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<document id="70E495557E0E72EC30A246D1A1B10E0C" ID-CLB-Dataset="30163" ID-DOI="10.11646/zootaxa.4415.1.1" ID-GBIF-Dataset="c803bce9-4469-43db-8acd-8291d035b6a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1241736" ID-ZooBank="5DAA824F-BCBD-47FF-9948-F7EC45829AEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525594886833" checkinUser="plazi" docAuthor="Wells, Alice &amp; Contents, Arturs Neboiss Table Of" docDate="2018" docId="03D6576233463F28908B263F9D4FBD32" docLanguage="en" docName="zootaxa.4415.1.1.pdf.imf" docOrigin="Zootaxa 4415 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcyphysa Wells &amp; Contents 2018, gen. nov." docType="treatment" docVersion="7" lastPageNumber="24" masterDocId="FFEF2F1A33533F3F901C23329F53B94F" masterDocTitle="Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus" masterLastPageNumber="44" masterPageNumber="1" pageNumber="22" updateTime="1698541036082" updateUser="ExternalLinkService">
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<mods:title id="98686151535C5C568597FFA7EA6F31CF">Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus</mods:title>
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<mods:namePart id="77F061ACD8C6EDEDD4DBE69292BC0891">Wells, Alice</mods:namePart>
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<mods:title id="D888F635C28F897C4F1D8D6A5FE3D83B">Zootaxa</mods:title>
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<mods:date id="33DA4D8AE726F5D605FFEF7B8A33B0F8">2018</mods:date>
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<treatment id="03D6576233463F28908B263F9D4FBD32" ID-DOI="http://doi.org/10.5281/zenodo.5966225" ID-GBIF-Taxon="143811135" ID-Zenodo-Dep="5966225" LSID="urn:lsid:plazi:treatment:03D6576233463F28908B263F9D4FBD32" httpUri="http://treatment.plazi.org/id/03D6576233463F28908B263F9D4FBD32" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection id="C365B5FF33463F2A908B263F9E10BC07" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BC0E67433463F2A908B263F9ED8BC68" blockId="21.[151,395,1293,1352]" box="[151,395,1293,1319]" pageId="21" pageNumber="22">
<heading id="D088511833463F2A908B263F9ED8BC68" bold="true" box="[151,395,1293,1319]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="B90B3A6633463F2A908B263F9ED8BC68" bold="true" box="[151,395,1293,1319]" pageId="21" pageNumber="22">
<taxonomicName id="4C7F9DF733463F2A908B263F9E45BC68" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[151,278,1293,1319]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B3A6633463F2A908B263F9E45BC68" bold="true" box="[151,278,1293,1319]" italics="true" pageId="21" pageNumber="22">Arcyphysa</emphasis>
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<taxonomicNameLabel id="A238871D33463F2A9101263F9ED8BC68" box="[285,395,1293,1319]" pageId="21" pageNumber="22" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="8BC0E67433463F2A908B261D9E10BC07" blockId="21.[151,395,1293,1352]" box="[151,323,1327,1352]" pageId="21" pageNumber="22">
(
<figureCitation id="1344FAF133463F2A9083261D9E69BC07" box="[159,314,1327,1352]" captionStart-0="FIGURES 102109" captionStart-1="FIGURES 110116" captionStart-2="FIGURES 117131" captionStart-3="FIGURES 132145" captionStart-4="FIGURES 146163" captionStart-5="FIGURES 164178" captionStart-6="FIGURES 179187" captionStartId-0="27.[151,264,1729,1752]" captionStartId-1="32.[151,264,1363,1386]" captionStartId-2="35.[151,264,1806,1829]" captionStartId-3="38.[151,264,1817,1840]" captionStartId-4="39.[151,264,1675,1698]" captionStartId-5="40.[151,264,1776,1799]" captionStartId-6="41.[151,264,1053,1076]" captionTargetBox-0="[150,1435,225,1691]" captionTargetBox-1="[155,1436,361,1352]" captionTargetBox-2="[223,1405,189,1778]" captionTargetBox-3="[151,1434,193,1797]" captionTargetBox-4="[151,1436,209,1614]" captionTargetBox-5="[151,1436,194,1751]" captionTargetBox-6="[151,1436,291,1027]" captionTargetId-0="figure@27.[150,1436,193,1708]" captionTargetId-1="figure@32.[151,1436,331,1352]" captionTargetId-2="figure@35.[176,1412,185,1793]" captionTargetId-3="figure@38.[151,1435,193,1797]" captionTargetId-4="figure@39.[151,1436,209,1620]" captionTargetId-5="figure@40.[151,1436,193,1756]" captionTargetId-6="figure@41.[151,1436,291,1032]" captionTargetPageId-0="27" captionTargetPageId-1="32" captionTargetPageId-2="35" captionTargetPageId-3="38" captionTargetPageId-4="39" captionTargetPageId-5="40" captionTargetPageId-6="41" captionText-0="FIGURES 102109. Arcyphysa gen. nov. spp., diagnostic features.102103, Arc.diamontona sp. nov., Vic., Diamond Creek, 7 km SE of Gembrook [PT-1006], ♂: 102, right fore- and hind wings, dorsal, 103, maxillary palp. 104109, ♂ genitalia. 104 105, Arc. angusta (Banks 1939; NSW, Styx R. [PT-1005]): 104, ventral, and 105, left lateral. 106107, Arc. diamontona sp. nov. [PT-1006]:106, ventral, and 107, left lateral.108109, Arc.fraserensis sp. nov. (Qld, Fraser Island, Pile Valley, nr Central Station [PT-1125]):108, ventral,and 109, leftlateral.Scale bars: Fig.102 = 1 mm; Fig.103 = 0.5 mm.Abbreviations:ac= anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; fI, fII, fIII, fIV, fV = forks IV; mc= median cell; har = harpago; ph = phallic apparatus; IX, X = abdominal segments IX, X" captionText-1="FIGURES 110116. Arcyphysa gen. nov. spp., diagnostic features. 110111, Arc. hystricosa (Neboiss 1979) (Qld, North Stradbroke Island [PT-512]): 110, ventral; 111, left lateral. 112, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek), ventral. 113115, Arc.candela sp. nov. (NSW, Brown Mountain [PT-1017]): 113, left lateral; 114, ventral; 115, dorsal. 116, Arc. diamontona sp. nov. (Vic., Diamond Creek, 7 km SE of Gembrook [PT-1047]), ♀ terminal abdominal segments, left lateral." captionText-2="FIGURES 117131. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 117118, Arc. caldera sp. nov. (Qld, The Crater,Atherton [PT-1040]):117, ventral; 118, leftlateral.119120, Arc.crescentinasp.nov. (Qld, TullyFalls [PT-1002]):119, ventral; 120, left lateral. 121122, Arc.hugginsi sp. nov. (Qld, Bellenden Ker Range [PT-1000]): 121, ventral; 122, left lateral. 123125, Arc. sybillae sp. nov. (Qld, Bloomfield River [PT-1016]): 123, ventral; 124, left lateral; 125, dorsal. 126127, Arc. carnarvona sp. nov. (holotype; Qld, Carnarvon Gorge National Park): 126, ventral; 127, left lateral. 128129, Arc. naumanni sp. nov. (holotype; Qld, 11 km W by N Bald Hill, McIllraith Range): 128, ventral; 129, left lateral. 130131, Arc.lovedayi sp. nov. (holotype): 130, ventral; 131, left lateral." captionText-3="FIGURES 132145. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 132133, Arc. acmea sp. nov. (Qld, Mt Bartle Frere, 0.5km N of South Peak [PT-999]): 132, ventral;133, left lateral.134135, Arc.anaplasis sp. nov. (Qld, 25 km along Mt Lewis Road, SW of Mossman [PT-1003]): 134, ventral; 135, left lateral. 136137, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]): 136, ventral; 137, left lateral. 138140, Arc. flinti sp. nov. (Qld, Palmerston National Park, Tchooratippa Creek [PT-2007]): 138, ventral; 139, left lateral; 140, dorsal. 141142, Arc. volsella sp. nov. (Qld, Queensland, Lamington National Park, Coomera Creek [PT-1013]): 141, ventral; 142, left lateral.143145, Arc. nebo sp. nov. (Qld, Mt Nebo [PT-1096]): 143, dorsal;144, left lateral; 145, ventral." captionText-4="FIGURES 146163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154163, ♂ genitalia. 154155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X." captionText-5="FIGURES 164178. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 164165, Arc. lovedayi sp. nov. (holotype): 164, ventral; 165, left lateral. 166167, Arc. crescentina sp. nov. (Qld, Tully Falls [PT-1002]): 166, ventral; 167, left lateral. 168169, Arc. hugginsi sp. nov. (Qld, Bellenden Ker Range [PT-1000]): 168, ventral; 169, left lateral. 170, Arc. sybillae sp. nov. (Qld, Bloomfield River [PT-1016]), ventral. 171173, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge): 171, ventral; 172, left lateral;173, dorsal. 174175, Arc.anaplasis sp. nov. (Qld, 25 km along Mt LewisRoad, SW of Mossman [PT-1003]): 174, ventral; 175, left lateral.176, Arc.acmea sp. nov. (Qld, Mt Bartle Frere, 0.5 km N of South Peak [PT-999]), ventral.177 178, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]): 177, ventral; 178, left lateral." captionText-6="FIGURES 179187. Arcyphysa gen. nov. spp., diagnostic features. 179182, ♀ terminal abdominal segments: 179180, Arc. angusta (NSW, Styx R. [PT-1005]): 179, ventral, and 180, left lateral; 181182, Arc. diamontona sp. nov. [PT-1006]: 181, ventral, and 182, left lateral. 183187, ♂ reticulate-walled internal sacs of abdominal segments VI and VII: 183, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]), ventral; 184185, Arc. acmea sp. nov. (Qld, Mt Bartle Frere,0.5 km Nof South Peak [PT-999]):184, rightlateral, and 185, ventral;186, Arc.anaplasis sp.nov. (Qld, 25 km along Mt Lewis Road, SW of Mossman [PT-1003]), ventral; 187, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]), ventral. Abbreviation:VIII = abdominal segment VIII." httpUri-0="https://zenodo.org/record/1241754/files/figure.png" httpUri-1="https://zenodo.org/record/1241756/files/figure.png" httpUri-2="https://zenodo.org/record/1241758/files/figure.png" httpUri-3="https://zenodo.org/record/1241760/files/figure.png" httpUri-4="https://zenodo.org/record/1241762/files/figure.png" httpUri-5="https://zenodo.org/record/1241764/files/figure.png" httpUri-6="https://zenodo.org/record/1241766/files/figure.png" pageId="21" pageNumber="22">Figs 102187</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C365B5FF33463F2A908B264B9C3FBCC1" box="[151,876,1400,1422]" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BC0E67433463F2A908B264B9C3FBCC1" blockId="21.[151,876,1400,1422]" box="[151,876,1400,1422]" pageId="21" pageNumber="22">
Type species:
<taxonomicName id="4C7F9DF733463F2A913A264A9D25BCC1" authority="Banks 1939" authorityName="Banks" authorityYear="1939" box="[294,630,1400,1422]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="angusta">
<emphasis id="B90B3A6633463F2A913A264A9EA6BCC1" box="[294,501,1400,1422]" italics="true" pageId="21" pageNumber="22">Diplectrona angusta</emphasis>
<bibRefCitation id="EFEE9B8533463F2A91E0264A9D25BCC1" box="[508,630,1400,1422]" pageId="21" pageNumber="22" refId="ref34905" refString="Banks, N. (1939) Trichoptera. Bulletin of the Museum of Comparative Zoology, Harvard, 85, 439 - 504." type="journal article">Banks 1939</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C365B5FF33463F2A908B26889DAABF70" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BC0E67433463F2A908B26889DAABF70" blockId="21.[151,1437,1466,1890]" pageId="21" pageNumber="22">
<emphasis id="B90B3A6633463F2A908B26889E42BC9C" bold="true" box="[151,273,1466,1491]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
This genus is distinguished from other Australian
<taxonomicName id="4C7F9DF733463F2A937B26889B5EBC9C" authorityName="Ulmer" authorityYear="1951" box="[871,1037,1466,1491]" class="Insecta" family="Hydropsychidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Diplectroninae">Diplectroninae</taxonomicName>
genera by the sharply and almost evenly divided anterolateral dorsal head warts, divided at right angles to the length of the body by a distinct suture, and in male genitalia abdominal sternite IX has one or more elongate, often elaborate, processes apicomedially, and a short, sometimes divided, harpago on each gonopod.
</paragraph>
</subSubSection>
<subSubSection id="C365B5FF33463F2990DB25789D28BEBA" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="8BC0E67433463F2A90DB25789CBFBE2D" blockId="21.[151,1437,1466,1890]" pageId="21" pageNumber="22">
<emphasis id="B90B3A6633463F2A90DB25789E0ABF2C" bold="true" box="[199,345,1610,1635]" pageId="21" pageNumber="22">Description.</emphasis>
Head (
<figureCitation id="1344FAF133463F2A91B325789D1DBF2C" box="[431,590,1610,1635]" captionStart="FIGURES 146163" captionStartId="39.[151,264,1675,1698]" captionTargetBox="[151,1436,209,1614]" captionTargetId="figure@39.[151,1436,209,1620]" captionTargetPageId="39" captionText="FIGURES 146163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154163, ♂ genitalia. 154155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X." httpUri="https://zenodo.org/record/1241762/files/figure.png" pageId="21" pageNumber="22">Figs 146150</figureCitation>
) dorsally with anterolateral warts divided obliquely, both parts similar in size, posterior parts not reaching midline; sutures pronounced. Antennae slender, about as long as forewings, segments cylindrical, with median suture and setae giving crenate appearance (
<figureCitation id="1344FAF133463F2A943025A09BC1BFE4" box="[1068,1170,1682,1707]" captionStart="FIGURES 146163" captionStartId="39.[151,264,1675,1698]" captionTargetBox="[151,1436,209,1614]" captionTargetId="figure@39.[151,1436,209,1620]" captionTargetPageId="39" captionText="FIGURES 146163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154163, ♂ genitalia. 154155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X." httpUri="https://zenodo.org/record/1241762/files/figure.png" pageId="21" pageNumber="22">Fig. 152</figureCitation>
). Maxillary palps (
<figureCitation id="1344FAF133463F2A957325A09F91BF80" captionStart="FIGURES 102109" captionStartId="27.[151,264,1729,1752]" captionTargetBox="[150,1435,225,1691]" captionTargetId="figure@27.[150,1436,193,1708]" captionTargetPageId="27" captionText="FIGURES 102109. Arcyphysa gen. nov. spp., diagnostic features.102103, Arc.diamontona sp. nov., Vic., Diamond Creek, 7 km SE of Gembrook [PT-1006], ♂: 102, right fore- and hind wings, dorsal, 103, maxillary palp. 104109, ♂ genitalia. 104 105, Arc. angusta (Banks 1939; NSW, Styx R. [PT-1005]): 104, ventral, and 105, left lateral. 106107, Arc. diamontona sp. nov. [PT-1006]:106, ventral, and 107, left lateral.108109, Arc.fraserensis sp. nov. (Qld, Fraser Island, Pile Valley, nr Central Station [PT-1125]):108, ventral,and 109, leftlateral.Scale bars: Fig.102 = 1 mm; Fig.103 = 0.5 mm.Abbreviations:ac= anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; fI, fII, fIII, fIV, fV = forks IV; mc= median cell; har = harpago; ph = phallic apparatus; IX, X = abdominal segments IX, X" httpUri="https://zenodo.org/record/1241754/files/figure.png" pageId="21" pageNumber="22">Fig. 103</figureCitation>
) with segments 2 and 3 about equal length, almost
<date id="FFC1C0B433463F2A92E325859C48BF80" box="[767,795,1719,1743]" pageId="21" pageNumber="22">2x</date>
length of segment 1, and about 1.3x length of segment 4, segment 5 exceeding length of all other segments together. Wing venation (
<figureCitation id="1344FAF133463F2A93EE25E89B0BBFBC" box="[1010,1112,1754,1779]" captionStart="FIGURES 102109" captionStartId="27.[151,264,1729,1752]" captionTargetBox="[150,1435,225,1691]" captionTargetId="figure@27.[150,1436,193,1708]" captionTargetPageId="27" captionText="FIGURES 102109. Arcyphysa gen. nov. spp., diagnostic features.102103, Arc.diamontona sp. nov., Vic., Diamond Creek, 7 km SE of Gembrook [PT-1006], ♂: 102, right fore- and hind wings, dorsal, 103, maxillary palp. 104109, ♂ genitalia. 104 105, Arc. angusta (Banks 1939; NSW, Styx R. [PT-1005]): 104, ventral, and 105, left lateral. 106107, Arc. diamontona sp. nov. [PT-1006]:106, ventral, and 107, left lateral.108109, Arc.fraserensis sp. nov. (Qld, Fraser Island, Pile Valley, nr Central Station [PT-1125]):108, ventral,and 109, leftlateral.Scale bars: Fig.102 = 1 mm; Fig.103 = 0.5 mm.Abbreviations:ac= anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; fI, fII, fIII, fIV, fV = forks IV; mc= median cell; har = harpago; ph = phallic apparatus; IX, X = abdominal segments IX, X" httpUri="https://zenodo.org/record/1241754/files/figure.png" pageId="21" pageNumber="22">Figs 102</figureCitation>
,
<figureCitation id="1344FAF133463F2A947A25E89BC2BFBC" box="[1126,1169,1754,1779]" captionStart="FIGURES 146163" captionStartId="39.[151,264,1675,1698]" captionTargetBox="[151,1436,209,1614]" captionTargetId="figure@39.[151,1436,209,1620]" captionTargetPageId="39" captionText="FIGURES 146163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154163, ♂ genitalia. 154155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X." httpUri="https://zenodo.org/record/1241762/files/figure.png" pageId="21" pageNumber="22">151</figureCitation>
) as in
<emphasis id="B90B3A6633463F2A94C325E99ACFBFBC" box="[1247,1436,1755,1779]" italics="true" pageId="21" pageNumber="22">
Arc.
<taxonomicName id="4C7F9DF733463F2A950A25E99ACFBFBC" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1302,1436,1755,1779]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="diamontona" status="sp. nov.">diamontona</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33463F2A908B25CC9FBCBE59" box="[151,239,1790,1815]" pageId="21" pageNumber="22" rank="species">
<emphasis id="B90B3A6633463F2A908B25CC9FBABE58" bold="true" box="[151,233,1790,1815]" pageId="21" pageNumber="22">sp. nov</emphasis>
.
</taxonomicNameLabel>
: Forewing fork 2 sessile; median cell length about 1.2x length of discoidal cell. Hind wing f1, f2, f3, and f5 present, median cell open; R1 curving strongly towards margin, as in
<taxonomicName id="4C7F9DF733463F2A93B724119B62BE74" authorityName="Wells &amp; Contents" authorityYear="2018" box="[939,1073,1827,1851]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B90B3A6633463F2A93B724119B62BE74" box="[939,1073,1827,1851]" italics="true" pageId="21" pageNumber="22">Diplectrona</emphasis>
</taxonomicName>
. Lateral filaments on sternite V present in both sexes, no more than half length of segment, often very short.
</paragraph>
<caption id="DF00B6FC33453F29908B26F09B44BFDC" httpUri="https://zenodo.org/record/1241750/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,264,1474,1497]" targetBox="[160,1424,193,1464]" targetPageId="22">
<paragraph id="8BC0E67433453F29908B26F09B44BFDC" blockId="22.[151,1437,1474,1686]" pageId="22" pageNumber="23">
<emphasis id="B90B3A6633453F29908B26F09E1EBC97" bold="true" box="[151,333,1474,1497]" pageId="22" pageNumber="23">FIGURES 7083</emphasis>
.
<taxonomicName id="4C7F9DF733453F29914626F19D33BC96" authority="Banks 1939" authorityName="Banks" authorityYear="1939" box="[346,608,1475,1497]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B90B3A6633453F29914626F19EB1BC96" box="[346,482,1475,1497]" italics="true" pageId="22" pageNumber="23">Austropsyche</emphasis>
<bibRefCitation id="EFEE9B8533453F2991F426F19D33BC96" box="[488,608,1475,1497]" pageId="22" pageNumber="23" refId="ref34905" refString="Banks, N. (1939) Trichoptera. Bulletin of the Museum of Comparative Zoology, Harvard, 85, 439 - 504." type="journal article">Banks 1939</bibRefCitation>
</taxonomicName>
spp.,
<specimenCount id="9D792DFD33453F29928026F69DFCBC96" box="[668,687,1476,1497]" pageId="22" pageNumber="23" type="male"></specimenCount>
diagnostic features.
<emphasis id="B90B3A6633453F29939D26F19C92BC97" bold="true" box="[897,961,1475,1496]" pageId="22" pageNumber="23">7075</emphasis>
, heads, dorsal:
<emphasis id="B90B3A6633453F29944326F19B2BBC97" bold="true" box="[1119,1144,1475,1496]" pageId="22" pageNumber="23">70</emphasis>
,
<taxonomicName id="4C7F9DF733453F29949926F19ACBBC96" authority="Banks 1939" authorityName="Banks. Diagnoses" authorityYear="1939" box="[1157,1432,1475,1497]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="victoriana">
<emphasis id="B90B3A6633453F29949926F19A45BC96" box="[1157,1302,1475,1497]" italics="true" pageId="22" pageNumber="23">Au. victoriana</emphasis>
<bibRefCitation id="EFEE9B8533453F29950026F19ACBBC96" box="[1308,1432,1475,1497]" pageId="22" pageNumber="23" refId="ref34905" refString="Banks, N. (1939) Trichoptera. Bulletin of the Museum of Comparative Zoology, Harvard, 85, 439 - 504." type="journal article">Banks 1939</bibRefCitation>
</taxonomicName>
;
<emphasis id="B90B3A6633453F29908B26D39FE3BCB9" bold="true" box="[151,176,1505,1526]" pageId="22" pageNumber="23">71</emphasis>
,
<taxonomicName id="4C7F9DF733453F2990A126D39EBCBCB8" authority="(Kimmins 1953)" authorityName="Kimmins" authorityYear="1953" baseAuthorityName="Kimmins, in Mosely &amp; Kimmins" baseAuthorityYear="1953" box="[189,495,1505,1527]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bifurcata">
<emphasis id="B90B3A6633453F2990A126D39E6FBCB8" box="[189,316,1505,1527]" italics="true" pageId="22" pageNumber="23">Au.bifurcata</emphasis>
(
<bibRefCitation id="EFEE9B8533453F29915626D39EB4BCB8" box="[330,487,1505,1527]" pageId="22" pageNumber="23" refId="ref35807" refString="Mosely, M. E. &amp; Kimmins, D. E. (1953) The Trichoptera (Caddis-flies) of Australia and New Zealand. British Museum (Natural History), London, 550 pp. https: // doi. org / 10.5962 / bhl. title. 118696" type="book">Kimmins 1953</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B90B3A6633453F2991E626D39D40BCB9" bold="true" box="[506,531,1505,1526]" pageId="22" pageNumber="23">72</emphasis>
,
<taxonomicName id="4C7F9DF733453F29923C26D09DCABCB8" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[544,665,1505,1527]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kaputar" status="sp. nov.">
<emphasis id="B90B3A6633453F29923C26D09DCABCB8" box="[544,665,1505,1527]" italics="true" pageId="22" pageNumber="23">Au. kaputar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F29928326D09DA2BCB8" box="[671,753,1506,1527]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F29928326D09DB9BCB8" bold="true" box="[671,746,1506,1527]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
;
<emphasis id="B90B3A6633453F2992E326D39C4BBCB9" bold="true" box="[767,792,1505,1526]" pageId="22" pageNumber="23">73</emphasis>
,
<taxonomicName id="4C7F9DF733453F29933926D09CD4BCB8" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[805,903,1506,1527]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuta" status="sp. nov.">
<emphasis id="B90B3A6633453F29933926D09CD4BCB8" box="[805,903,1506,1527]" italics="true" pageId="22" pageNumber="23">Au. acuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F29939226D09C8CBCB8" box="[910,991,1506,1527]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F29939226D09C89BCB8" bold="true" box="[910,986,1506,1527]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
; 74,
<taxonomicName id="4C7F9DF733453F29940826D09BC4BCB8" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1044,1175,1505,1527]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ambigua" status="sp. nov.">
<emphasis id="B90B3A6633453F29940826D09BC4BCB8" box="[1044,1175,1505,1527]" italics="true" pageId="22" pageNumber="23">Au. ambigua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F29948126D09BBCBCB8" box="[1181,1263,1506,1527]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F29948126D09BBABCB8" bold="true" box="[1181,1257,1506,1527]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
;
<emphasis id="B90B3A6633453F2994E126D09A45BCB8" bold="true" box="[1277,1302,1506,1527]" pageId="22" pageNumber="23">75</emphasis>
,
<taxonomicName id="4C7F9DF733453F29953826D09ACFBCB8" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1316,1436,1506,1527]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="morana" status="sp. nov.">
<emphasis id="B90B3A6633453F29953826D09ACFBCB8" box="[1316,1436,1506,1527]" italics="true" pageId="22" pageNumber="23">Au. morana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F29908B25329FB4BF59" box="[151,231,1536,1558]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F29908B25329FB2BF5A" bold="true" box="[151,225,1536,1557]" pageId="22" pageNumber="23">
sp.
<date id="FFC1C0B433453F2990A025329FB2BF5A" box="[188,225,1536,1557]" pageId="22" pageNumber="23" value="1976-11">nov</date>
</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B90B3A6633453F2990F225329E7DBF5A" bold="true" box="[238,302,1536,1557]" pageId="22" pageNumber="23">7679</emphasis>
, right fore- and hind wings, dorsal:
<emphasis id="B90B3A6633453F29928125329DE5BF5A" bold="true" box="[669,694,1536,1557]" pageId="22" pageNumber="23">76</emphasis>
,
<taxonomicName id="4C7F9DF733453F2992D825339C07BF59" authorityName="Banks. Diagnoses" authorityYear="1939" box="[708,852,1537,1558]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="victoriana">
<emphasis id="B90B3A6633453F2992D825339C07BF59" box="[708,852,1537,1558]" italics="true" pageId="22" pageNumber="23">Au. victoriana</emphasis>
</taxonomicName>
;
<emphasis id="B90B3A6633453F29937E25329C28BF5A" bold="true" box="[866,891,1536,1557]" pageId="22" pageNumber="23">77</emphasis>
,
<taxonomicName id="4C7F9DF733453F29939525339B52BF59" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[905,1025,1536,1558]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kaputar" status="sp. nov.">
<emphasis id="B90B3A6633453F29939525339B52BF59" box="[905,1025,1536,1558]" italics="true" pageId="22" pageNumber="23">Au. kaputar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F29941B25329B0BBF59" box="[1031,1112,1536,1558]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F29941B25329B01BF5A" bold="true" box="[1031,1106,1536,1557]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
;
<emphasis id="B90B3A6633453F29947A25329B2CBF5A" bold="true" box="[1126,1151,1536,1557]" pageId="22" pageNumber="23">78</emphasis>
,
<taxonomicName id="4C7F9DF733453F29949025339BBDBF59" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1164,1262,1537,1558]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuta" status="sp. nov.">
<emphasis id="B90B3A6633453F29949025339BBDBF59" box="[1164,1262,1537,1558]" italics="true" pageId="22" pageNumber="23">Au. acuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F2994E825329A16BF59" box="[1268,1349,1536,1558]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F2994E825329A6CBF5A" bold="true" box="[1268,1343,1536,1557]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
;
<emphasis id="B90B3A6633453F29954F25329A20BF5A" bold="true" box="[1363,1395,1536,1557]" pageId="22" pageNumber="23">79,</emphasis>
<taxonomicName id="4C7F9DF733453F29956525339FB5BF7A" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="morana" status="sp. nov.">
<emphasis id="B90B3A6633453F29956525339FB5BF7A" italics="true" pageId="22" pageNumber="23">Au. morana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F2990ED252D9E15BF7B" box="[241,326,1567,1588]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F2990ED252D9E12BF7B" bold="true" box="[241,321,1567,1588]" pageId="22" pageNumber="23">
sp.
<date id="FFC1C0B433453F299107252D9E12BF7B" box="[283,321,1567,1588]" pageId="22" pageNumber="23" value="1980-11">nov</date>
</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B90B3A6633453F29914E252D9E38BF7B" bold="true" box="[338,363,1567,1588]" pageId="22" pageNumber="23">80</emphasis>
,
<taxonomicName id="4C7F9DF733453F299161252D9EB0BF7A" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[381,483,1567,1589]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuta" status="sp. nov.">
<emphasis id="B90B3A6633453F299161252D9ECABF7B" box="[381,409,1567,1588]" italics="true" pageId="22" pageNumber="23">Au</emphasis>
.
<emphasis id="B90B3A6633453F2991B725129EB0BF7A" box="[427,483,1568,1589]" italics="true" pageId="22" pageNumber="23">acuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F2991F2252D9D17BF7B" box="[494,580,1567,1588]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F2991F2252D9D6DBF7B" bold="true" box="[494,574,1567,1588]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
, antennal segments.
<emphasis id="B90B3A6633453F29933B252D9C13BF7B" bold="true" box="[807,832,1567,1588]" pageId="22" pageNumber="23">81</emphasis>
,
<taxonomicName id="4C7F9DF733453F29934E252D9CEBBF7A" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[850,952,1567,1589]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuta" status="sp. nov.">
<emphasis id="B90B3A6633453F29934E252D9C3DBF7B" box="[850,878,1567,1588]" italics="true" pageId="22" pageNumber="23">Au</emphasis>
.
<emphasis id="B90B3A6633453F29939C25129CEBBF7A" box="[896,952,1568,1589]" italics="true" pageId="22" pageNumber="23">acuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F2993D8252D9B4ABF7B" box="[964,1049,1567,1588]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F2993D8252D9B40BF7B" bold="true" box="[964,1043,1567,1588]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
, maxillary palp.
<emphasis id="B90B3A6633453F2994C8252D9BBEBF7B" bold="true" box="[1236,1261,1567,1588]" pageId="22" pageNumber="23">82</emphasis>
,
<taxonomicName id="4C7F9DF733453F29951C252D9AC6BF7A" authorityName="Banks. Diagnoses" authorityYear="1939" box="[1280,1429,1567,1589]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="victoriana">
<emphasis id="B90B3A6633453F29951C252D9A4FBF7B" box="[1280,1308,1567,1588]" italics="true" pageId="22" pageNumber="23">Au</emphasis>
.
<emphasis id="B90B3A6633453F29953125129AC6BF7A" box="[1325,1429,1568,1589]" italics="true" pageId="22" pageNumber="23">victoriana</emphasis>
</taxonomicName>
, reticulate-walled internal sacs of abdominal segments VI and VII, ventral.
<emphasis id="B90B3A6633453F2993AD250F9C99BF1D" bold="true" box="[945,970,1597,1618]" pageId="22" pageNumber="23">83</emphasis>
,
<taxonomicName id="4C7F9DF733453F2993C7250C9B6CBF1C" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[987,1087,1598,1619]" class="Insecta" family="Hydropsychidae" genus="Austropsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuta" status="sp. nov.">
<emphasis id="B90B3A6633453F2993C7250C9CA4BF1C" box="[987,1015,1598,1619]" italics="true" pageId="22" pageNumber="23">Au</emphasis>
.
<emphasis id="B90B3A6633453F29941B250C9B6CBF1C" box="[1031,1087,1598,1619]" italics="true" pageId="22" pageNumber="23">acuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33453F299456250C9BCDBF1C" box="[1098,1182,1598,1619]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B90B3A6633453F299456250C9BCBBF1C" bold="true" box="[1098,1176,1598,1619]" pageId="22" pageNumber="23">sp. nov</emphasis>
.
</taxonomicNameLabel>
, left lateral filament on segment V, ventral. Scale bars =
<quantity id="4C874B9133453F2991FF256E9D71BF3D" box="[483,546,1628,1650]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="23" unit="mm" value="2.0">2 mm</quantity>
. Abbreviations:
<emphasis id="B90B3A6633453F2992D4256F9DB3BF3D" box="[712,736,1629,1650]" italics="true" pageId="22" pageNumber="23">ac</emphasis>
= anal cell; alw = anterolateral wart; amw = anteromesal wart; A1+2+3 = fused anal veins A1, A2, A3;
<emphasis id="B90B3A6633453F2991DB254F9E8CBFDC" box="[455,479,1661,1683]" italics="true" pageId="22" pageNumber="23">dc</emphasis>
= discoidal cell;
<emphasis id="B90B3A6633453F299291254C9DF9BFDC" box="[653,682,1662,1683]" italics="true" pageId="22" pageNumber="23">mc</emphasis>
= median cell; pw = posterior wart.
</paragraph>
</caption>
<paragraph id="8BC0E67433453F2990DB258F9E12BEC5" blockId="22.[151,1437,1725,2038]" pageId="22" pageNumber="23">
Male. Abdomen with paired reticulate-walled internal sacs in segments VI and VII (
<figureCitation id="1344FAF133453F29946C258F9A5FBF99" box="[1136,1292,1725,1750]" captionStart="FIGURES 179187" captionStartId="41.[151,264,1053,1076]" captionTargetBox="[151,1436,291,1027]" captionTargetId="figure@41.[151,1436,291,1032]" captionTargetPageId="41" captionText="FIGURES 179187. Arcyphysa gen. nov. spp., diagnostic features. 179182, ♀ terminal abdominal segments: 179180, Arc. angusta (NSW, Styx R. [PT-1005]): 179, ventral, and 180, left lateral; 181182, Arc. diamontona sp. nov. [PT-1006]: 181, ventral, and 182, left lateral. 183187, ♂ reticulate-walled internal sacs of abdominal segments VI and VII: 183, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]), ventral; 184185, Arc. acmea sp. nov. (Qld, Mt Bartle Frere,0.5 km Nof South Peak [PT-999]):184, rightlateral, and 185, ventral;186, Arc.anaplasis sp.nov. (Qld, 25 km along Mt Lewis Road, SW of Mossman [PT-1003]), ventral; 187, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]), ventral. Abbreviation:VIII = abdominal segment VIII." httpUri="https://zenodo.org/record/1241766/files/figure.png" pageId="22" pageNumber="23">Figs 183187</figureCitation>
) about equal length, rarely longer than length of respective segment. Genitalia (
<figureCitation id="1344FAF133453F29939425D39B76BFB5" box="[904,1061,1761,1786]" captionStart-0="FIGURES 102109" captionStart-1="FIGURES 110116" captionStartId-0="27.[151,264,1729,1752]" captionStartId-1="32.[151,264,1363,1386]" captionTargetBox-0="[150,1435,225,1691]" captionTargetBox-1="[155,1436,361,1352]" captionTargetId-0="figure@27.[150,1436,193,1708]" captionTargetId-1="figure@32.[151,1436,331,1352]" captionTargetPageId-0="27" captionTargetPageId-1="32" captionText-0="FIGURES 102109. Arcyphysa gen. nov. spp., diagnostic features.102103, Arc.diamontona sp. nov., Vic., Diamond Creek, 7 km SE of Gembrook [PT-1006], ♂: 102, right fore- and hind wings, dorsal, 103, maxillary palp. 104109, ♂ genitalia. 104 105, Arc. angusta (Banks 1939; NSW, Styx R. [PT-1005]): 104, ventral, and 105, left lateral. 106107, Arc. diamontona sp. nov. [PT-1006]:106, ventral, and 107, left lateral.108109, Arc.fraserensis sp. nov. (Qld, Fraser Island, Pile Valley, nr Central Station [PT-1125]):108, ventral,and 109, leftlateral.Scale bars: Fig.102 = 1 mm; Fig.103 = 0.5 mm.Abbreviations:ac= anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; fI, fII, fIII, fIV, fV = forks IV; mc= median cell; har = harpago; ph = phallic apparatus; IX, X = abdominal segments IX, X" captionText-1="FIGURES 110116. Arcyphysa gen. nov. spp., diagnostic features. 110111, Arc. hystricosa (Neboiss 1979) (Qld, North Stradbroke Island [PT-512]): 110, ventral; 111, left lateral. 112, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek), ventral. 113115, Arc.candela sp. nov. (NSW, Brown Mountain [PT-1017]): 113, left lateral; 114, ventral; 115, dorsal. 116, Arc. diamontona sp. nov. (Vic., Diamond Creek, 7 km SE of Gembrook [PT-1047]), ♀ terminal abdominal segments, left lateral." httpUri-0="https://zenodo.org/record/1241754/files/figure.png" httpUri-1="https://zenodo.org/record/1241756/files/figure.png" pageId="22" pageNumber="23">Figs 104115</figureCitation>
, 117131,
<figureCitation id="1344FAF133453F2994BE25D39A5ABFB5" box="[1186,1289,1761,1786]" captionStart-0="FIGURES 146163" captionStart-1="FIGURES 164178" captionStartId-0="39.[151,264,1675,1698]" captionStartId-1="40.[151,264,1776,1799]" captionTargetBox-0="[151,1436,209,1614]" captionTargetBox-1="[151,1436,194,1751]" captionTargetId-0="figure@39.[151,1436,209,1620]" captionTargetId-1="figure@40.[151,1436,193,1756]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionText-0="FIGURES 146163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154163, ♂ genitalia. 154155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X." captionText-1="FIGURES 164178. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 164165, Arc. lovedayi sp. nov. (holotype): 164, ventral; 165, left lateral. 166167, Arc. crescentina sp. nov. (Qld, Tully Falls [PT-1002]): 166, ventral; 167, left lateral. 168169, Arc. hugginsi sp. nov. (Qld, Bellenden Ker Range [PT-1000]): 168, ventral; 169, left lateral. 170, Arc. sybillae sp. nov. (Qld, Bloomfield River [PT-1016]), ventral. 171173, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge): 171, ventral; 172, left lateral;173, dorsal. 174175, Arc.anaplasis sp. nov. (Qld, 25 km along Mt LewisRoad, SW of Mossman [PT-1003]): 174, ventral; 175, left lateral.176, Arc.acmea sp. nov. (Qld, Mt Bartle Frere, 0.5 km N of South Peak [PT-999]), ventral.177 178, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]): 177, ventral; 178, left lateral." httpUri-0="https://zenodo.org/record/1241762/files/figure.png" httpUri-1="https://zenodo.org/record/1241764/files/figure.png" pageId="22" pageNumber="23">154178</figureCitation>
): Tergites IX and X fused, membranous, somewhat rounded apically with small median incision; sternite IX with upper lateral angle extended posteriorly into processes of various shapes; inferior appendages short, robust, harpago undivided or bifid; phallic apparatus in lateral view gradually curved downward, in ventral view with base abruptly widened, without spines.
</paragraph>
<paragraph id="8BC0E67433453F2990DB24A79D28BEBA" blockId="22.[151,1437,1725,2038]" pageId="22" pageNumber="23">
Female (
<figureCitation id="1344FAF133453F29913124A79EC5BEE1" box="[301,406,1941,1966]" captionStart="FIGURES 110116" captionStartId="32.[151,264,1363,1386]" captionTargetBox="[155,1436,361,1352]" captionTargetId="figure@32.[151,1436,331,1352]" captionTargetPageId="32" captionText="FIGURES 110116. Arcyphysa gen. nov. spp., diagnostic features. 110111, Arc. hystricosa (Neboiss 1979) (Qld, North Stradbroke Island [PT-512]): 110, ventral; 111, left lateral. 112, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek), ventral. 113115, Arc.candela sp. nov. (NSW, Brown Mountain [PT-1017]): 113, left lateral; 114, ventral; 115, dorsal. 116, Arc. diamontona sp. nov. (Vic., Diamond Creek, 7 km SE of Gembrook [PT-1047]), ♀ terminal abdominal segments, left lateral." httpUri="https://zenodo.org/record/1241756/files/figure.png" pageId="22" pageNumber="23">Figs 116</figureCitation>
,
<figureCitation id="1344FAF133453F2991BA24A49D5EBEE1" box="[422,525,1942,1966]" captionStart="FIGURES 179187" captionStartId="41.[151,264,1053,1076]" captionTargetBox="[151,1436,291,1027]" captionTargetId="figure@41.[151,1436,291,1032]" captionTargetPageId="41" captionText="FIGURES 179187. Arcyphysa gen. nov. spp., diagnostic features. 179182, ♀ terminal abdominal segments: 179180, Arc. angusta (NSW, Styx R. [PT-1005]): 179, ventral, and 180, left lateral; 181182, Arc. diamontona sp. nov. [PT-1006]: 181, ventral, and 182, left lateral. 183187, ♂ reticulate-walled internal sacs of abdominal segments VI and VII: 183, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]), ventral; 184185, Arc. acmea sp. nov. (Qld, Mt Bartle Frere,0.5 km Nof South Peak [PT-999]):184, rightlateral, and 185, ventral;186, Arc.anaplasis sp.nov. (Qld, 25 km along Mt Lewis Road, SW of Mossman [PT-1003]), ventral; 187, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]), ventral. Abbreviation:VIII = abdominal segment VIII." httpUri="https://zenodo.org/record/1241766/files/figure.png" pageId="22" pageNumber="23">179182</figureCitation>
). Abdominal sternite VIII well divided mesoventrally and produced posterad, laterodistal angle produced slightly, rounded; segment IX with large dorsoventrally elongate sclerotic cavity, inner surface of which with granular appearance.
</paragraph>
</subSubSection>
<subSubSection id="C365B5FF33443F2890DB23A59FA4B99A" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BC0E67433443F2890DB23A59FA4B99A" blockId="23.[151,1437,151,1149]" pageId="23" pageNumber="24">
<emphasis id="B90B3A6633443F2890DB23A59E33B9FF" bold="true" box="[199,352,151,176]" pageId="23" pageNumber="24">Distribution.</emphasis>
Most species are from north-eastern
<collectingCountry id="F368A6E433443F28931323A59C2AB9FF" box="[783,889,151,176]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
, but one species is found in central and eastern
<collectingRegion id="49BB289633443F28908B238E9FA7B99A" box="[151,244,188,213]" country="Australia" name="Victoria" pageId="23" pageNumber="24">Victoria</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C365B5FF33443F2890DB23ED9CD0B80F" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BC0E67433443F2890DB23ED9CD0B80F" blockId="23.[151,1437,151,1149]" pageId="23" pageNumber="24">
<emphasis id="B90B3A6633443F2890DB23ED9E1DB9B7" bold="true" box="[199,334,223,248]" pageId="23" pageNumber="24">Etymology.</emphasis>
<taxonomicName id="4C7F9DF733443F28914523D29E9DB9B7" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[345,462,224,248]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="42" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B3A6633443F28914523D29E9DB9B7" box="[345,462,224,248]" italics="true" pageId="23" pageNumber="24">Arcyphysa</emphasis>
</taxonomicName>
, the name derived from Greek,
<emphasis id="B90B3A6633443F28934D23D29CDDB9B7" box="[849,910,224,248]" italics="true" pageId="23" pageNumber="24">arkys</emphasis>
being descriptive of bubble- or bellows-like appearance of lateral lobes in ventral view of male genitalia, and
<emphasis id="B90B3A6633443F28938C22369C82B853" box="[912,977,260,284]" italics="true" pageId="23" pageNumber="24">physa</emphasis>
, net or mesh, for the appearance of the internal reticulate-walled sacs of the male. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C365B5FF33443F2890DB227E9D4FBD32" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BC0E67433443F2890DB227E9FA6BB2F" blockId="23.[151,1437,151,1149]" pageId="23" pageNumber="24">
<emphasis id="B90B3A6633443F2890DB227E9E68B82A" bold="true" box="[199,315,332,357]" pageId="23" pageNumber="24">Remarks.</emphasis>
Relationships of the pair of male genitalic processes above the phallic apparatus and gonopods are often unclear. They are referred to by Mosely (in
<bibRefCitation id="EFEE9B8533443F2892FD225D9B5FB8C7" box="[737,1036,367,392]" pageId="23" pageNumber="24" refId="ref35807" refString="Mosely, M. E. &amp; Kimmins, D. E. (1953) The Trichoptera (Caddis-flies) of Australia and New Zealand. British Museum (Natural History), London, 550 pp. https: // doi. org / 10.5962 / bhl. title. 118696" type="book">Mosely &amp; Kimmins 1953</bibRefCitation>
) in the description of
<taxonomicName id="4C7F9DF733443F28950922429FB1B8E3" authorityName="Mosely, in Mosely &amp; Kimmins" authorityYear="1953" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="satana">
<emphasis id="B90B3A6633443F28950922429FB1B8E3" italics="true" pageId="23" pageNumber="24">Diplectrona satana</emphasis>
</taxonomicName>
as an 'upper penis-cover forming a pair of processes' and by
<bibRefCitation id="EFEE9B8533443F28938F22A69B11B8E3" box="[915,1090,404,429]" pageId="23" pageNumber="24" refId="ref35916" refString="Neboiss, A. (1979) A review of caddis flies from three coastal islands of south-eastern Queensland (Insecta: Trichoptera). Australian Journal of Marine and Freshwater Research, 29 (6) [&quot; December 1978 &quot;], 825 - 843. [Commonwealth Scientific and Industrial Research Organization Library stamp &quot; 25 JAN 1979 &quot;]" type="journal article">Neboiss (1979)</bibRefCitation>
, in
<taxonomicName id="4C7F9DF733443F28947022A79A52B8E3" authorityName="Neboiss" authorityYear="1979" box="[1132,1281,404,428]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="hystricosa">
<emphasis id="B90B3A6633443F28947022A79A52B8E3" box="[1132,1281,404,428]" italics="true" pageId="23" pageNumber="24">D. hystricosa</emphasis>
</taxonomicName>
, as a 'pointed process' arising from each lower margin of tergite X. In some species the one or two processes appear clearly to be lobes on the lateral margins of tergite X, in others to be individual structures arising from the apicolateral margins of segment IX. Tergite X is sometimes more or less fused with tergite IX. These variously expressed structures may not be homologues, but to avoid confusion, they are simply referred to as 'apicolateral processes' in the descriptions and key.
</paragraph>
<paragraph id="8BC0E67433443F2890DB215E9D4FBD32" blockId="23.[151,1437,151,1149]" pageId="23" pageNumber="24">
Neboiss in his draft notes on Australian diplectronines, recognised three separate sets among species that here are assigned to this new genus,
<taxonomicName id="4C7F9DF733443F28921721A29DD3BBE7" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[523,640,656,680]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="42" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B3A6633443F28921721A29DD3BBE7" box="[523,640,656,680]" italics="true" pageId="23" pageNumber="24">Arcyphysa</emphasis>
</taxonomicName>
. One set, here termed the
<emphasis id="B90B3A6633443F2893A421A29C8CBBE8" box="[952,991,656,679]" italics="true" pageId="23" pageNumber="24">Arc</emphasis>
.
<taxonomicName id="4C7F9DF733443F2893EC21A39B1ABBE7" authorityName="Banks" authorityYear="1939" box="[1008,1097,657,680]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="angusta">
<emphasis id="B90B3A6633443F2893EC21A39B1ABBE7" box="[1008,1097,657,680]" italics="true" pageId="23" pageNumber="24">angusta</emphasis>
</taxonomicName>
Group, included
<taxonomicName id="4C7F9DF733443F28950921A29FA3BB83" authorityName="Banks" authorityYear="1939" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="angusta">
<emphasis id="B90B3A6633443F28950921A29FA3BB83" italics="true" pageId="23" pageNumber="24">Diplectrona angusta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F9DF733443F2890E321869D13BB84" authority="Arc." authorityName="Arc." authorityYear="2018" box="[255,576,692,716]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="hystricosa" status="sp. nov.">
<emphasis id="B90B3A6633443F2890E321869D52BB83" box="[255,513,692,716]" italics="true" pageId="23" pageNumber="24">Diplectrona hystricosa</emphasis>
,
<emphasis id="B90B3A6633443F28920D21869D13BB84" box="[529,576,692,715]" italics="true" pageId="23" pageNumber="24">Arc.</emphasis>
</taxonomicName>
<taxonomicName id="4C7F9DF733443F28925421869D91BB83" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[584,706,692,716]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fraserensis" status="sp. nov.">
<emphasis id="B90B3A6633443F28925421869D91BB83" box="[584,706,692,716]" italics="true" pageId="23" pageNumber="24">fraserensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238871D33443F2892D721869C76BB83" box="[715,805,692,717]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F2892D721869C4CBB82" bold="true" box="[715,799,692,717]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
, and
<emphasis id="B90B3A6633443F28937B21869B77BB83" box="[871,1060,692,716]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28938221869B77BB83" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[926,1060,692,716]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="diamontona" status="sp. nov.">diamontona</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28943121869BDBBB83" box="[1069,1160,692,717]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28943121869BD2BB82" bold="true" box="[1069,1153,692,717]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
Five further species are added now to this group:
<emphasis id="B90B3A6633443F2891A821EA9D11BBBF" box="[436,578,728,752]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2891F521EA9D11BBBF" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[489,578,728,752]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="candela" status="sp. nov.">candela</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28925521E59DF2BBA0" box="[585,673,727,752]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28925521E59DC8BBBF" bold="true" box="[585,667,727,752]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F2892AC21EA9C17BBBF" box="[688,836,728,752]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2892F921EA9C17BBBF" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[741,836,728,752]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="lovedayi" status="sp. nov.">lovedayi</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28935721E59CF0BBA0" box="[843,931,727,752]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28935721E59CCDBBBF" bold="true" box="[843,926,727,752]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F2893AE21EA9B04BBA0" box="[946,1111,728,751]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2893FB21EA9B04BBA0" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[999,1111,728,751]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="naumanni" status="sp. nov.">naumanni</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28944221E59BE5BBA0" box="[1118,1206,727,752]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28944221E59BE2BBBF" bold="true" box="[1118,1201,727,752]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F2894D921EA9A63BBA0" box="[1221,1328,728,751]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2894E721EA9A63BBA0" authorityName="Kimmins, Kimmins (in Mosely &amp; Kimmins" authorityYear="1953" box="[1275,1328,728,751]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="rossi">rossi</taxonomicName>
</emphasis>
, and
<emphasis id="B90B3A6633443F28957121EA9FB1BA5B" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28908B21CF9FB1BA5B" authorityName="Mosely, in Mosely &amp; Kimmins" authorityYear="1953" box="[151,226,765,788]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="satana">satana</taxonomicName>
</emphasis>
. Neboiss in his preliminary work did not assign
<emphasis id="B90B3A6633443F28931B21CE9C21BA5B" box="[775,882,764,788]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28932121CF9C21BA5B" authorityName="Kimmins, Kimmins (in Mosely &amp; Kimmins" authorityYear="1953" box="[829,882,765,788]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="rossi">rossi</taxonomicName>
</emphasis>
or
<emphasis id="B90B3A6633443F28938521CE9B49BA5B" box="[921,1050,764,788]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2893D321CF9B49BA5B" authorityName="Mosely, in Mosely &amp; Kimmins" authorityYear="1953" box="[975,1050,765,788]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="satana">satana</taxonomicName>
</emphasis>
to any of his groups; however, on the basis of head and general genitalic features they cluster with members of the
<emphasis id="B90B3A6633443F28943120129BEEBA77" box="[1069,1213,800,824]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28947820139BEEBA77" authorityName="Banks" authorityYear="1939" box="[1124,1213,801,824]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
</emphasis>
Group. The second set, here the
<emphasis id="B90B3A6633443F28913920769E1FBA14" box="[293,332,836,859]" italics="true" pageId="23" pageNumber="24">Arc</emphasis>
.
<taxonomicName id="4C7F9DF733443F28914620769EE9BA13" authorityName="Wells &amp; Contents" authorityYear="2018" box="[346,442,836,860]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="hugginsi">
<emphasis id="B90B3A6633443F28914620769EE9BA13" box="[346,442,836,860]" italics="true" pageId="23" pageNumber="24">hugginsi</emphasis>
</taxonomicName>
Group, included
<emphasis id="B90B3A6633443F28926320769C47BA13" box="[639,788,836,860]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2892A820769C47BA13" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[692,788,836,860]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="hugginsi" status="sp. nov.">hugginsi</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28930020769C27BA13" box="[796,884,836,861]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28930020769C3CBA12" bold="true" box="[796,879,836,861]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F28939820769B43BA13" box="[900,1040,836,860]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2893A520769B43BA13" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[953,1040,836,860]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="sybillae" status="sp. nov.">sybillae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28940B20769B23BA13" box="[1047,1136,836,861]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28940B20769B39BA12" bold="true" box="[1047,1130,836,861]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F28946320769A67BA13" box="[1151,1332,836,860]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2894A820779A67BA13" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1204,1332,837,860]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="crescentina" status="sp. nov.">crescentina</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28952020769AC7BA13" box="[1340,1428,836,861]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28952020769ADDBA12" bold="true" box="[1340,1422,836,861]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
, and
<emphasis id="B90B3A6633443F2890DB205A9E00BACF" box="[199,339,872,896]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F2890E2205A9E00BACF" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[254,339,872,896]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="caldera" status="sp. nov.">caldera</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28914020559EE5BA30" box="[348,438,871,896]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28914020559EFCBACF" bold="true" box="[348,431,871,896]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
; a fifth species is now added,
<emphasis id="B90B3A6633443F28930D205A9C98BACF" box="[785,971,872,896]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F289354205B9C98BACF" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[840,971,873,896]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="carnarvona" status="sp. nov.">carnarvona</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F2893CF20559B7EBA30" box="[979,1069,871,896]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F2893CF20559B75BACF" bold="true" box="[979,1062,871,896]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
The third group, here called the
<emphasis id="B90B3A6633443F28908B20BE9FEDBAEC" box="[151,190,908,931]" italics="true" pageId="23" pageNumber="24">Arc</emphasis>
.
<taxonomicName id="4C7F9DF733443F2890D320BF9E4BBAEB" authorityName="Wells &amp; Contents" authorityYear="2018" box="[207,280,909,932]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="acmea">
<emphasis id="B90B3A6633443F2890D320BF9E4BBAEB" box="[207,280,909,932]" italics="true" pageId="23" pageNumber="24">acmea</emphasis>
</taxonomicName>
Group, comprised
<emphasis id="B90B3A6633443F2891E420BE9D2ABAEB" box="[504,633,908,932]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28922C20BF9D2ABAEB" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[560,633,909,932]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="acmea" status="sp. nov.">acmea</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28929F20BE9D8DBAEB" box="[643,734,908,933]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28929F20BE9D8BBAEA" bold="true" box="[643,728,908,933]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F2892F320BE9CC1BAEB" box="[751,914,908,932]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28933B20BE9CC1BAEB" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[807,914,908,932]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="anaplasis" status="sp. nov.">anaplasis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28938020BE9CABBAEB" box="[924,1016,908,933]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28938020BE9CA1BAEA" bold="true" box="[924,1010,908,933]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F28941520BE9BDEBAEB" box="[1033,1165,908,932]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28945D20BE9BDEBAEB" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1089,1165,908,932]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tortula" status="sp. nov.">tortula</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28948B20BE9BA1BAEB" box="[1175,1266,908,933]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28948B20BE9BBFBAEA" bold="true" box="[1175,1260,908,933]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F28951820BE9A22BAEB" box="[1284,1393,908,932]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28952020BE9A22BAEB" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1340,1393,908,932]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="flinti" status="sp. nov.">flinti</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F28956620BE9F95BA88" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F28956620BE9F92BA87" bold="true" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="B90B3A6633443F2890C620829E39BA87" box="[218,362,944,968]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28910F20829E39BA87" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[275,362,944,968]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="volsella" status="sp. nov.">volsella</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F289169209D9E81BA88" box="[373,466,943,968]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F289169209D9E9FBA87" bold="true" box="[373,460,943,968]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
, and
<emphasis id="B90B3A6633443F28920420829DDABA87" box="[536,649,944,968]" italics="true" pageId="23" pageNumber="24">
Arc.
<taxonomicName id="4C7F9DF733443F28924E20829DDABA87" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[594,649,944,968]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nebo" status="sp. nov.">nebo</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A238871D33443F289288209D9DA2BA88" box="[660,753,943,968]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B90B3A6633443F289288209D9DB9BA87" bold="true" box="[660,746,943,968]" pageId="23" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
Neboiss proposed, tentatively, that these be recognised as separate new genera. However, features of head, antennae, maxillary palpi, and wings unite all species here assigned to
<taxonomicName id="4C7F9DF733443F28913D20CA9EC5BD5F" authority="Wells &amp; Contents, 2018" authorityName="Wells &amp; Contents" authorityYear="2018" box="[289,406,1016,1040]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="42" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B3A6633443F28913D20CA9EC5BD5F" box="[289,406,1016,1040]" italics="true" pageId="23" pageNumber="24">Arcyphysa</emphasis>
</taxonomicName>
. They differ in aspects of male genitalia, but, except for the
<emphasis id="B90B3A6633443F28947C20CA9BD4BD40" box="[1120,1159,1016,1039]" italics="true" pageId="23" pageNumber="24">Arc</emphasis>
.
<taxonomicName id="4C7F9DF733443F28948520CA9BAABD5F" authorityName="Wells &amp; Contents" authorityYear="2018" box="[1177,1273,1016,1040]" class="Insecta" family="Hydropsychidae" genus="Arcyphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="hugginsi">
<emphasis id="B90B3A6633443F28948520CA9BAABD5F" box="[1177,1273,1016,1040]" italics="true" pageId="23" pageNumber="24">hugginsi</emphasis>
</taxonomicName>
Group (all of which have a pair of parameres basally on the phallic apparatus), no clear synapomorphies can be recognised to unite members. For the present, a single new genus is recognised. The order of treatments here is alphabetic within each of the 3 loose species groups.
</paragraph>
</subSubSection>
</treatment>
</document>