA5FDA7E8AF" bold="true" box="[151,550,373,399]" fontSize="11" level="1" pageId="14" pageNumber="81" reason="1">
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<taxonomicName id="95CAE6977E54E50E46A6FEA5FDA7E8AF" authority="Jerdon, 1870" authorityName="Jerdon" authorityYear="1870" box="[151,550,373,399]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="deccanensis">
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<emphasis id="60BE41067E54E50E46A6FEA5FDA7E8AF" bold="true" box="[151,550,373,399]" pageId="14" pageNumber="81">
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<emphasis id="60BE41067E54E50E46A6FEA5FEFCE8AF" bold="true" box="[151,381,373,399]" italics="true" pageId="14" pageNumber="81">Sitana deccanensis</emphasis>
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<bibRefCitation id="365BE0E57E54E50E47B5FEA5FDA7E8AF" author="Jerdon" box="[388,550,373,399]" pageId="14" pageNumber="81" refString="Jerdon, T. C. (1870) Notes on Indian Herpetology. Proceedings of the Asiatic Society Bengal, 1870, 66 - 85." type="proceedings paper" year="1870">Jerdon, 1870</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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<paragraph id="52759D147E54E50E46A6FE48FE53E890" blockId="14.[151,550,373,433]" box="[151,466,408,433]" pageId="14" pageNumber="81">
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(
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<figureCitation id="CAF181917E54E50E46AEFE48FF6CE891" box="[159,237,408,433]" captionStart="FIGURE 5" captionStartId="15.[151,250,1538,1560]" captionTargetBox="[311,1275,193,1516]" captionTargetId="figure@15.[311,1275,193,1517]" captionTargetPageId="15" captionText="FIGURE 5. Sitana deccanensis Jerdon, 1870 syntype (BMNH 1946.8.27.39) (A) dorsal view of head; (B) lateral view of head; (C) ventral view of head; (D) ventral view of thigh and tail base (photo: P. D. Campbell); (E) Sitana deccanensis live breeding male (not collected) with expanded throat-fan at Satara, Maharashtra State, India (photo: J. Katre in 2011)." httpUri="https://zenodo.org/record/287776/files/figure.png" pageId="14" pageNumber="81">Figs. 5</figureCitation>
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,
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<figureCitation id="CAF181917E54E50E46CBFE48FE8BE891" box="[250,266,408,433]" captionStart="FIGURE 6" captionStartId="16.[151,250,1829,1851]" captionTargetBox="[311,1275,484,1807]" captionTargetId="figure@16.[311,1275,484,1808]" captionTargetPageId="16" captionText="FIGURE 6. Sitana deccanensis live breeding male (not collected) with expanded throat-fan at Belgaum, Karnataka State, India (photo: N. Sant)." httpUri="https://zenodo.org/record/287777/files/figure.png" pageId="14" pageNumber="81">6</figureCitation>
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,
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<figureCitation id="CAF181917E54E50E4727FE48FEA7E890" box="[278,294,408,432]" captionStart="FIGURE 8" captionStartId="20.[151,250,1229,1251]" captionTargetBox="[311,1275,193,1204]" captionTargetId="figure@20.[311,1275,193,1208]" captionTargetPageId="20" captionText="FIGURE 8. Sitana deccanensis syntype (BMNH 1946.8.27.39) (A) lateral body scales; (B) pectoral scales; (C) abdominal scales; Sitana ponticeriana lectotype (MNHN 6901) (D) lateral body scales; (E) pectoral scales; (F) abdominal scales (illustrated by A. A. T. Amarasinghe)." httpUri="https://zenodo.org/record/287779/files/figure.png" pageId="14" pageNumber="81">8</figureCitation>
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;
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<tableCitation id="1F48A8AF7E54E50E4702FE48FE10E890" box="[307,401,408,433]" captionStart="TABLE 1. A" captionStartId="8.[151,239,152,175]" captionTargetBox="[159,1414,234,1331]" captionTargetPageId="8" captionText="TABLE 1. A comparison of morphometric and meristic characters of male specimens of Sitana ponticeriana, S. deccanensis, S. bahiri sp. nov., and S. devakai sp. nov. based on examined materials." httpUri="http://table.plazi.org/id/06B5CD9C7E52E50846A6FF48FBA5E9EF" pageId="14" pageNumber="81" tableUuid="06B5CD9C7E52E50846A6FF48FBA5E9EF">Tables 1</tableCitation>
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,
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<tableCitation id="1F48A8AF7E54E50E47AEFE48FE2EE890" box="[415,431,408,432]" captionStart="TABLE 2" captionStartId="8.[151,239,1396,1419]" captionTargetBox="[159,1424,1478,1655]" captionTargetPageId="8" captionText="TABLE 2. External morphological character variation in Sitana ponticeriana, S. deccanensis, S. bahiri sp. nov., and S. devakai sp. nov. based on examined male specimens." httpUri="http://table.plazi.org/id/06B5CD9C7E52E50846A6FAA4FD51EC8B" pageId="14" pageNumber="81" tableUuid="06B5CD9C7E52E50846A6FAA4FD51EC8B">2</tableCitation>
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,
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||
<tableCitation id="1F48A8AF7E54E50E478AFE48FE4BE890" box="[443,458,408,432]" captionStart="TABLE 4" captionStartId="26.[151,239,404,427]" captionTargetBox="[159,1419,486,828]" captionTargetPageId="26" captionText="TABLE 4. Clarification of specimens data which were held at the BMNH collection prior to 1885 based on Boulenger (1885) and the museum registry; M, male; F, female; (J), juvenile." httpUri="http://table.plazi.org/id/06B5CD9C7E40E51A46A6FE44FCD9E8EB" pageId="14" pageNumber="81" tableUuid="06B5CD9C7E40E51A46A6FE44FCD9E8EB">4</tableCitation>
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)
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</paragraph>
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||
<paragraph id="52759D147E54E50E46A6FE31FD50E8D6" blockId="14.[151,1149,480,563]" box="[151,721,480,502]" pageId="14" pageNumber="81">
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<taxonomicName id="95CAE6977E54E50E46A6FE31FE5EE8D6" box="[151,479,480,502]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="subSpecies" species="ponticeriana" subSpecies="deccanensis">
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<emphasis id="60BE41067E54E50E46A6FE31FE5EE8D6" box="[151,479,480,502]" italics="true" pageId="14" pageNumber="81">Sitana ponticeriana deccanensis</emphasis>
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</taxonomicName>
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—
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<bibRefCitation id="365BE0E57E54E50E47CBFE30FD4CE8D6" author="Deraniyagala" box="[506,717,480,502]" pageId="14" pageNumber="81" refString="Deraniyagala, P. E. P. (1953) A Colored Atlas of some vertebrates from Ceylon, Tetrapod Reptilia. Vol. 2. National Museums of Sri Lanka, Colombo, 101 pp." type="book" year="1953">Deraniyagala (1953)</bibRefCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="1AD0CE9F7E54E50E46A6FE2FFECBEA55" pageId="14" pageNumber="81" type="reference_group">
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||
<paragraph id="52759D147E54E50E46A6FE2FFC49EB13" blockId="14.[151,1149,480,563]" pageId="14" pageNumber="81">
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||
<taxonomicName id="95CAE6977E54E50E46A6FE2FFE9BEB35" box="[151,282,511,533]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="minor">
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<emphasis id="60BE41067E54E50E46A6FE2FFE9BEB35" box="[151,282,511,533]" italics="true" pageId="14" pageNumber="81">Sitana minor</emphasis>
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</taxonomicName>
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—Schleich & Kästle (1998), Schleich
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<emphasis id="60BE41067E54E50E4491FDD0FD4DEB34" box="[672,716,510,533]" italics="true" pageId="14" pageNumber="81">et al</emphasis>
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||
. (1998),
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||
<bibRefCitation id="365BE0E57E54E50E4518FE2EFB9DEB34" author="Anders" box="[809,1052,510,532]" pageId="14" pageNumber="81" refString="Anders, C. & Kastle, W. (2002) Amphibians and Reptiles of Nepal - Lizards and Crocodiles. Koeltz Scientific Books, Konigstein, 39 pp. [Schleich, H. H. & Kastle, W. (Eds.)]" type="book" year="2002">Anders & Kästle (2002)</bibRefCitation>
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||
[
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||
<emphasis id="60BE41067E54E50E421DFDD0FBF1EB35" box="[1068,1136,512,533]" italics="true" pageId="14" pageNumber="81">partim</emphasis>
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||
].
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||
<taxonomicName id="95CAE6977E54E50E46A6FDCEFEDEEB13" box="[151,351,542,563]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="ponticeriana">
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||
<emphasis id="60BE41067E54E50E46A6FDCEFEDEEB13" box="[151,351,542,563]" italics="true" pageId="14" pageNumber="81">Sitana ponticeriana</emphasis>
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||
</taxonomicName>
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||
—Das & de Silva (2005) [
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||
<emphasis id="60BE41067E54E50E445AFDCEFD2EEB13" box="[619,687,542,563]" italics="true" pageId="14" pageNumber="81">partim</emphasis>
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],
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||
<bibRefCitation id="365BE0E57E54E50E44F2FDCDFCE6EB13" author="Manthey" box="[707,871,541,563]" pageId="14" pageNumber="81" refString="Manthey, U. (2010) Agamid Lizards of Southern Asia, Draconinae 2 & Leiolepidinae. Edition Chimaira & A. C. S. Glaser, Germany, Frankfurt am Main, 168 pp." type="book" year="2010">Manthey (2010)</bibRefCitation>
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[
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||
<emphasis id="60BE41067E54E50E4547FDCEFC3BEB13" box="[886,954,542,563]" italics="true" pageId="14" pageNumber="81">partim</emphasis>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="52759D147E54E50E46A6FDB0FE0FEBC5" blockId="14.[151,1437,608,2001]" pageId="14" pageNumber="81">
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<emphasis id="60BE41067E54E50E46A6FDB0FE88EB59" bold="true" box="[151,265,608,633]" pageId="14" pageNumber="81">
|
||
<typeStatus id="8D7123B67E54E50E46A6FDB0FE84EB59" box="[151,261,608,633]" pageId="14" pageNumber="81" type="syntype">Syntypes</typeStatus>
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||
.
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||
</emphasis>
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||
Male, BMNH 1946.8.27.39, SVL
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||
<quantity id="953230F17E54E50E44A0FDB0FD79EB58" box="[657,760,608,633]" metricMagnitude="-2" metricUnit="m" metricValue="6.76" pageId="14" pageNumber="81" unit="mm" value="67.6">67.6 mm</quantity>
|
||
,
|
||
<collectingCountry id="2ADDDD847E54E50E4535FDB0FCC1EB59" box="[772,832,608,633]" name="India" pageId="14" pageNumber="81">India</collectingCountry>
|
||
, presented by J. E. Gray; male, BMNH 1946.8.27.40, SVL 60.0 mm,
|
||
<collectingCountry id="2ADDDD847E54E50E4761FD54FE0DEBBD" box="[336,396,644,669]" name="India" pageId="14" pageNumber="81">India</collectingCountry>
|
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, presented by J. E. Gray.
|
||
<emphasis id="60BE41067E54E50E44F1FD54FD7EEBBD" bold="true" box="[704,767,644,669]" pageId="14" pageNumber="81">Note.</emphasis>
|
||
We discovered the original
|
||
<typeStatus id="8D7123B67E54E50E427CFD55FBFCEBBD" box="[1101,1149,645,669]" pageId="14" pageNumber="81">type</typeStatus>
|
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series of
|
||
<taxonomicName id="95CAE6977E54E50E42C4FD55FA1AEBBD" box="[1269,1435,644,669]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E54E50E42C4FD55FA1AEBBD" box="[1269,1435,644,669]" italics="true" pageId="14" pageNumber="81">S. deccanensis</emphasis>
|
||
</taxonomicName>
|
||
(comprising two
|
||
<typeStatus id="8D7123B67E54E50E476BFD78FE3FEBE0" box="[346,446,680,704]" pageId="14" pageNumber="81" type="syntype">syntypes</typeStatus>
|
||
) among the
|
||
<typeStatus id="8D7123B67E54E50E4461FD79FD33EBE1" box="[592,690,681,705]" pageId="14" pageNumber="81" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="95CAE6977E54E50E44EAFD79FC69EBE1" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[731,1000,680,705]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="minor">
|
||
<emphasis id="60BE41067E54E50E44EAFD79FCBDEBE0" box="[731,828,681,704]" italics="true" pageId="14" pageNumber="81">S. minor</emphasis>
|
||
<bibRefCitation id="365BE0E57E54E50E4574FD78FC69EBE1" author="Gunther" box="[837,1000,680,705]" pageId="14" pageNumber="81" refString="Gunther, A. C. L. G. (1864) The Reptiles of British India. The Ray Society, London, 452 pp." type="book" year="1864">Günther, 1864</bibRefCitation>
|
||
</taxonomicName>
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||
and they probably got mixed up later (probably after 1870).
|
||
</paragraph>
|
||
<paragraph id="52759D147E54E50E46F6FD20FECBEA55" blockId="14.[151,1437,608,2001]" pageId="14" pageNumber="81">
|
||
<emphasis id="60BE41067E54E50E46F6FD20FD96EA29" bold="true" box="[199,535,752,777]" pageId="14" pageNumber="81">Other Specimens examined.</emphasis>
|
||
Males, MNHN 2568, MNHN 6903–4, respective SVL
|
||
<quantity id="953230F17E54E50E42A0FD20FB62EA28" box="[1169,1251,752,777]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="14" pageNumber="81" unit="mm" value="62.0">62 mm</quantity>
|
||
,
|
||
<quantity id="953230F17E54E50E42C1FD20FAC3EA28" box="[1264,1346,752,777]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="14" pageNumber="81" unit="mm" value="69.0">69 mm</quantity>
|
||
, and
|
||
<quantity id="953230F17E54E50E43B1FD20FF44EA0C" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="81" unit="mm" value="78.0">78 mm</quantity>
|
||
, “Indes Orientales”, coll. Jacquemont; Male, MNHN 6023, SVL
|
||
<quantity id="953230F17E54E50E4597FCC5FC76EA0C" box="[934,1015,789,813]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="14" pageNumber="81" unit="mm" value="74.0">74 mm</quantity>
|
||
, “Bellari (
|
||
<collectingCountry id="2ADDDD847E54E50E425DFCC4FB25EA0D" box="[1132,1188,788,813]" name="India" pageId="14" pageNumber="81">Inde</collectingCountry>
|
||
)”, coll. Chaper; Male, BMNH 1946.8.27.41 (now
|
||
<typeStatus id="8D7123B67E54E50E47FCFCE8FDE5EA71" box="[461,612,824,849]" pageId="14" pageNumber="81" type="paralectotype">paralectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="95CAE6977E54E50E44BBFCE9FD68EA70" box="[650,745,825,848]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="minor">
|
||
<emphasis id="60BE41067E54E50E44BBFCE9FD68EA70" box="[650,745,825,848]" italics="true" pageId="14" pageNumber="81">S. minor</emphasis>
|
||
</taxonomicName>
|
||
; see below),
|
||
<collectingCountry id="2ADDDD847E54E50E45B1FCE8FC38EA71" box="[896,953,824,849]" name="India" pageId="14" pageNumber="81">India</collectingCountry>
|
||
(Madras, in error), SVL
|
||
<quantity id="953230F17E54E50E42E5FCE8FAA4EA70" box="[1236,1317,824,849]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="81" unit="mm" value="60.0">60 mm</quantity>
|
||
, presented by T. C. Jerdon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E54E50E46F6FC50FD79EE61" pageId="14" pageNumber="81" type="diagnosis">
|
||
<paragraph id="52759D147E54E50E46F6FC50FD79EE61" blockId="14.[151,1437,608,2001]" pageId="14" pageNumber="81">
|
||
<emphasis id="60BE41067E54E50E46F6FC50FE7AEAB9" bold="true" box="[199,507,896,921]" pageId="14" pageNumber="81">Diagnosis for adult males.</emphasis>
|
||
<taxonomicName id="95CAE6977E54E50E4433FC51FD56EAB9" box="[514,727,896,921]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E54E50E4433FC51FD56EAB9" box="[514,727,896,921]" italics="true" pageId="14" pageNumber="81">Sitana deccanensis</emphasis>
|
||
</taxonomicName>
|
||
differs from each congener by having the following characters:
|
||
<emphasis id="60BE41067E54E50E46A6FC74FEDFEA9D" bold="true" box="[151,350,932,957]" pageId="14" pageNumber="81">
|
||
<taxonomicName id="95CAE6977E54E50E46A6FC74FF7AEA9D" box="[151,251,932,957]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="bahiri" status="sp. nov.">
|
||
<emphasis id="60BE41067E54E50E46A6FC74FF7AEA9D" bold="true" box="[151,251,932,957]" italics="true" pageId="14" pageNumber="81">S. bahiri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E54E50E4735FC74FEDFEA9D" box="[260,350,932,957]" pageId="14" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: larger SVL above 60.0 mm (vs. smaller SVL 40.0–50.0 mm), snout shorter ES 53.1–53.4% of HW (vs. longer, ES 54.7–57.4% of HW), throat-fan longer ADW 50.3–60.2% of AG (vs. shorter, ADW 28.3–29.2% of AG), femur longer FEL 79.7–82.9% of TBL (vs. shorter, FEL 70.6–75.3% of TBL), foot shorter FOL 131.9–144.6% of HL (vs. longer, FOL 158.9–159.8 of HL), supralabials 12 (vs. 7–8), midbody scales 66–70 (vs. 49–56), subdigital lamellae on toe
|
||
<date id="2674BBD47E54E50E4478FBE5FD06ED6D" box="[585,647,1077,1101]" pageId="14" pageNumber="81" value="2013-03">III 13</date>
|
||
(vs. 14–17), subdigital lamellae on toe
|
||
<date id="2674BBD47E54E50E420EFBE5FB28ED6D" box="[1087,1193,1077,1101]" pageId="14" pageNumber="81" value="1919-04-23">IV 19–23</date>
|
||
(vs. 23–26), enlarged scales above the tympanum absent (vs. present), equal and regular lateral scales without intermediate enlarged scales (vs. unequal and irregular lateral scales with enlarged scales), breeding-males with bright red patch extending to vent (vs. bleached orange colour patch), 3–4 bright stripes from snout and behind the eye up to the shoulders (vs. a single pale stripe from snout up to the shoulder);
|
||
<emphasis id="60BE41067E54E50E454FFB14FBD9EDFD" bold="true" box="[894,1112,1220,1245]" pageId="14" pageNumber="81">
|
||
<taxonomicName id="95CAE6977E54E50E454FFB14FC77EDFD" box="[894,1014,1220,1245]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="devakai" status="sp. nov.">
|
||
<emphasis id="60BE41067E54E50E454FFB14FC77EDFD" bold="true" box="[894,1014,1220,1245]" italics="true" pageId="14" pageNumber="81">S. devakai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E54E50E45CCFB14FBD9EDFD" box="[1021,1112,1220,1245]" pageId="14" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: larger SVL above 60.0 mm (vs. smaller SVL 40.0–
|
||
<quantity id="953230F17E54E50E47AAFB38FD83EC20" box="[411,514,1256,1281]" metricMagnitude="-2" metricUnit="m" metricValue="4.56" pageId="14" pageNumber="81" unit="mm" value="45.6">45.6 mm</quantity>
|
||
), snout shorter ES 53.1–53.4% of HW (vs. longer, ES 57.2–63.2% of HW), throatfan longer ADW 50.3–60.2% of AG (vs. shorter, ADW 29.9–32.5% of AG), femur longer FEL 79.7–82.9% of TBL (vs. shorter, FEL 72.2–78.4% of TBL), foot shorter FOL 131.9–144.6% of HL (vs. longer, FOL 154.7–180.2 of HL), supralabials 12 (vs. 8–9), midbody scale rows 66–70 (vs. 57–59), ventral scales 82–92 (vs. 100–108), subdigital lamellae on toe
|
||
<date id="2674BBD47E54E50E47FCFAA9FD91ECB0" box="[461,528,1400,1424]" pageId="14" pageNumber="81" value="2013-03">III 13</date>
|
||
(vs. 15–16), enlarged scales above the tympanum absent (vs. present), lateral scales without intermediate enlarged scales (vs. with enlarged scales), breeding-males with bright red patch extending to vent (vs. bright red colour patch restricted to mid throat-fan), 3–4 bright stripes from snout and behind the eye up to the shoulders (vs. a single pale stripe from snout up to the shoulder);
|
||
<emphasis id="60BE41067E54E50E4268FA34FA95ECDD" bold="true" box="[1113,1300,1508,1533]" pageId="14" pageNumber="81">
|
||
<taxonomicName id="95CAE6977E54E50E4268FA34FA8BECDD" box="[1113,1290,1508,1533]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="ponticeriana">
|
||
<emphasis id="60BE41067E54E50E4268FA34FA8BECDD" bold="true" box="[1113,1290,1508,1533]" italics="true" pageId="14" pageNumber="81">S. ponticeriana</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
larger SVL above 60.0 mm (vs. smaller SVL 48.0–49.0 mm), snout shorter ES 53.1–53.4% of HW (vs. longer, ES 61.5–68.1% of HW), throat-fan longer ADW 50.3–60.2% of AG (vs. shorter, ADW 31.1–54.7% of AG), foot shorter FOL 131.9–144.6% of HL (vs. longer, FOL 150.9–175.5% of HL), supralabials 12 (vs. 8–10), midbody scale rows 66–70 (vs. 50–54), ventral scales 82–92 (vs. 65–69), no enlarged scales above the tympanum (vs. present), equal and regular lateral scales without intermediate enlarged scales (vs. unequal and irregular lateral scales with enlarged scales), breeding-males with bright red patch extending to vent (vs. bright red colour patch restricted to mid throat-fan);
|
||
<emphasis id="60BE41067E54E50E4768F90FFD14EFD9" bold="true" box="[345,661,1759,1785]" pageId="14" pageNumber="81">
|
||
<taxonomicName id="95CAE6977E54E50E4768F90FFD9DEFD8" box="[345,540,1759,1784]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="sivalensis">
|
||
<emphasis id="60BE41067E54E50E4768F90FFD9DEFD8" bold="true" box="[345,540,1759,1784]" italics="true" pageId="14" pageNumber="81">Sitana sivalensis</emphasis>
|
||
</taxonomicName>
|
||
complex:
|
||
</emphasis>
|
||
larger SVL above 60.0 mm (vs. smaller SVL of 35.0–
|
||
<quantity id="953230F17E54E50E4315F930FA11EFD8" box="[1316,1424,1760,1785]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" pageId="14" pageNumber="81" unit="mm" value="44.5">44.5 mm</quantity>
|
||
), throat-fan extending up to the abdomen (vs. not extending to the level of the axilla), breeding-males with bright red patch extending to vent (vs. no such red colour patch).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E54E51146F6F89CFAD1EACF" lastPageId="17" lastPageNumber="84" pageId="14" pageNumber="81" type="description">
|
||
<paragraph id="52759D147E54E51046F6F89CFB8BE88F" blockId="14.[151,1437,608,2001]" lastBlockId="16.[151,1436,151,431]" lastPageId="16" lastPageNumber="83" pageId="14" pageNumber="81">
|
||
<emphasis id="60BE41067E54E50E46F6F89CFE61EE45" bold="true" box="[199,480,1868,1893]" pageId="14" pageNumber="81">
|
||
Description of
|
||
<typeStatus id="8D7123B67E54E50E474DF89CFE5BEE45" box="[380,474,1868,1893]" pageId="14" pageNumber="81" type="syntype">syntype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(Based on BMNH 1946.8.27.39). An adult male,
|
||
<quantity id="953230F17E54E50E4219F89CFB0EEE44" box="[1064,1167,1868,1893]" metricMagnitude="-2" metricUnit="m" metricValue="6.76" pageId="14" pageNumber="81" unit="mm" value="67.6">67.6 mm</quantity>
|
||
SVL; head moderately large (HL 28.7% of SVL), elongate (HW 62.4% of HL), narrow (HW 17.9% of SVL), distinct from neck; snout short (ES 53.1% of HW); snout length somewhat greater than eye diameter (ED 91.7% of ES); interorbital distance broad (IO 20.1% of HL); eye large (ED 30.4% of HL); pupil rounded; ear-opening shallow, its greatest diameter being dorsoventrally, ear-opening with enlarged unkeeled scales anteriorly but not posteriorly, tympanum smaller than orbit (
|
||
<collectingCountry id="2ADDDD847E55E50F472DF977FED6EF9F" box="[284,343,1703,1727]" name="Chad" pageId="15" pageNumber="82">TYD</collectingCountry>
|
||
40.7% of ED); no spines or enlarged scales above the tympanum, but the scales above the tympanum are keeled; diameter of eyes greater than eye to ear distance (ED 122.9% of TYE); forehead concave; scales on snout keeled, smaller than those of occipital region; scales on interorbital and supercilium area keeled; no nuchal or dorsal crest; rostral scale width greater than height, ventro-posteriorly in contact with the first supralabial, in contact posteriorly with the prenasal and three equal sized postrostral scales; two supranasals around nostrils on each side, three postnasals with the lower smallest and the middle largest, a single upper supralabial and one prenasal; nostrils oval and located in the middle of an undivided nasal plate;
|
||
<emphasis id="60BE41067E55E50F4215F8AEFB60EEB7" box="[1060,1249,1918,1943]" italics="true" pageId="15" pageNumber="82">canthus rostralis</emphasis>
|
||
and superciliary edges sharp; parietal plate lozenge shaped (anterior sides shorter than posterior sides) including a grey-blue coloured “pineal eye”, parietal plate larger than adjacent plates. Mental subtriangular, approximately equal in length and width, posteriorlaterally in contact with two enlarged postmentals separated by a third smaller median gular scale preventing contact between them; first postmental pair bordered posteriorly by four smooth scales exclusive of infralabial. Throat-fan large, posterior scales reach mid venter (DWL 70.6% of SVL, ADW 50.3% of AG), margin of the anterior part of the throat-fan covered with smooth, overlapping, bluntly pointed scales, posterior part (from the middle of the throat to the midventer) with long, elongate and lanceolate bluntly pointed scales; throat scales rounded, smooth and overlapping; three scale rows separate orbit from supralabials; supralabials 12 (9th in mid orbit position); infralabials 10, decreasing in size towards gape; scales on the throat-fan smooth but adjacent ventral scales slightly keeled and mucronate, overlapping.
|
||
</paragraph>
|
||
<caption id="06B5CD9C7E55E50F46A6F9D2FB4FEF75" httpUri="https://zenodo.org/record/287776/files/figure.png" pageId="15" pageNumber="82" targetBox="[311,1275,193,1516]" targetPageId="15">
|
||
<paragraph id="52759D147E55E50F46A6F9D2FB4FEF75" blockId="15.[151,1436,1538,1621]" pageId="15" pageNumber="82">
|
||
<emphasis id="60BE41067E55E50F46A6F9D2FE93EF38" bold="true" box="[151,274,1538,1560]" pageId="15" pageNumber="82">FIGURE 5.</emphasis>
|
||
<taxonomicName id="95CAE6977E55E50F4729F9D3FDE3EF38" authority="Jerdon, 1870" authorityName="Jerdon" authorityYear="1870" box="[280,610,1538,1560]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="82" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E55E50F4729F9D3FE58EF38" box="[280,473,1538,1560]" italics="true" pageId="15" pageNumber="82">Sitana deccanensis</emphasis>
|
||
Jerdon, 1870
|
||
</taxonomicName>
|
||
syntype (BMNH 1946.8.27.39) (
|
||
<emphasis id="60BE41067E55E50F4583F9D2FC4DEF39" bold="true" box="[946,972,1538,1561]" pageId="15" pageNumber="82">A)</emphasis>
|
||
dorsal view of head; (
|
||
<emphasis id="60BE41067E55E50F429FF9D2FB47EF39" bold="true" box="[1198,1222,1538,1561]" pageId="15" pageNumber="82">B)</emphasis>
|
||
lateral view of head; (
|
||
<emphasis id="60BE41067E55E50F46AEF9F0FF3BEF17" bold="true" box="[159,186,1568,1591]" pageId="15" pageNumber="82">C)</emphasis>
|
||
ventral view of head; (
|
||
<emphasis id="60BE41067E55E50F4799F9F0FE43EF17" bold="true" box="[424,450,1568,1591]" pageId="15" pageNumber="82">D)</emphasis>
|
||
ventral view of thigh and tail base (photo: P.D. Campbell); (
|
||
<emphasis id="60BE41067E55E50F421FF9F1FBBEEF16" bold="true" box="[1070,1087,1569,1590]" pageId="15" pageNumber="82">E</emphasis>
|
||
)
|
||
<taxonomicName id="95CAE6977E55E50F427FF9F2FA8EEF17" box="[1102,1295,1569,1591]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="82" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E55E50F427FF9F2FA8EEF17" box="[1102,1295,1569,1591]" italics="true" pageId="15" pageNumber="82">Sitana deccanensis</emphasis>
|
||
</taxonomicName>
|
||
live breeding male (not collected) with expanded throat-fan at Satara, Maharashtra State, India (photo: J. Katre in 2011).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="06B5CD9C7E4AE51046A6F8F5FEC3EE7A" httpUri="https://zenodo.org/record/287777/files/figure.png" pageId="16" pageNumber="83" targetBox="[311,1275,484,1807]" targetPageId="16">
|
||
<paragraph id="52759D147E4AE51046A6F8F5FEC3EE7A" blockId="16.[151,1435,1829,1882]" pageId="16" pageNumber="83">
|
||
<emphasis id="60BE41067E4AE51046A6F8F5FE93EE1A" bold="true" box="[151,274,1829,1851]" pageId="16" pageNumber="83">FIGURE 6.</emphasis>
|
||
<taxonomicName id="95CAE6977E4AE5104729F8F6FE58EE1B" box="[280,473,1829,1851]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E4AE5104729F8F6FE58EE1B" box="[280,473,1829,1851]" italics="true" pageId="16" pageNumber="83">Sitana deccanensis</emphasis>
|
||
</taxonomicName>
|
||
live breeding male (not collected) with expanded throat-fan at Belgaum, Karnataka State, India (photo: N. Sant).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="52759D147E4AE51146F6F851FB97E83C" blockId="16.[151,1436,1921,2018]" lastBlockId="17.[151,1437,151,1115]" lastPageId="17" lastPageNumber="84" pageId="16" pageNumber="83">Body slender (AG 45.6% of SVL); mid dorsal scales equal, keeled, elongate with pointed dorsal scales at midbody, keels straight forming longitudinal continuous parallel ridges; scales on dorsum at midbody larger than those of venter at same level; lateral body scales equal, regular, keeled, mucronate, smaller than dorsals and without randomly distributed enlarged scales; upper dorsolateral scales directed backwards and upwards, dorsoventral ones directed backwards and downwards; 70 scales around midbody; pectoral scales not enlarged, keeled, mucronate, short and overlapping; abdominal scales enlarged and bluntly-pointed; ventral keels forming regular and parallel continuous ridges; no preanal or femoral pores. Ventrals 92.</paragraph>
|
||
<paragraph id="52759D147E4BE51146F6FEF8FE70EB1C" blockId="17.[151,1437,151,1115]" pageId="17" pageNumber="84">Upper arm somewhat shorter than lower arm (UAL 19.5% of SVL; LAL 19.8% of SVL); femur shorter than tibia (FEL 26.5% of SVL; TBL 31.9% of SVL; FEL 82.9% of TBL). Dorsal scales on fore and hind limbs not enlarged, keeled, overlapping, and mucronate; ventral scales on upper and lower arm not enlarged, keeled, overlapping, and mucronate; scales on dorsal and ventral surface of thigh keeled, overlapping and mucronate; keels on dorsal and ventral arm and foot forming a series of continuous parallel ridges. Digits elongate, slender, all bearing slightly recurved claws; claws are sharp and elongate; subdigital lamellae entire and mucronate, 13 subdigital lamellae on the toe III, and 19 on toe IV; inter-digital webbing absent; relative length of fingers 3> 4> 2> 5> 1, and toes 4> 3> 2> 1.</paragraph>
|
||
<paragraph id="52759D147E4BE51146F6FD97FDC6EB88" blockId="17.[151,1437,151,1115]" pageId="17" pageNumber="84">Tail incomplete (broken); tail base swollen, ventral scales and dorsal scales on tail keeled, elongate, overlapping, directed backwards, keels forming continuous parallel ridges; remnant tail with subcaudals on median row not enlarged, keeled, overlapping.</paragraph>
|
||
<paragraph id="52759D147E4BE51146F6FD64FB12EA37" blockId="17.[151,1437,151,1115]" pageId="17" pageNumber="84">
|
||
<emphasis id="60BE41067E4BE51146F6FD64FDBBEBED" bold="true" box="[199,570,692,717]" pageId="17" pageNumber="84">Colour in preserved specimen.</emphasis>
|
||
Based on
|
||
<typeStatus id="8D7123B67E4BE5114487FD65FC91EBED" box="[694,784,693,717]" pageId="17" pageNumber="84" type="syntype">syntype</typeStatus>
|
||
, BMNH 1946.8.27.39. Colouration and markings almost faded. General colouration whitish dorsally and ventrally with some scattered darker marks irregularly placed. Throat-fan with three colours: anterior 3/4th light blue and black and posterior 1/4th whitish.
|
||
</paragraph>
|
||
<paragraph id="52759D147E4BE51146F6FCF2FB8EEAA2" blockId="17.[151,1437,151,1115]" pageId="17" pageNumber="84">
|
||
<emphasis id="60BE41067E4BE51146F6FCF2FEFEEA1B" bold="true" box="[199,383,802,827]" pageId="17" pageNumber="84">Colour in life.</emphasis>
|
||
See following corresponding figures of
|
||
<taxonomicName id="95CAE6977E4BE511454BFCF3FBA7EA1B" box="[890,1062,802,827]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E4BE511454BFCF3FBA7EA1B" box="[890,1062,802,827]" italics="true" pageId="17" pageNumber="84">S. deccanensis</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="365BE0E57E4BE511426DFCF2FA19EA1B" author="Manthey" box="[1116,1432,802,827]" pageId="17" pageNumber="84" refString="Manthey, U. (2010) Agamid Lizards of Southern Asia, Draconinae 2 & Leiolepidinae. Edition Chimaira & A. C. S. Glaser, Germany, Frankfurt am Main, 168 pp." type="book" year="2010">Manthey (2010: 155–157)</bibRefCitation>
|
||
: RA04103–111 identified as “
|
||
<taxonomicName id="95CAE6977E4BE51147D3FC97FDABEA7E" box="[482,554,839,862]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="60BE41067E4BE51147D3FC97FDABEA7E" box="[482,554,839,862]" italics="true" pageId="17" pageNumber="84">Sitana</emphasis>
|
||
</taxonomicName>
|
||
sp. A”. See also the illustration [Atlas - plate 16, figure 2] of
|
||
<taxonomicName id="95CAE6977E4BE51142DEFC97FA1DEA7F" box="[1263,1436,839,863]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="species" species="ponticeriana">
|
||
<emphasis id="60BE41067E4BE51142DEFC97FA1DEA7F" box="[1263,1436,839,863]" italics="true" pageId="17" pageNumber="84">S. ponticeriana</emphasis>
|
||
</taxonomicName>
|
||
which was made by
|
||
<bibRefCitation id="365BE0E57E4BE511474DFCBBFDBCEAA3" author="Duvernoy" box="[380,573,874,899]" pageId="17" pageNumber="84" refString="Duvernoy, L. G. (1846) Atlas (Les reptiles). Le Regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee, plate 16." type="book chapter" year="1846">Duvernoy (1846)</bibRefCitation>
|
||
but which corresponds to
|
||
<taxonomicName id="95CAE6977E4BE5114554FCBBFB89EAA3" box="[869,1032,874,899]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E4BE5114554FCBBFB89EAA3" box="[869,1032,874,899]" italics="true" pageId="17" pageNumber="84">S. deccanensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="52759D147E4BE51146F6FC5EFDB2EAEB" blockId="17.[151,1437,151,1115]" pageId="17" pageNumber="84">
|
||
<emphasis id="60BE41067E4BE51146F6FC5EFCF7EA87" bold="true" box="[199,886,910,935]" pageId="17" pageNumber="84">
|
||
Variation in the second
|
||
<typeStatus id="8D7123B67E4BE51147CAFC5EFDD6EA87" box="[507,599,910,935]" pageId="17" pageNumber="84" type="syntype">syntype</typeStatus>
|
||
(BMNH 1946.8.27.40).
|
||
</emphasis>
|
||
66 scale rows around midbody; ventrals, 83; subdigital lamellae on the toe IV, 23.
|
||
</paragraph>
|
||
<paragraph id="52759D147E4BE51146F6FC06FAD1EACF" blockId="17.[151,1437,151,1115]" box="[199,1360,982,1007]" pageId="17" pageNumber="84">
|
||
<emphasis id="60BE41067E4BE51146F6FC06FD86EACF" bold="true" box="[199,519,982,1007]" pageId="17" pageNumber="84">Suggested common names.</emphasis>
|
||
English name: Deccan Fan-throat Lizard; French name: Sitane du Deccan.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E4BE51146F6FC2AFDF1ED7B" pageId="17" pageNumber="84" type="distribution">
|
||
<paragraph id="52759D147E4BE51146F6FC2AFDF1ED7B" blockId="17.[151,1437,151,1115]" pageId="17" pageNumber="84">
|
||
<emphasis id="60BE41067E4BE51146F6FC2AFEE1ED33" bold="true" box="[199,352,1018,1043]" pageId="17" pageNumber="84">Distribution.</emphasis>
|
||
See map
|
||
<figureCitation id="CAF181917E4BE51147E9FC2AFD9EED33" box="[472,543,1018,1043]" captionStart="FIGURE 9" captionStartId="21.[151,250,1528,1550]" captionTargetBox="[311,1267,179,1495]" captionTargetId="figure@21.[311,1275,179,1503]" captionTargetPageId="21" captionText="FIGURE 9. Distribution map of S. ponticeriana (yellow), S. deccanensis (green), Sitana cf. devakai (purple), S. cf. ponticeriana (orange), Sitana sivalensis complex (grey), S. bahiri sp. nov. (blue); S. devakai sp. nov. (black); and S. ponticeriana mucronata (red). Holotype locality of each Sri Lankan species marked with squares." httpUri="https://zenodo.org/record/287780/files/figure.png" pageId="17" pageNumber="84">Fig. 9</figureCitation>
|
||
. South-western
|
||
<collectingCountry id="2ADDDD847E4BE51144EBFC2AFC97ED33" box="[730,790,1018,1043]" name="India" pageId="17" pageNumber="84">India</collectingCountry>
|
||
, observed (not collected) populations referred to
|
||
<taxonomicName id="95CAE6977E4BE5114365FC2BFE9FED17" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E4BE5114365FC2BFE9FED17" italics="true" pageId="17" pageNumber="84">Sitana deccanensis</emphasis>
|
||
</taxonomicName>
|
||
from Belgaum in Karnataka State and Satara in Maharashtra State. Also see MNHN 6023 which was collected from Bellary in Karnataka State.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |