307 lines
48 KiB
XML
307 lines
48 KiB
XML
<document id="86D66E92DBC43C550BAB04C1D0DAADEC" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FFF2E30EFF78FA87FBBC8058" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Harpes lahceni Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="47" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="46" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:mods id="0ABF00A86AAB458FFB72A9130E1978D8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D95724FD4CE65E88270ADE561C7523A6">
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<mods:title id="87ECF3B5934BC00A1881170C711DEBED">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<mods:name id="DC74F711AC01A16D84D44F72D32C7204" type="personal">
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<mods:roleTerm id="074F9BA1E6155760FE8B83B4F762FCFB">Author</mods:roleTerm>
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<mods:namePart id="7F568CFB2DDC6FCD9A3277F856C50250">Johnson, Robert G.</mods:namePart>
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<mods:typeOfResource id="786958157FCEFB875CE2CE255106C1E3">text</mods:typeOfResource>
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<mods:title id="652BB2A59C94EE27FC2A70BDDF3789AD">Zootaxa</mods:title>
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<mods:part id="C77FCE5D70E08573D5DBBDF10C089AD1">
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<mods:date id="24849D059494713F35EC56C916301A34">2024</mods:date>
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<mods:detail id="2E1636C3E11CDB77128595283A3CB31B" type="pubDate">
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<mods:number id="9899BD1DEA81AD1699D8007CFF6D9BD0">2024-05-15</mods:number>
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<mods:number id="FD9CE5FF073EEF30CB9FBA85ACC0BCC2">5450</mods:number>
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<mods:number id="21062D06C95D597366A0367B2102FC91">1</mods:number>
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<mods:start id="F3BED02629FD84018571AB328D612251">1</mods:start>
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<mods:location id="3DBA92A0224996208E29EDADC487162D">
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<mods:url id="E16CB66366C00953E0D5BBBA864EC7FE">http://dx.doi.org/10.11646/zootaxa.5450.1.1</mods:url>
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</mods:location>
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<mods:classification id="B811F11C1E7B9F1CD854F3B90091773D">journal article</mods:classification>
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<mods:identifier id="7105113AB847E8EFFABE4CF700A141E0" type="DOI">10.11646/zootaxa.5450.1.1</mods:identifier>
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<mods:identifier id="1285CE278B86383901A3712D58DE8695" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="8DA22A7CC6CA1740E346A8B5ABD5F132" type="Zenodo-Dep">11232500</mods:identifier>
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<mods:identifier id="A97D78114C3FB89F064980CB84AE40A0" type="ZooBank">1B5D192F-1D5B-4460-9133-9AEAE9C920BF</mods:identifier>
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<treatment id="03FD8227FFF2E30EFF78FA87FBBC8058" ID-DOI="http://doi.org/10.5281/zenodo.11232543" ID-Zenodo-Dep="11232543" LSID="urn:lsid:plazi:treatment:03FD8227FFF2E30EFF78FA87FBBC8058" httpUri="http://treatment.plazi.org/id/03FD8227FFF2E30EFF78FA87FBBC8058" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<subSubSection id="C34E60BAFFF2E30DFF78FA87FE2F8464" box="[151,410,762,789]" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="8BEB3331FFF2E30DFF78FA87FE2F8464" blockId="45.[151,410,762,825]" box="[151,410,762,789]" pageId="45" pageNumber="46">
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<heading id="D0A3845DFFF2E30DFF78FA87FE2F8464" bold="true" box="[151,410,762,789]" fontSize="11" level="2" pageId="45" pageNumber="46" reason="4">
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<emphasis id="B920EF23FFF2E30DFF78FA87FE2F8464" bold="true" box="[151,410,762,789]" pageId="45" pageNumber="46">
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||
<taxonomicName id="4C5448B2FFF2E30DFF78FA87FEE48464" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,337,762,789]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="lahceni" status="sp. nov.">
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<emphasis id="B920EF23FFF2E30DFF78FA87FEE48464" bold="true" box="[151,337,762,789]" italics="true" pageId="45" pageNumber="46">Harpes lahceni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2135258FFF2E30DFEB8FA87FE2F8464" box="[343,410,763,789]" pageId="45" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E60BAFFF2E30DFF78FB63FEAE8448" box="[151,283,798,825]" pageId="45" pageNumber="46" type="description">
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<paragraph id="8BEB3331FFF2E30DFF78FB63FEAE8448" blockId="45.[151,410,762,825]" box="[151,283,798,825]" pageId="45" pageNumber="46">
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<figureCitation id="136F2FB4FFF2E30DFF78FB63FEAE8448" box="[151,283,798,825]" captionStart="PLATE 5" captionStartId="111.[151,234,1683,1707]" captionTargetBox="[237,1351,181,1659]" captionTargetId="figure-15@111.[237,1351,181,1659]" captionTargetPageId="111" captionText="PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232547" httpUri="https://zenodo.org/record/11232547/files/figure.png" pageId="45" pageNumber="46">Plate 5A–N</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34E60BAFFF2E30DFF78FB1AFC7F8344" pageId="45" pageNumber="46" type="diagnosis">
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<paragraph id="8BEB3331FFF2E30DFF78FB1AFC7F8344" blockId="45.[151,1437,870,2013]" pageId="45" pageNumber="46">
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<emphasis id="B920EF23FFF2E30DFF78FB1AFEA784F1" bold="true" box="[151,274,870,896]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
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Wide cephalon, width (tr.) across alae about 1.4 times length (sag.). Glabella oval and strongly inflated. Brim barely sloping, slightly concave and narrowing moderately strongly posterolaterally. Alae inflated and axial furrows not extending across alae. Alar furrows become shallower distally. Fine pointed tubercles scattered along top of glabella, on dorsal surface of eye lobes and occipital ring and along dorsal surfaces of external and internal rims. Prolongations up to 15% longer (exsag.) than cephalon (sag.). In lateral view, external rim posteriorly curving upwards and internal rim wave-like in shape. Over 25 thoracic segments.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFF2E30DFF28FC43FF6F830C" pageId="45" pageNumber="46" type="etymology">
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<paragraph id="8BEB3331FFF2E30DFF28FC43FF6F830C" blockId="45.[151,1437,870,2013]" pageId="45" pageNumber="46">
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<emphasis id="B920EF23FFF2E30DFF28FC43FEFB8328" bold="true" box="[199,334,1087,1113]" pageId="45" pageNumber="46">Etymology.</emphasis>
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For Ait Hssaine Lahcen, a Moroccan trilobite worker who assisted with fieldwork in the Anti-Atlas.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFF2E30DFF28FCFAFAE68208" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="8BEB3331FFF2E30DFF28FCFAFBBE8240" blockId="45.[151,1437,870,2013]" pageId="45" pageNumber="46">
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<emphasis id="B920EF23FFF2E30DFF28FCFAFE8383D1" bold="true" box="[199,310,1158,1184]" pageId="45" pageNumber="46">Material.</emphasis>
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<materialsCitation id="3B3C396CFFF2E30DFEAEFCFBFCC383B4" collectionCode="NHMUK" collectorName="It & Section & El Otfal Formation & Eifelian & Jbel Issoumour & Dib" pageId="45" pageNumber="46" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8D93FFF2E30DFEAEFCFBFE1B83D0" box="[321,430,1159,1185]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFF2E30DFE54FCFBFD9083D0" box="[443,549,1159,1185]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="45" pageNumber="46" type="Museum">NHMUK</collectionCode>
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<collectorName id="26A156E7FFF2E30DFDDFFCFBFDF483D0" box="[560,577,1159,1185]" pageId="45" pageNumber="46">It</collectorName>
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29232,
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<figureCitation id="136F2FB4FFF2E30DFD4CFCFBFCBC83D0" box="[675,777,1158,1185]" captionStart="PLATE 5" captionStartId="111.[151,234,1683,1707]" captionTargetBox="[237,1351,181,1659]" captionTargetId="figure-15@111.[237,1351,181,1659]" captionTargetPageId="111" captionText="PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232547" httpUri="https://zenodo.org/record/11232547/files/figure.png" pageId="45" pageNumber="46">Pl. 5A–J</figureCitation>
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, from horizon 3,
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<collectorName id="26A156E7FFF2E30DFC36FCFBFB9883D0" box="[985,1069,1159,1185]" pageId="45" pageNumber="46">Section</collectorName>
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2 (
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<figureCitation id="136F2FB4FFF2E30DFBB7FCFBFB1583D0" box="[1112,1184,1159,1185]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[159,1431,194,1849]" captionTargetId="figure-15@13.[151,1435,181,1857]" captionTargetPageId="13" captionText="FIGURE 7. Section 2 Jbel Issoumour" figureDoi="http://doi.org/10.5281/zenodo.11232517" httpUri="https://zenodo.org/record/11232517/files/figure.png" pageId="45" pageNumber="46">Fig. 7</figureCitation>
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),
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<collectorName id="26A156E7FFF2E30DFB56FCFBFA2C83D0" box="[1209,1433,1159,1185]" pageId="45" pageNumber="46">El Otfal Formation</collectorName>
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,
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<collectorName id="26A156E7FFF2E30DFF78FCD7FF4683B4" box="[151,243,1195,1221]" pageId="45" pageNumber="46">Eifelian</collectorName>
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,
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<collectorName id="26A156E7FFF2E30DFEEEFCD7FE1B83B4" box="[257,430,1195,1221]" pageId="45" pageNumber="46">Jbel Issoumour</collectorName>
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||
near bou
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<collectorName id="26A156E7FFF2E30DFDC7FCD7FDE783B4" box="[552,594,1195,1221]" pageId="45" pageNumber="46">Dîb</collectorName>
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||
(
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<figureCitation id="136F2FB4FFF2E30DFD8BFCD7FD0A83B4" box="[612,703,1195,1221]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="45" pageNumber="46">Fig. 2B</figureCitation>
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, Map 7, site 6)
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</materialsCitation>
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.
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<materialsCitation id="3B3C396CFFF2E30DFC6AFCD7FBAB8398" collectionCode="NHMUK" collectorName="It & El Otfal Formation & Jbel Oufatene" pageId="45" pageNumber="46" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54EF8D93FFF2E30DFC6AFCD7FC5E83B4" box="[901,1003,1195,1221]" pageId="45" pageNumber="46" type="paratype">Paratype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFF2E30DFC16FCD7FBD683B4" box="[1017,1123,1195,1221]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="45" pageNumber="46" type="Museum">NHMUK</collectionCode>
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<collectorName id="26A156E7FFF2E30DFB82FCD7FBCB83B4" box="[1133,1150,1195,1221]" pageId="45" pageNumber="46">It</collectorName>
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29233,
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<figureCitation id="136F2FB4FFF2E30DFB30FCD7FAE283B4" box="[1247,1367,1194,1221]" captionStart="PLATE 5" captionStartId="111.[151,234,1683,1707]" captionTargetBox="[237,1351,181,1659]" captionTargetId="figure-15@111.[237,1351,181,1659]" captionTargetPageId="111" captionText="PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232547" httpUri="https://zenodo.org/record/11232547/files/figure.png" pageId="45" pageNumber="46">Pl. 5M, N</figureCitation>
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||
, from “
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<emphasis id="B920EF23FFF2E30DFF4BFCB3FEBD8398" box="[164,264,1231,1257]" italics="true" pageId="45" pageNumber="46">Phylonix</emphasis>
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||
Couche”,
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<collectorName id="26A156E7FFF2E30DFE6EFCB3FDE98398" box="[385,604,1231,1257]" pageId="45" pageNumber="46">El Otfal Formation</collectorName>
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||
,
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||
<collectorName id="26A156E7FFF2E30DFD87FCB3FCB08398" box="[616,773,1231,1257]" pageId="45" pageNumber="46">Jbel Oufatene</collectorName>
|
||
(
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||
<figureCitation id="136F2FB4FFF2E30DFCFAFCB3FCDB8398" box="[789,878,1231,1257]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="45" pageNumber="46">Fig. 2B</figureCitation>
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, Map 8, site 6)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B3C396CFFF2E30DFBC5FCB3FBB28240" collectionCode="NHMUK" collectorName="It & El Otfal Formation & Eifelian & Timerzit & All" pageId="45" pageNumber="46" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="54EF8D93FFF2E30DFBC5FCB3FB258398" box="[1066,1168,1231,1257]" pageId="45" pageNumber="46" type="paratype">Paratype</typeStatus>
|
||
:
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||
<collectionCode id="ED45ABF4FFF2E30DFB72FCB3FAB28398" box="[1181,1287,1231,1257]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="45" pageNumber="46" type="Museum">NHMUK</collectionCode>
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||
<collectorName id="26A156E7FFF2E30DFAE0FCB3FA958398" box="[1295,1312,1231,1257]" pageId="45" pageNumber="46">It</collectorName>
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||
29234,
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||
<figureCitation id="136F2FB4FFF2E30DFA91FCB3FF6F827C" captionStart="PLATE 5" captionStartId="111.[151,234,1683,1707]" captionTargetBox="[237,1351,181,1659]" captionTargetId="figure-15@111.[237,1351,181,1659]" captionTargetPageId="111" captionText="PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232547" httpUri="https://zenodo.org/record/11232547/files/figure.png" pageId="45" pageNumber="46">Pl. 5K, L</figureCitation>
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||
,
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||
<collectorName id="26A156E7FFF2E30DFF09FC8FFE0B827C" box="[230,446,1267,1293]" pageId="45" pageNumber="46">El Otfal Formation</collectorName>
|
||
,
|
||
<collectorName id="26A156E7FFF2E30DFE26FC8FFD97827C" box="[457,546,1267,1293]" pageId="45" pageNumber="46">Eifelian</collectorName>
|
||
from “
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||
<taxonomicName id="4C5448B2FFF2E30DFD9DFC8FFC96827C" authority="Couche" authorityName="Couche" box="[626,803,1267,1293]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B920EF23FFF2E30DFD9DFC8FFD71827D" box="[626,708,1267,1292]" italics="true" pageId="45" pageNumber="46">Harpes</emphasis>
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Couche
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</taxonomicName>
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” just above the
|
||
<taxonomicName id="4C5448B2FFF2E30DFC39FC8FFBDD827C" box="[982,1128,1267,1293]" class="Trilobita" family="Lichidae" genus="Acanthopyge" kingdom="Animalia" order="Lichida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EF23FFF2E30DFC39FC8FFBDD827C" box="[982,1128,1267,1293]" italics="true" pageId="45" pageNumber="46">Acanthopyge</emphasis>
|
||
</taxonomicName>
|
||
horizon,
|
||
<collectorName id="26A156E7FFF2E30DFB3DFC8FFA84827C" box="[1234,1329,1267,1293]" pageId="45" pageNumber="46">Timerzit</collectorName>
|
||
(
|
||
<figureCitation id="136F2FB4FFF2E30DFAAFFC8FFA2D827C" box="[1344,1432,1267,1293]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="45" pageNumber="46">Fig. 2B</figureCitation>
|
||
, Map 10, site 2).
|
||
<collectorName id="26A156E7FFF2E30DFEA2FD6BFEC48240" box="[333,369,1303,1329]" pageId="45" pageNumber="46">All</collectorName>
|
||
articulated dorsal exoskeletons with thorax partly enrolled
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF2E30DFF28FD47FAE68208" blockId="45.[151,1437,870,2013]" pageId="45" pageNumber="46">
|
||
Other material:
|
||
<taxonomicName id="4C5448B2FFF2E30DFE9CFD47FD3F8224" authority=", NHMUK It" authorityName="NHMUK It" box="[371,650,1339,1365]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF2E30DFE9CFD47FE3A8225" box="[371,399,1339,1364]" italics="true" pageId="45" pageNumber="46">H.</emphasis>
|
||
cf
|
||
<emphasis id="B920EF23FFF2E30DFE42FD47FE4A8224" box="[429,511,1339,1365]" italics="true" pageId="45" pageNumber="46">lahceni</emphasis>
|
||
, NHMUK It
|
||
</taxonomicName>
|
||
29235,
|
||
<figureCitation id="136F2FB4FFF2E30DFD30FD47FC8B8224" box="[735,830,1339,1365]" captionStart="PLATE 6" captionStartId="112.[151,234,1747,1771]" captionTargetBox="[213,1375,181,1723]" captionTargetId="figure-20@112.[213,1375,181,1723]" captionTargetPageId="112" captionText="PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232549" httpUri="https://zenodo.org/record/11232549/files/figure.png" pageId="45" pageNumber="46">Pl. 6I–N</figureCitation>
|
||
, from “
|
||
<taxonomicName id="4C5448B2FFF2E30DFC61FD47FBC38224" authority="Couche" authorityName="Couche" box="[910,1142,1339,1365]" class="Trilobita" family="Acastidae" genus="Erbenochile" kingdom="Animalia" order="Phacopida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EF23FFF2E30DFC61FD47FBA28224" box="[910,1047,1339,1365]" italics="true" pageId="45" pageNumber="46">Erbenochile</emphasis>
|
||
Couche
|
||
</taxonomicName>
|
||
”, Timrhanrhart Formation, Emsian,
|
||
<quantity id="4CAC9ED4FFF2E30DFF16FD23FEF58208" box="[249,320,1375,1401]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="45" pageNumber="46" unit="km" value="24.0">24 km</quantity>
|
||
S.W. of Foum Zguid (
|
||
<figureCitation id="136F2FB4FFF2E30DFDD3FD23FD208208" box="[572,661,1375,1401]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="45" pageNumber="46">Fig. 2A</figureCitation>
|
||
, Map 2, site 6). Partial dorsal exoskeleton of an upper lamella.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFF2E30EFF28FDFEFD4784F0" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
|
||
<paragraph id="8BEB3331FFF2E30DFF28FDFEFDC38190" blockId="45.[151,1437,870,2013]" pageId="45" pageNumber="46">
|
||
<emphasis id="B920EF23FFF2E30DFF28FDFEFEEC82ED" bold="true" box="[199,345,1410,1436]" pageId="45" pageNumber="46">Description.</emphasis>
|
||
In dorsal view, ovoid cephalon widest at mid-point between level of eye lobes and alae, where length (sag.) is around 70% of width (tr.). Cephalon and glabella particularly strongly vaulted. Glabella anterior to S0 more oval than cone-shaped, rounded anteriorly and with sparse tubercles along posterior two thirds of crest. S1 sloping posteriorly before curving anteriorly near crest. No furrows or visible scars at S2 and S3. Preglabellar furrow shallow and preglabellar genal area narrow, with length (sag. exsag.) ≈20% of width (tr.). Axial furrows convex, weakly convergent and not crossing junction of ala and L1. Alae are large, inflated, laterally directed and reaching more than 70% of way across (tr.) genal area. Alar furrows shallow distally. No interalar furrows or ridges but alar depressions present. Occipital furrow deep, broad and medially straight. Occipital ring narrow (sag. and exsag.) and occipital node small, set back from anterior edge of occipital ring. Posterior border very short (tr.), around 20% of width (tr.) of occipital ring.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF2E30DFF28FE97FE6580AC" blockId="45.[151,1437,870,2013]" pageId="45" pageNumber="46">
|
||
Genal area narrow, width (tr.) around 53% of length (exsag.). Eye lobes nearly conical in shape and nearly reach across genal area to near inner margin of fringe. Scattered fine tubercles on dorsal surfaces. Eye position ratio about 0.89. Two horizontal oval eye lenses face outwards anterolaterally and posterolaterally respectively (
|
||
<figureCitation id="136F2FB4FFF2E30DFB13FF4FFAF1803C" box="[1276,1348,1842,1869]" captionStart="PLATE 5" captionStartId="111.[151,234,1683,1707]" captionTargetBox="[237,1351,181,1659]" captionTargetId="figure-15@111.[237,1351,181,1659]" captionTargetPageId="111" captionText="PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232547" httpUri="https://zenodo.org/record/11232547/files/figure.png" pageId="45" pageNumber="46">Pl. 5E</figureCitation>
|
||
). Broad but low ridge from between the two lenses, reaches down onto genal area, where it narrows and forms a very faint genal ridge extending posterolaterally to cross inner margin of fringe level with anterior of ala. Genal roll convex and wide, narrowing moderately posteriorly and steep, sloping anteriorly at over
|
||
<superScript id="7C219E79FFF2E30DFBDFFFE3FBE180DD" attach="left" box="[1072,1108,1950,1977]" fontSize="6" pageId="45" pageNumber="46">600</superScript>
|
||
and laterally at over
|
||
<superScript id="7C219E79FFF2E30DFAA8FFE3FADE80DD" attach="none" box="[1351,1387,1950,1977]" fontSize="6" pageId="45" pageNumber="46">700</superScript>
|
||
. No anterior boss or girder kink.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28F8EBFE1186DC" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
Brim gently sloping anteriorly at under
|
||
<superScript id="7C219E79FFF1E30EFD7BF8EBFD0D87D5" attach="left" box="[660,696,150,177]" fontSize="6" pageId="46" pageNumber="47">100</superScript>
|
||
and with a width (sag.) of about 28% of cephalic length (sag.); brim width ratio around 0.70. Row of large perforations either side of girder. Average diameter of perforations on central part of standardised brim ≈190 μm (
|
||
<figureCitation id="136F2FB4FFF1E30EFD44F8A3FD4D8788" box="[683,760,222,249]" captionStart="PLATE 5" captionStartId="111.[151,234,1683,1707]" captionTargetBox="[237,1351,181,1659]" captionTargetId="figure-15@111.[237,1351,181,1659]" captionTargetPageId="111" captionText="PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232547" httpUri="https://zenodo.org/record/11232547/files/figure.png" pageId="46" pageNumber="47">Pl. 5H</figureCitation>
|
||
). Above girder, perforations quickly decrease in size to that of perforations on main part of genal roll, which are finer than those of brim. At inner margin of fringe, in front of glabella, a row of larger perforations about the same size as those immediately above girder. Row of coarser perforations around external margin of brim. External rim is raised and broad with dorsal rim suture located closer to outside edge than inside edge. Tubercles on dorsal surface of rim densely packed. Marginal band not visible on material presented here.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28F9CBFA398510" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
Internal rim set at about
|
||
<superScript id="7C219E79FFF1E30EFE07F9CAFDAF86B5" attach="left" box="[488,538,438,465]" fontSize="6" pageId="46" pageNumber="47">1500</superScript>
|
||
to posterior border and curving adaxially posteriorly so that distal ends point obliquely adaxially. In lateral view, profile of internal rim strongly wave-like and upturned distally. Dorsal surface of internal rim broad with densely-packed tubercles. Row of perforations immediately under internal rim larger than those on brim below. Extension of girder meeting internal rim about one third of the way down prolongation. External rim turning upwards posteriorly to meet internal rim at end of prolongation. Genal spine short and stout.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28FA17FE508464" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">The thorax has 23 thoracic segments and tapers posteriorly so that last segment is just under 40% of width (tr.) of fifth segment (widest point). Axis strongly vaulted and with axial rings becoming narrower (tr.) posteriorly, width (tr.) of last being about 40% of the first. Anterior and posterior bands of pleura narrow (exsag.) and of the same width (exsag.). Pleural furrow deep, widening distally and extending onto downturned outer portion of pleura, which has rounded distal end.</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28FB63FD4784F0" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
Pygidium (
|
||
<figureCitation id="136F2FB4FFF1E30EFEAAFB63FE328448" box="[325,391,798,825]" captionStart="PLATE 5" captionStartId="111.[151,234,1683,1707]" captionTargetBox="[237,1351,181,1659]" captionTargetId="figure-15@111.[237,1351,181,1659]" captionTargetPageId="111" captionText="PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232547" httpUri="https://zenodo.org/record/11232547/files/figure.png" pageId="46" pageNumber="47">Pl. 5J</figureCitation>
|
||
) short (sag.) with three axial rings and pleurae and a very short (sag.) terminal piece. Axial rings wide (tr.), first axial ring being over 40% of width (tr.) of anterior of pygidium. Interpleural furrows deep, pleural furrows shallow and pleurae distal ends acute.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFF1E30EFF28FBF7FBBC8058" pageId="46" pageNumber="47" type="discussion">
|
||
<paragraph id="8BEB3331FFF1E30EFF28FBF7FCD284B8" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
<emphasis id="B920EF23FFF1E30EFF28FBF7FE8E84D4" bold="true" box="[199,315,907,933]" pageId="46" pageNumber="47">Remarks.</emphasis>
|
||
<emphasis id="B920EF23FFF1E30EFEABFBF7FE4E84D4" box="[324,507,907,933]" italics="true" pageId="46" pageNumber="47">
|
||
<taxonomicName id="4C5448B2FFF1E30EFEABFBF7FE2384D5" box="[324,406,907,932]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Harpes</taxonomicName>
|
||
lahceni’
|
||
</emphasis>
|
||
s morphology complies with the genus diagnostics with the exception of the brim width ratio, which is 10% above the range proposed in
|
||
<tableCitation id="C6D6068AFFF1E30EFCEEFBD3FCD684B8" box="[769,867,942,969]" captionStart="TABLE 5" captionStartId="182.[151,239,503,529]" captionText="TABLE 5b. Proposed diagnostic characters for Devonian harpetid genera (continued)" pageId="46" pageNumber="47">Table 5a</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28FBAFFE31827C" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
<taxonomicName id="4C5448B2FFF1E30EFF28FBAFFEC7849C" box="[199,370,979,1005]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFF28FBAFFEC7849C" box="[199,370,979,1005]" italics="true" pageId="46" pageNumber="47">Harpes lahceni</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C5448B2FFF1E30EFDEBFBAFFD13849C" authorityName="Whittington" authorityYear="1950" box="[516,678,979,1005]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="whidbornei">
|
||
<emphasis id="B920EF23FFF1E30EFDEBFBAFFD13849C" box="[516,678,979,1005]" italics="true" pageId="46" pageNumber="47">H. whidbornei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFC54EC0FFF1E30EFD5BFBAFFC3E849C" author="Whittington, H. B." box="[692,907,978,1005]" pageId="46" pageNumber="47" refId="ref91104" refString="Whittington, H. B. (1950 a) British Trilobites of the order Harpidae. The Palaeontographical Society, London, 57 pp., 7 pls." type="book" year="1950">Whittington 1950a</bibRefCitation>
|
||
, pl. VII, figs 5–12) in the strong vaulting of the cephalon, the steep slope of the anterior of the genicranium and the inflated glabella. The alae of
|
||
<taxonomicName id="4C5448B2FFF1E30EFB17FB8BFA288360" authorityName="Whittington" authorityYear="1950" box="[1272,1437,1015,1041]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="whidbornei">
|
||
<emphasis id="B920EF23FFF1E30EFB17FB8BFA288360" box="[1272,1437,1015,1041]" italics="true" pageId="46" pageNumber="47">H. whidbornei</emphasis>
|
||
</taxonomicName>
|
||
are more weakly-defined and lack the inflation of those of
|
||
<taxonomicName id="4C5448B2FFF1E30EFCA9FC67FC0B8344" box="[838,958,1051,1077]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFCA9FC67FC0B8344" box="[838,958,1051,1077]" italics="true" pageId="46" pageNumber="47">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
. Also,
|
||
<emphasis id="B920EF23FFF1E30EFBFCFC67FB748344" box="[1043,1217,1051,1077]" italics="true" pageId="46" pageNumber="47">H. whidbornei’</emphasis>
|
||
s prolongations are shorter and the internal rims, which in lateral view are nearly straight, descend steeply to meet the external rims at the distal ends. The internal rims of
|
||
<taxonomicName id="4C5448B2FFF1E30EFDBFFC1FFD73830C" box="[592,710,1123,1149]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFDBFFC1FFD73830C" box="[592,710,1123,1149]" italics="true" pageId="46" pageNumber="47">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
initially arch up from the posterior border before curving down and then, towards the distal ends, curving up again forming a wave-like shape. The poor state of the
|
||
<taxonomicName id="4C5448B2FFF1E30EFB14FCFBFA2883D0" authorityName="Whittington" authorityYear="1950" box="[1275,1437,1159,1185]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="whidbornei">
|
||
<emphasis id="B920EF23FFF1E30EFB14FCFBFA2883D0" box="[1275,1437,1159,1185]" italics="true" pageId="46" pageNumber="47">H. whidbornei</emphasis>
|
||
</taxonomicName>
|
||
material makes comparisons of the anatomy of the two species difficult. The thorax of
|
||
<taxonomicName id="4C5448B2FFF1E30EFB80FCD7FAA783B4" authorityName="Whittington" authorityYear="1950" box="[1135,1298,1195,1221]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="whidbornei">
|
||
<emphasis id="B920EF23FFF1E30EFB80FCD7FAA783B4" box="[1135,1298,1195,1221]" italics="true" pageId="46" pageNumber="47">H. whidbornei</emphasis>
|
||
</taxonomicName>
|
||
is unknown and the pygidium attributed by
|
||
<bibRefCitation id="EFC54EC0FFF1E30EFE18FCB3FD088398" author="Whidborne, G. F." box="[503,701,1231,1257]" pageId="46" pageNumber="47" refId="ref90906" refString="Whidborne, G. F. (1889 - 1892) A Monograph of the Devonian Fauna of the South of England. The Palaeontology Society, London, 344 pp., xxxi pls. https: // doi. org / 10.5962 / bhl. title. 25922" type="book" year="1889">Whidborne (1889</bibRefCitation>
|
||
–1892) to
|
||
<taxonomicName id="4C5448B2FFF1E30EFCDEFCB3FC668398" authorityName="Whittington" authorityYear="1950" box="[817,979,1231,1257]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="whidbornei">
|
||
<emphasis id="B920EF23FFF1E30EFCDEFCB3FC668398" box="[817,979,1231,1257]" italics="true" pageId="46" pageNumber="47">H. whidbornei</emphasis>
|
||
</taxonomicName>
|
||
was considered doubtful by Whittington (
|
||
<bibRefCitation id="EFC54EC0FFF1E30EFF70FC8FFECD827C" author="Whittington, H. B." box="[159,376,1266,1293]" pageId="46" pageNumber="47" refId="ref91104" refString="Whittington, H. B. (1950 a) British Trilobites of the order Harpidae. The Palaeontographical Society, London, 57 pp., 7 pls." type="book" year="1950">Whittington 1950a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28FD6BFB7D8104" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
<taxonomicName id="4C5448B2FFF1E30EFF28FD6BFE8E8240" box="[199,315,1303,1329]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFF28FD6BFE8E8240" box="[199,315,1303,1329]" italics="true" pageId="46" pageNumber="47">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
is also similar to
|
||
<taxonomicName id="4C5448B2FFF1E30EFDEEFD6BFDC78240" authorityName="Johnson" authorityYear="2024" box="[513,626,1303,1329]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lentigo">
|
||
<emphasis id="B920EF23FFF1E30EFDEEFD6BFDC78240" box="[513,626,1303,1329]" italics="true" pageId="46" pageNumber="47">H. lentigo</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136F2FB4FFF1E30EFD6EFD6BFCB48240" box="[641,769,1303,1329]" captionStart="PLATE 7" captionStartId="113.[151,234,1741,1765]" captionTargetBox="[213,1374,181,1717]" captionTargetId="figure-15@113.[213,1374,181,1717]" captionTargetPageId="113" captionText="PLATE 7. A–M Harpes lentigo n. sp., from a coral horizon 5 m (12.2 m down slope) below the “Morocconites Couche”, Tazoulait Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 4). A–G dorsal exoskeleton, thorax disarticulated and partially rolled, holotype, NHMUK It 29236.A, dorsal, B, anterior, C, posterior and D, lateral view. E, lateral view of eye lobe (uncoated), F, laterodorsal view of eye lobe (uncoated) and G, dorsal view of brim perforations. H–M Harpes lentigo n. sp., plate of three disarticulated dorsal exoskeletons, paratypes, NHMUK It 29237 (1 to 3) (Fig. 2B, Map 8, site 5). H, dorsal, I, anterodorsal and J, lateral view. K, dorsal view of brim perforations, L, lateral view of pleurae and M, anterior view of rim. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232553" httpUri="https://zenodo.org/record/11232553/files/figure.png" pageId="46" pageNumber="47">Pl. F7A–M</figureCitation>
|
||
) but it has: a glabella that is inset with much finer tubercles a furrow at S1 curving around the top of the muscle insertion area; a preglabellar furrow that in anterior view is straight rather than being bowed up; a broader occipital furrow; axial furrows that do not cross the alae and alae that are wider, more inflated with no interalar furrows; a narrower genal area with scattered fine tubercles but no caeca or pits; larger eye lobes that are cone shaped rather than oval; an inner margin of the fringe the course of which across the anterior of the genal areas is straight not convex; no anterior boss or girder kink; a genal roll with perforations that are the same size as those on the brim not smaller as in the case of
|
||
<taxonomicName id="4C5448B2FFF1E30EFC27FD93FB8C8178" authorityName="Johnson" authorityYear="2024" box="[968,1081,1519,1545]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lentigo">
|
||
<emphasis id="B920EF23FFF1E30EFC27FD93FB8C8178" box="[968,1081,1519,1545]" italics="true" pageId="46" pageNumber="47">H. lentigo</emphasis>
|
||
</taxonomicName>
|
||
; a brim that slopes less steeply and has a higher brim width ratio, no caeca and perforations that do not become smaller away from the girder; an external rim that has fine tubercles on its dorsal surface; prolongations that are longer and in lateral view have a strong wave like profile; a thorax that is more tapered, inner portion of pleura in line with outer.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28FE03FF5481CC" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
A comparison between
|
||
<taxonomicName id="4C5448B2FFF1E30EFE25FE03FD8981E8" box="[458,572,1663,1689]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFE25FE03FD8981E8" box="[458,572,1663,1689]" italics="true" pageId="46" pageNumber="47">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
and its sister species,
|
||
<taxonomicName id="4C5448B2FFF1E30EFCC3FE03FC0381E8" authorityName="Johnson" authorityYear="2024" box="[812,950,1663,1689]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFF1E30EFCC3FE03FC0381E8" box="[812,950,1663,1689]" italics="true" pageId="46" pageNumber="47">H. prescheri</emphasis>
|
||
</taxonomicName>
|
||
, is given in the systematics for
|
||
<taxonomicName id="4C5448B2FFF1E30EFAE3FE03FA2381E8" authorityName="Johnson" authorityYear="2024" box="[1292,1430,1663,1689]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="prescheri">
|
||
<emphasis id="B920EF23FFF1E30EFAE3FE03FA2381E8" box="[1292,1430,1663,1689]" italics="true" pageId="46" pageNumber="47">H. prescheri</emphasis>
|
||
</taxonomicName>
|
||
, below.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF1E30EFF28FEBBFBBC8058" blockId="46.[151,1437,150,1833]" pageId="46" pageNumber="47">
|
||
<taxonomicName id="4C5448B2FFF1E30EFF28FEBBFE2F8190" box="[199,410,1735,1761]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFF28FEBBFEAC8191" box="[199,281,1735,1760]" italics="true" pageId="46" pageNumber="47">Harpes</emphasis>
|
||
cf.
|
||
<emphasis id="B920EF23FFF1E30EFEA7FEBBFE2F8190" box="[328,410,1735,1761]" italics="true" pageId="46" pageNumber="47">lahceni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136F2FB4FFF1E30EFE43FEBBFDAB8190" box="[428,542,1735,1761]" captionStart="PLATE 6" captionStartId="112.[151,234,1747,1771]" captionTargetBox="[213,1375,181,1723]" captionTargetId="figure-20@112.[213,1375,181,1723]" captionTargetPageId="112" captionText="PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232549" httpUri="https://zenodo.org/record/11232549/files/figure.png" pageId="46" pageNumber="47">Pl. F6I–N</figureCitation>
|
||
) differs from
|
||
<taxonomicName id="4C5448B2FFF1E30EFD50FEBBFC808190" box="[703,821,1735,1761]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFD50FEBBFC808190" box="[703,821,1735,1761]" italics="true" pageId="46" pageNumber="47">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
in having a brim that is significantly wider at S0 and standardised brim perforations that are over 40% larger. The axial furrow is expressed at L1, the alae are less inflated and there are interalar furrows and ridges not present on the alae of
|
||
<taxonomicName id="4C5448B2FFF1E30EFC62FF73FBB48058" box="[909,1025,1807,1833]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lahceni">
|
||
<emphasis id="B920EF23FFF1E30EFC62FF73FBB48058" box="[909,1025,1807,1833]" italics="true" pageId="46" pageNumber="47">H. lahceni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |