819 lines
100 KiB
XML
819 lines
100 KiB
XML
<document id="1B5C606C3B746746B756D7FE54C97ECE" ID-DOI="10.1080/00222933.2020.1736679" ID-ISSN="1464-5262" ID-Zenodo-Dep="4608898" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615923489356" checkinUser="carolina" docAuthor="Mahony, Stephen, Kamei, Rachunliu G., Teeling, Emma C. & Biju, S. D." docDate="2020" docId="03C78783FF816A5CFE78CCC8FD4DFBE5" docLanguage="en" docName="JNATHIST.54.1-4.119-194.pdf" docOrigin="Journal of Natural History 54 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2020.1736679" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Megophrys (Xenophrys) awuh Mahony & Kamei & Teeling & Biju 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="148" masterDocId="FFFEFFFBFF996A42FFDECE02FF83FFAB" masterDocTitle="Taxonomic review of the Asian Horned Frogs (Amphibia: Megophrys Kuhl & Van Hasselt) of Northeast India and Bangladesh previously misidentified as M. parva (Boulenger), with descriptions of three new species" masterLastPageNumber="194" masterPageNumber="119" pageNumber="142" updateTime="1698887099588" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DB995ED45AFCE7932D1AB12DEB658BD2">Taxonomic review of the Asian Horned Frogs (Amphibia: Megophrys Kuhl & Van Hasselt) of Northeast India and Bangladesh previously misidentified as M. parva (Boulenger), with descriptions of three new species</mods:title>
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<mods:name id="F83AEF1EFF0BD7006C101831845F488F" type="personal">
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<mods:roleTerm id="36529F97A9580406D02F92FF5F68A6C4">Author</mods:roleTerm>
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<mods:namePart id="4C612B9184A04D59FC78AFA468899A2C">Mahony, Stephen</mods:namePart>
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<mods:roleTerm id="131F33CF71A18D8AA38E7E529ACE0608">Author</mods:roleTerm>
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<mods:namePart id="2493FC83547738301743A866F78F81E3">Kamei, Rachunliu G.</mods:namePart>
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<mods:roleTerm id="446D1505E7A0643AC7993B4CC72E9049">Author</mods:roleTerm>
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<mods:namePart id="2F183E4BB45196A69B1DC8A68618101E">Teeling, Emma C.</mods:namePart>
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</mods:name>
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<mods:name id="766F90AE5808C93600EA691BFD87B5D1" type="personal">
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<mods:roleTerm id="AA19EDF38E7600E21C40389F2D2D19BE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D7ABB4E01867046A0FF319E7003A104C">Biju, S. D.</mods:namePart>
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<mods:typeOfResource id="7EB017B34827351810FF261C7FD44F25">text</mods:typeOfResource>
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<mods:title id="A160F05181F941E37D2440DFA2BD6AD5">Journal of Natural History</mods:title>
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<mods:part id="28795B01215A559CEA73E28D36450729">
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<mods:date id="F84A7A034383CAD61888AEB58F97B9F0">2020</mods:date>
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<mods:number id="AA2C4A87D6E2DCBA5BB15557F25584CB">2020-04-28</mods:number>
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<mods:detail id="EC7F7FDC3BBD0C85295B73D6B1EF985D" type="volume">
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<mods:number id="49CF5D61F4FB2CAA529E89DE268CA582">54</mods:number>
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<mods:number id="BB72EE070E378975E560530FD7DB609D">1 - 4</mods:number>
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<mods:start id="E6031198428CEF1800AF30403DD9EBA8">119</mods:start>
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<mods:url id="55C45ED3A2921ED79DC1E22357AB7BB5">http://dx.doi.org/10.1080/00222933.2020.1736679</mods:url>
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<mods:classification id="81DE6574E1AF1F0BA48F20E4F96C8159">journal article</mods:classification>
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<mods:identifier id="27B614648490A4D89CBF87C99184F3B2" type="DOI">10.1080/00222933.2020.1736679</mods:identifier>
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<mods:identifier id="B457359035299D490B26415BA92C858B" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03C78783FF816A5CFE78CCC8FD4DFBE5" ID-DOI="http://doi.org/10.5281/zenodo.4608876" ID-Zenodo-Dep="4608876" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5" httpUri="http://treatment.plazi.org/id/03C78783FF816A5CFE78CCC8FD4DFBE5" lastPageId="30" lastPageNumber="148" pageId="24" pageNumber="142">
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<subSubSection id="C374651EFF816A5AFE78CCC8FCFEFD48" box="[422,893,714,739]" pageId="24" pageNumber="142" type="nomenclature">
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<paragraph id="8BD13695FF816A5AFE78CCC8FCFEFD48" blockId="24.[422,893,714,739]" box="[422,893,714,739]" pageId="24" pageNumber="142">
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<heading id="D09981F9FF816A5AFE78CCC8FCFEFD48" box="[422,893,714,739]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="142" reason="2">
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<emphasis id="B91AEA87FF816A5AFE78CCC8FCFEFD48" bold="true" box="[422,893,714,739]" pageId="24" pageNumber="142">
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<taxonomicName id="4C6E4D16FF816A5AFE78CCC8FC91FD48" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[422,786,714,739]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="24" pageNumber="142" phylum="Chordata" rank="species" species="awuh" status="sp. nov." subGenus="Xenophrys">
|
||
<emphasis id="B91AEA87FF816A5AFE78CCC8FDADFD48" bold="true" box="[422,558,714,739]" italics="true" pageId="24" pageNumber="142">Megophrys</emphasis>
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||
(
|
||
<emphasis id="B91AEA87FF816A5AFDE0CCC8FD3CFD48" bold="true" box="[574,703,714,739]" italics="true" pageId="24" pageNumber="142">Xenophrys</emphasis>
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||
)
|
||
<emphasis id="B91AEA87FF816A5AFD11CCC8FC91FD48" bold="true" box="[719,786,714,739]" italics="true" pageId="24" pageNumber="142">awuh</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A22957FCFF816A5AFCC5CCC8FCFEFD48" box="[795,893,714,739]" pageId="24" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
|
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C374651EFF816A5EFF7ECD22FC44FF47" lastPageId="28" lastPageNumber="146" pageId="24" pageNumber="142" type="materials_examined">
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<paragraph id="8BD13695FF816A5AFF7ECD22FCE7FCD4" blockId="24.[160,1155,800,895]" pageId="24" pageNumber="142">
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||
<materialsCitation id="3B063CC8FF816A5AFF7ECD22FCE7FCD4" collectingDate="2007-07-05" collectionCode="RGK, VK" country="India" county="Northeast" elevation="2220" latitude="25.464443" location="Adult" longLatPrecision="1" longitude="94.70389" municipality="Phek district" pageId="24" pageNumber="142" specimenCode="BNHS 6070, SDBDU 2007.163" specimenCount="1" stateProvince="Nagaland" typeStatus="holotype">
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<emphasis id="B91AEA87FF816A5AFF7ECD22FE91FC91" bold="true" box="[160,274,800,826]" italics="true" pageId="24" pageNumber="142">
|
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<typeStatus id="54D58837FF816A5AFF7ECD22FE8EFC91" box="[160,269,800,826]" pageId="24" pageNumber="142" type="holotype">Holotype</typeStatus>
|
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.
|
||
</emphasis>
|
||
<location id="8EB1604EFF816A5AFEFFCD20FEDDFC91" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFEFFCD20FEDDFC91" box="[289,350,802,826]" country="India" county="Northeast" latitude="25.464443" longLatPrecision="1" longitude="94.70389" municipality="Phek district" name="Adult" pageId="24" pageNumber="142" stateProvince="Nagaland">Adult</location>
|
||
male (
|
||
<specimenCode id="DBC89EEEFF816A5AFE77CD20FDABFC91" box="[425,552,802,826]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="24" pageNumber="142">BNHS 6070</specimenCode>
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[field tag
|
||
<specimenCode id="DBC89EEEFF816A5AFD45CD20FCD9FC91" box="[667,858,802,826]" collectionCode="SDBDU" pageId="24" pageNumber="142">SDBDU 2007.163</specimenCode>
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] [
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<figureCitation id="13552A10FF816A5AFCB0CD20FC49FC91" box="[878,970,802,826]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="24" pageNumber="142">Figure 5</figureCitation>
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]) from
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<location id="8EB1604EFF816A5AFBC3CD20FB00FC91" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFBC3CD20FB00FC91" box="[1053,1155,802,826]" country="India" county="Northeast" latitude="25.464443" longLatPrecision="1" longitude="94.70389" municipality="Phek district" name="Star Lake" pageId="24" pageNumber="142" stateProvince="Nagaland">Star Lake</location>
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(
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<geoCoordinate id="EE5A5052FF816A5AFF78CD46FE9DFCF7" box="[166,286,836,860]" degrees="25.464444" direction="north" orientation="latitude" pageId="24" pageNumber="142" precision="1" value="25.464443">25.464444</geoCoordinate>
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,
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<geoCoordinate id="EE5A5052FF816A5AFEF6CD46FE1DFCF7" box="[296,414,836,860]" degrees="94.703889" direction="east" orientation="longitude" pageId="24" pageNumber="142" precision="1" value="94.70389">94.703889</geoCoordinate>
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,
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<elevation id="0043D1A6FF816A5AFE77CD46FDA9FCF7" box="[425,554,836,860]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="24" pageNumber="142" unit="m" value="2220.0">
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<quantity id="4C969B70FF816A5AFE77CD46FE7DFCF7" box="[425,510,836,860]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="24" pageNumber="142" unit="m" value="2220.0">2220 m</quantity>
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asl
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</elevation>
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.),
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<location id="8EB1604EFF816A5AFD9ECD46FD58FCF7" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFD9ECD46FD58FCF7" box="[576,731,836,860]" country="India" county="Northeast" latitude="25.464443" longLatPrecision="1" longitude="94.70389" municipality="Phek district" name="Reguri village" pageId="24" pageNumber="142" stateProvince="Nagaland">Reguri village</location>
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,
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<location id="8EB1604EFF816A5AFD39CD46FCADFCF7" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFD39CD46FCADFCF7" box="[743,814,836,860]" country="India" county="Northeast" latitude="25.464443" longLatPrecision="1" longitude="94.70389" municipality="Phek district" name="Meluri" pageId="24" pageNumber="142" stateProvince="Nagaland">Meluri</location>
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circle,
|
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<collectingMunicipality id="6BB5ACEFFF816A5AFCA1CD46FB8EFCF7" box="[895,1037,836,860]" pageId="24" pageNumber="142">Phek district</collectingMunicipality>
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,
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<collectingRegion id="49AAF877FF816A5AFBC6CD46FF5AFCD4" country="India" name="Nagaland" pageId="24" pageNumber="142">Nagaland state</collectingRegion>
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,
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<collectingCounty id="62B04E19FF816A5AFF3DCD65FED2FCD4" box="[227,337,871,895]" pageId="24" pageNumber="142">Northeast</collectingCounty>
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<collectingCountry id="F3797605FF816A5AFE87CD65FE10FCD4" box="[345,403,871,895]" name="India" pageId="24" pageNumber="142">India</collectingCountry>
|
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, collected by
|
||
<collectionCode id="ED7FAE50FF816A5AFDF0CD65FDDFFCD4" box="[558,604,871,895]" pageId="24" pageNumber="142">RGK</collectionCode>
|
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and
|
||
<collectionCode id="ED7FAE50FF816A5AFD4BCD65FD31FCD4" box="[661,690,871,895]" pageId="24" pageNumber="142">VK</collectionCode>
|
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on
|
||
<date id="FFD01055FF816A5AFD01CD65FCE3FCD4" box="[735,864,871,895]" pageId="24" pageNumber="142" value="2007-07-05">
|
||
<collectingDate id="EF94E9BDFF816A5AFD01CD65FCE3FCD4" box="[735,864,871,895]" pageId="24" pageNumber="142" value="2007-07-05">5 July 2007</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
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</paragraph>
|
||
<paragraph id="8BD13695FF816A5AFF7ECDA9FCF5FC4C" blockId="24.[160,1155,939,999]" pageId="24" pageNumber="142">
|
||
<materialsCitation id="3B063CC8FF816A5AFF7ECDA9FCF1FC4C" collectionCode="BNHS, SDBDU" pageId="24" pageNumber="142" specimenCode="BNHS 6069, SDBDU 2007.161, BNHS 6071, SDBDU 2007.162" specimenCount="2" typeStatus="paratype">
|
||
<emphasis id="B91AEA87FF816A5AFF7ECDA9FE9FFC6E" bold="true" box="[160,284,939,965]" italics="true" pageId="24" pageNumber="142">
|
||
<typeStatus id="54D58837FF816A5AFF7ECDA9FE94FC6E" box="[160,279,939,965]" pageId="24" pageNumber="142" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult female (
|
||
<specimenCode id="DBC89EEEFF816A5AFE17CDAFFDCBFC6E" box="[457,584,941,965]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="24" pageNumber="142">BNHS 6069</specimenCode>
|
||
[field tag
|
||
<specimenCode id="DBC89EEEFF816A5AFD67CDAFFCF9FC6E" box="[697,890,941,965]" collectionCode="SDBDU" pageId="24" pageNumber="142">SDBDU 2007.161</specimenCode>
|
||
]) and adult male (
|
||
<specimenCode id="DBC89EEEFF816A5AFB9CCDAFFF54FC4C" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="24" pageNumber="142">BNHS 6071</specimenCode>
|
||
[field tag
|
||
<specimenCode id="DBC89EEEFF816A5AFE94CDCDFD8FFC4C" box="[330,524,975,999]" collectionCode="SDBDU" pageId="24" pageNumber="142">SDBDU 2007.162</specimenCode>
|
||
]) collected along with
|
||
<typeStatus id="54D58837FF816A5AFCD5CDCDFCF1FC4C" box="[779,882,975,999]" pageId="24" pageNumber="142" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF816A5AFF7ECA11FE77FACE" blockId="24.[160,1156,1043,1381]" pageId="24" pageNumber="142">
|
||
<emphasis id="B91AEA87FF816A5AFF7ECA11FE0FFB86" bold="true" box="[160,396,1043,1069]" italics="true" pageId="24" pageNumber="142">Referred specimens.</emphasis>
|
||
<materialsCitation id="3B063CC8FF816A5AFE42CA17FE73FACE" collectingDate="2007-06-18" collectingDateMax="2007-07-10" collectingDateMin="2007-06-18" collectionCode="NEI, RGK, VK, SDB" county="Jakhama village" elevation="1650" latitude="25.64139" location="river Dzutsuru" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" pageId="24" pageNumber="142" specimenCode="SDBDU 2007.111, SDBDU 2007.079, SDBDU 2007.080, SDBDU 2007.192" specimenCount="1" stateProvince="Nagaland">
|
||
Adult male (
|
||
<specimenCode id="DBC89EEEFF816A5AFDF8CA17FD66FB86" box="[550,741,1045,1069]" collectionCode="SDBDU" pageId="24" pageNumber="142">SDBDU 2007.111</specimenCode>
|
||
[
|
||
<figureCitation id="13552A10FF816A5AFD2BCA17FCD1FB86" box="[757,850,1045,1069]" captionStart="Figure 6" captionStartId="26.[160,225,1399,1422]" captionTargetBox="[164,1152,155,1354]" captionTargetId="figure-52@26.[163,1153,154,1368]" captionTargetPageId="26" captionText="Figure 6. Megophrys awuh sp. nov. referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b–f). specimens in preservation from Dzüleke village, showing variation: (b–d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e–f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view." figureDoi="http://doi.org/10.5281/zenodo.4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="24" pageNumber="142">Figure 6</figureCitation>
|
||
(a)]) from about
|
||
<quantity id="4C969B70FF816A5AFBD3CA17FBC5FB86" box="[1037,1094,1045,1069]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="142" unit="km" value="1.0">1 km</quantity>
|
||
from Dzükou tourist entry office (
|
||
<geoCoordinate id="EE5A5052FF816A5AFE3ECA35FDDBFBE4" box="[480,600,1079,1103]" degrees="25.571667" direction="north" orientation="latitude" pageId="24" pageNumber="142" precision="1" value="25.571667">25.571667</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A5052FF816A5AFDBACA35FD59FBE4" box="[612,730,1079,1103]" degrees="94.120278" direction="east" orientation="longitude" pageId="24" pageNumber="142" precision="1" value="94.12028">94.120278</geoCoordinate>
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||
, ca.
|
||
<elevation id="0043D1A6FF816A5AFCD3CA35FC0CFBE4" box="[781,911,1079,1103]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="24" pageNumber="142" unit="m" value="1650.0">
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<quantity id="4C969B70FF816A5AFCD3CA35FCE0FBE4" box="[781,867,1079,1103]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="24" pageNumber="142" unit="m" value="1650.0">1650 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.), off highway NH 39,
|
||
<collectingCounty id="62B04E19FF816A5AFF7ECA58FED8FBD9" box="[160,347,1114,1138]" pageId="24" pageNumber="142">Jakhama village</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BB5ACEFFF816A5AFEB2CA58FD9DFBD9" box="[364,542,1114,1138]" pageId="24" pageNumber="142">Kohima district</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49AAF877FF816A5AFDF0CA58FD63FBD9" box="[558,736,1114,1138]" country="India" name="Nagaland" pageId="24" pageNumber="142">Nagaland state</collectingRegion>
|
||
,
|
||
<collectionCode id="ED7FAE50FF816A5AFD2FCA58FC9AFBD9" box="[753,793,1114,1138]" pageId="24" pageNumber="142">NEI</collectionCode>
|
||
, collected by
|
||
<collectionCode id="ED7FAE50FF816A5AFC18CA58FC77FBD9" box="[966,1012,1114,1138]" pageId="24" pageNumber="142">RGK</collectionCode>
|
||
and
|
||
<collectionCode id="ED7FAE50FF816A5AFBE5CA58FBDBFBD9" box="[1083,1112,1114,1138]" pageId="24" pageNumber="142">VK</collectionCode>
|
||
on
|
||
<date id="FFD01055FF816A5AFF7ECA7FFEBCFB3E" box="[160,319,1149,1173]" pageId="24" pageNumber="142" value="2007-06-24">
|
||
<collectingDate id="EF94E9BDFF816A5AFF7ECA7FFEBCFB3E" box="[160,319,1149,1173]" pageId="24" pageNumber="142" value="2007-06-24">24 June 2007</collectingDate>
|
||
</date>
|
||
; adult male (
|
||
<specimenCode id="DBC89EEEFF816A5AFE07CA7FFD1FFB3E" box="[473,668,1149,1173]" collectionCode="SDBDU" pageId="24" pageNumber="142">SDBDU 2007.079</specimenCode>
|
||
[
|
||
<figureCitation id="13552A10FF816A5AFD71CA7FFC8DFB3E" box="[687,782,1149,1173]" captionStart="Figure 6" captionStartId="26.[160,225,1399,1422]" captionTargetBox="[164,1152,155,1354]" captionTargetId="figure-52@26.[163,1153,154,1368]" captionTargetPageId="26" captionText="Figure 6. Megophrys awuh sp. nov. referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b–f). specimens in preservation from Dzüleke village, showing variation: (b–d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e–f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view." figureDoi="http://doi.org/10.5281/zenodo.4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="24" pageNumber="142">Figure 6</figureCitation>
|
||
(e
|
||
<emphasis id="B91AEA87FF816A5AFCFDCA7EFCB3FB3F" box="[803,816,1148,1172]" italics="true" pageId="24" pageNumber="142">–</emphasis>
|
||
f)]) and adult female (
|
||
<specimenCode id="DBC89EEEFF816A5AFBEFCA7FFE86FB1C" collectionCode="SDBDU" pageId="24" pageNumber="142">SDBDU 2007.080</specimenCode>
|
||
[
|
||
<figureCitation id="13552A10FF816A5AFEC9CA9DFEF4FB1C" box="[279,375,1183,1207]" captionStart="Figure 6" captionStartId="26.[160,225,1399,1422]" captionTargetBox="[164,1152,155,1354]" captionTargetId="figure-52@26.[163,1153,154,1368]" captionTargetPageId="26" captionText="Figure 6. Megophrys awuh sp. nov. referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b–f). specimens in preservation from Dzüleke village, showing variation: (b–d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e–f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view." figureDoi="http://doi.org/10.5281/zenodo.4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="24" pageNumber="142">Figure 6</figureCitation>
|
||
(b
|
||
<emphasis id="B91AEA87FF816A5AFE53CA9DFE19FB1C" box="[397,410,1183,1207]" italics="true" pageId="24" pageNumber="142">–</emphasis>
|
||
d)]) from the
|
||
<location id="8EB1604EFF816A5AFDE7CA9DFD59FB1C" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFDE7CA9DFD59FB1C" box="[569,730,1183,1207]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="river Dzutsuru" pageId="24" pageNumber="142" stateProvince="Nagaland">river Dzütsürü</location>
|
||
(
|
||
<geoCoordinate id="EE5A5052FF816A5AFD34CA9DFCE2FB1C" box="[746,865,1183,1207]" degrees="25.613611" direction="north" orientation="latitude" pageId="24" pageNumber="142" precision="1" value="25.613611">25.613611</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A5052FF816A5AFCB1CA9DFC66FB1C" box="[879,997,1183,1207]" degrees="93.969444" direction="east" orientation="longitude" pageId="24" pageNumber="142" precision="1" value="93.969444">93.969444</geoCoordinate>
|
||
,
|
||
<elevation id="0043D1A6FF816A5AFC2FCA9DFBF5FB1C" box="[1009,1142,1183,1207]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="24" pageNumber="142" unit="m" value="2010.0">
|
||
<quantity id="4C969B70FF816A5AFC2FCA9DFBCBFB1C" box="[1009,1096,1183,1207]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="24" pageNumber="142" unit="m" value="2010.0">2010 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.),
|
||
<location id="8EB1604EFF816A5AFF7ECAC0FEC9FB71" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFF7ECAC0FEC9FB71" box="[160,330,1218,1242]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="Dzuleke village" pageId="24" pageNumber="142" stateProvince="Nagaland">Dzüleke village</location>
|
||
, Sechu-Zubza circle,
|
||
<location id="8EB1604EFF816A5AFDEFCAC0FD58FB71" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFDEFCAC0FD58FB71" box="[561,731,1218,1242]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="Kohima district" pageId="24" pageNumber="142" stateProvince="Nagaland">Kohima district</location>
|
||
,
|
||
<collectingRegion id="49AAF877FF816A5AFD3ACAC0FC0DFB71" box="[740,910,1218,1242]" country="India" name="Nagaland" pageId="24" pageNumber="142">Nagaland state</collectingRegion>
|
||
,
|
||
<collectionCode id="ED7FAE50FF816A5AFC49CAC0FC3CFB71" box="[919,959,1218,1242]" pageId="24" pageNumber="142">NEI</collectionCode>
|
||
, collected by
|
||
<collectionCode id="ED7FAE50FF816A5AFB8BCAC0FB00FB71" box="[1109,1155,1218,1242]" pageId="24" pageNumber="142">RGK</collectionCode>
|
||
and
|
||
<collectionCode id="ED7FAE50FF816A5AFF0DCAE7FF73FB56" box="[211,240,1253,1277]" pageId="24" pageNumber="142">VK</collectionCode>
|
||
on
|
||
<date id="FFD01055FF816A5AFEC0CAE7FE3BFB56" box="[286,440,1253,1277]" pageId="24" pageNumber="142" value="2007-06-18">
|
||
<collectingDate id="EF94E9BDFF816A5AFEC0CAE7FE3BFB56" box="[286,440,1253,1277]" pageId="24" pageNumber="142" value="2007-06-18">18 June 2007</collectingDate>
|
||
</date>
|
||
; tissue sample without associated specimen (
|
||
<specimenCode id="DBC89EEEFF816A5AFC65CAE7FBFEFB56" box="[955,1149,1253,1277]" collectionCode="SDBDU" pageId="24" pageNumber="142">SDBDU 2007.192</specimenCode>
|
||
) from
|
||
<location id="8EB1604EFF816A5AFF3FCB05FE1EFAB4" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFF3FCB05FE1EFAB4" box="[225,413,1287,1311]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="Aradurah Forest" pageId="24" pageNumber="142" stateProvince="Nagaland">Aradurah Forest</location>
|
||
, above
|
||
<location id="8EB1604EFF816A5AFE25CB05FD1DFAB4" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFE25CB05FD1DFAB4" box="[507,670,1287,1311]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="New Ministers" pageId="24" pageNumber="142" stateProvince="Nagaland">New Ministers</location>
|
||
<emphasis id="B91AEA87FF816A5AFD40CB05FD27FAB4" box="[670,676,1287,1311]" italics="true" pageId="24" pageNumber="142">’</emphasis>
|
||
<location id="8EB1604EFF816A5AFD71CB05FD50FAB4" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFD71CB05FD50FAB4" box="[687,723,1287,1311]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="Hill" pageId="24" pageNumber="142" stateProvince="Nagaland">Hill</location>
|
||
(
|
||
<geoCoordinate id="EE5A5052FF816A5AFD3BCB05FCDEFAB4" box="[741,861,1287,1311]" degrees="25.641389" direction="north" orientation="latitude" pageId="24" pageNumber="142" precision="1" value="25.64139">25.641389</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A5052FF816A5AFCB5CB05FC62FAB4" box="[875,993,1287,1311]" degrees="94.094167" direction="east" orientation="longitude" pageId="24" pageNumber="142" precision="1" value="94.09417">94.094167</geoCoordinate>
|
||
,
|
||
<elevation id="0043D1A6FF816A5AFC31CB05FBF5FAB4" box="[1007,1142,1287,1311]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="24" pageNumber="142" unit="m" value="1980.0">
|
||
<quantity id="4C969B70FF816A5AFC31CB05FBC4FAB4" box="[1007,1095,1287,1311]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="24" pageNumber="142" unit="m" value="1980.0">1980 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.),
|
||
<location id="8EB1604EFF816A5AFF7ECB28FEBEFAE9" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFF7ECB28FEBEFAE9" box="[160,317,1322,1346]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="Kohima town" pageId="24" pageNumber="142" stateProvince="Nagaland">Kohima town</location>
|
||
,
|
||
<location id="8EB1604EFF816A5AFE92CB28FE6FFAE9" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFE92CB28FE6FFAE9" box="[332,492,1322,1346]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="Kohima Sadar" pageId="24" pageNumber="142" stateProvince="Nagaland">Kohima Sadar</location>
|
||
circle,
|
||
<location id="8EB1604EFF816A5AFD9DCB28FD70FAE9" LSID="urn:lsid:plazi:treatment:03C78783FF816A5CFE78CCC8FD4DFBE5:8EB1604EFF816A5AFD9DCB28FD70FAE9" box="[579,755,1322,1346]" county="Jakhama village" latitude="25.64139" longLatPrecision="1" longitude="94.09417" municipality="Kohima district" name="Kohima district" pageId="24" pageNumber="142" stateProvince="Nagaland">Kohima district</location>
|
||
,
|
||
<collectingRegion id="49AAF877FF816A5AFCDFCB28FC32FAE9" box="[769,945,1322,1346]" country="India" name="Nagaland" pageId="24" pageNumber="142">Nagaland state</collectingRegion>
|
||
,
|
||
<collectionCode id="ED7FAE50FF816A5AFC1ECB28FC6BFAE9" box="[960,1000,1322,1346]" pageId="24" pageNumber="142">NEI</collectionCode>
|
||
, collected by
|
||
<collectionCode id="ED7FAE50FF816A5AFF7ECB4FFF4DFACE" box="[160,206,1357,1381]" pageId="24" pageNumber="142">RGK</collectionCode>
|
||
and
|
||
<collectionCode id="ED7FAE50FF816A5AFED6CB4FFEB6FACE" box="[264,309,1357,1381]" pageId="24" pageNumber="142">SDB</collectionCode>
|
||
on
|
||
<date id="FFD01055FF816A5AFEBFCB4FFE73FACE" box="[353,496,1357,1381]" pageId="24" pageNumber="142" value="2007-07-10">
|
||
<collectingDate id="EF94E9BDFF816A5AFEBFCB4FFE73FACE" box="[353,496,1357,1381]" pageId="24" pageNumber="142" value="2007-07-10">10 July 2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF816A5BFF7ECB92FF7FF915" blockId="24.[160,1156,1424,1727]" lastBlockId="25.[160,1155,1494,1726]" lastPageId="25" lastPageNumber="143" pageId="24" pageNumber="142">
|
||
<emphasis id="B91AEA87FF816A5AFF7ECB92FD35FA01" bold="true" box="[160,694,1424,1450]" italics="true" pageId="24" pageNumber="142">
|
||
<typeStatus id="54D58837FF816A5AFF7ECB92FE88FA01" box="[160,267,1424,1450]" pageId="24" pageNumber="142" type="holotype">Holotype</typeStatus>
|
||
description (measurements in mm).
|
||
</emphasis>
|
||
Mature male (
|
||
<collectionCode id="ED7FAE50FF816A5AFCBECB90FC08FA01" box="[864,907,1426,1450]" pageId="24" pageNumber="142">SVL</collectionCode>
|
||
37.2) (
|
||
<figureCitation id="13552A10FF816A5AFC06CB90FBB7FA01" box="[984,1076,1426,1450]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="24" pageNumber="142">Figure 5</figureCitation>
|
||
). Head moderately large, as long as wide (
|
||
<collectionCode id="ED7FAE50FF816A5AFDE3CBB7FDE4FA66" box="[573,615,1461,1485]" pageId="24" pageNumber="142">HW</collectionCode>
|
||
13.0, HL 13.0,
|
||
<collectionCode id="ED7FAE50FF816A5AFCC7CBB7FCBAFA66" box="[793,825,1461,1485]" country="Nigeria" lsid="urn:lsid:biocol.org:col:13788" name="Obafemi Awolowo University" pageId="24" pageNumber="142" type="Herbarium">IFE</collectionCode>
|
||
6.3,
|
||
<collectionCode id="ED7FAE50FF816A5AFCA8CBB7FC1BFA66" box="[886,920,1461,1485]" country="Spain" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="24" pageNumber="142">IBE</collectionCode>
|
||
10.3); snout bluntly pointed in dorsal view, obtusely protruding in lateral view, rostral appendage absent (
|
||
<figureCitation id="13552A10FF816A5AFF79CBF8FE81F9B9" box="[167,258,1530,1554]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="24" pageNumber="142">Figure 5</figureCitation>
|
||
(c)); loreal region acute, concave; canthus rostralis angular; dorsal region of snout slightly concave; eye larger than maximum tympanum diameter, and equal to snout length (
|
||
<collectionCode id="ED7FAE50FF816A5AFF28C83DFE91F9FC" box="[246,274,1599,1623]" pageId="24" pageNumber="142">EL</collectionCode>
|
||
4.7,
|
||
<collectionCode id="ED7FAE50FF816A5AFE98C83DFEF0F9FC" box="[326,371,1599,1623]" pageId="24" pageNumber="142">TYD</collectionCode>
|
||
2.8, SL 4.7); eye
|
||
<emphasis id="B91AEA87FF816A5AFDFBC83DFDB1F9FC" box="[549,562,1599,1623]" italics="true" pageId="24" pageNumber="142">–</emphasis>
|
||
tympanum distance (
|
||
<collectionCode id="ED7FAE50FF816A5AFCC1C83DFCC9F9FC" box="[799,842,1599,1623]" pageId="24" pageNumber="142">TYE</collectionCode>
|
||
2.2) shorter than maximum tympanum diameter; tympanum distinctly oval shaped, upper border not concealed by supratympanic ridge; pupil oval in preservation, orientated vertically; nostril oval, orientated laterally, closer to eye than to snout (
|
||
<collectionCode id="ED7FAE50FF816A5AFDABC8A5FD16F914" box="[629,661,1703,1727]" pageId="24" pageNumber="142">EN</collectionCode>
|
||
2.2,
|
||
<collectionCode id="ED7FAE50FF816A5AFD19C8A5FD65F914" box="[711,742,1703,1727]" pageId="24" pageNumber="142">NS</collectionCode>
|
||
2.6); internarial distance greater than eyelid width, and equal to narrowest point between upper eyelids (
|
||
<collectionCode id="ED7FAE50FF806A5BFC7CCBD4FC38FA45" box="[930,955,1494,1518]" pageId="25" pageNumber="143">IN</collectionCode>
|
||
3.9,
|
||
<collectionCode id="ED7FAE50FF806A5BFC2CCBD4FBA4FA45" box="[1010,1063,1494,1518]" pageId="25" pageNumber="143">UEW</collectionCode>
|
||
3.7,
|
||
<collectionCode id="ED7FAE50FF806A5BFB80CBD4FB00FA45" box="[1118,1155,1494,1518]" pageId="25" pageNumber="143">IUE</collectionCode>
|
||
3.9); pineal ocellus not visible externally; vomerine ridges weak, forming long narrow ridges without swollen posterior end, orientated acutely, extending from near anterior choanae border, terminating posterior to choanae, posterior ends equidistant from each other and choanae; vomerine teeth absent; maxillary teeth present; tongue large, covers almost entire floor of mouth, cordiform (with weak notch posteriorly), no medial lingual process.
|
||
</paragraph>
|
||
<caption id="DF11661DFF806A5BFF41CB21FDE4FADE" ID-DOI="http://doi.org/10.5281/zenodo.4608909" ID-Zenodo-Dep="4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="25" pageNumber="143" startId="25.[159,224,1315,1338]" targetBox="[161,1151,137,1284]" targetPageId="25">
|
||
<paragraph id="8BD13695FF806A5BFF41CB21FDE4FADE" blockId="25.[159,1155,1314,1397]" pageId="25" pageNumber="143">
|
||
<emphasis id="B91AEA87FF806A5BFF41CB21FF7AFA91" bold="true" box="[159,249,1315,1338]" pageId="25" pageNumber="143">Figure 5.</emphasis>
|
||
<taxonomicName id="4C6E4D16FF806A5BFEDFCB20FE24FA92" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[257,423,1314,1337]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="25" pageNumber="143" phylum="Chordata" rank="species" species="awuh" status="sp. nov.">
|
||
<emphasis id="B91AEA87FF806A5BFEDFCB20FE24FA92" box="[257,423,1314,1337]" italics="true" pageId="25" pageNumber="143">Megophrys awuh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF806A5BFE73CB21FD83FA91" box="[429,512,1315,1338]" pageId="25" pageNumber="143" rank="species">
|
||
<emphasis id="B91AEA87FF806A5BFE73CB21FE78FA91" bold="true" box="[429,507,1315,1338]" pageId="25" pageNumber="143">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11661DFF836A58FF7ECB75FC95F9A8" ID-DOI="http://doi.org/10.5281/zenodo.4608911" ID-Zenodo-Dep="4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="26" pageNumber="144" startId="26.[160,225,1399,1422]" targetBox="[164,1152,155,1354]" targetPageId="26">
|
||
<paragraph id="8BD13695FF836A58FF7ECB75FC95F9A8" blockId="26.[160,1156,1398,1539]" pageId="26" pageNumber="144">
|
||
<emphasis id="B91AEA87FF836A58FF7ECB75FF7AFA25" bold="true" box="[160,249,1399,1422]" pageId="26" pageNumber="144">Figure 6.</emphasis>
|
||
<taxonomicName id="4C6E4D16FF836A58FF20CB74FE21FA26" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[254,418,1398,1421]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="26" pageNumber="144" phylum="Chordata" rank="species" species="awuh" status="sp. nov.">
|
||
<emphasis id="B91AEA87FF836A58FF20CB74FE21FA26" box="[254,418,1398,1421]" italics="true" pageId="26" pageNumber="144">Megophrys awuh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF836A58FE79CB75FE74FA25" box="[423,503,1399,1422]" pageId="26" pageNumber="144" rank="species">
|
||
<emphasis id="B91AEA87FF836A58FE79CB75FE71FA25" bold="true" box="[423,498,1399,1422]" pageId="26" pageNumber="144">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b
|
||
<emphasis id="B91AEA87FF836A58FC0ECB96FC5EFA00" box="[976,989,1428,1451]" italics="true" pageId="26" pageNumber="144">–</emphasis>
|
||
f). specimens in preservation from Dzüleke village, showing variation: (b
|
||
<emphasis id="B91AEA87FF836A58FD3CCBB3FD6CFA63" box="[738,751,1457,1480]" italics="true" pageId="26" pageNumber="144">–</emphasis>
|
||
d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e
|
||
<emphasis id="B91AEA87FF836A58FC41CBCDFC2FFA4D" box="[927,940,1487,1510]" italics="true" pageId="26" pageNumber="144">–</emphasis>
|
||
f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD13695FF836A59FF1EC854FD4EFE17" blockId="26.[160,1155,1621,1715]" lastBlockId="27.[160,1156,142,1726]" lastPageId="27" lastPageNumber="145" pageId="26" pageNumber="144">
|
||
Forelimbs moderately long, thin (
|
||
<figureCitation id="13552A10FF836A58FDE4C854FD15F9C5" box="[570,662,1622,1646]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="26" pageNumber="144">Figure 5</figureCitation>
|
||
(a
|
||
<emphasis id="B91AEA87FF836A58FD74C857FD34F9C6" box="[682,695,1621,1645]" italics="true" pageId="26" pageNumber="144">–</emphasis>
|
||
b)); forearms slightly enlarged relative to upper forelimbs, forearm shorter than hand length (
|
||
<collectionCode id="ED7FAE50FF836A58FD00C87AFC89F93B" box="[734,778,1656,1680]" pageId="26" pageNumber="144">FAL</collectionCode>
|
||
9.1,
|
||
<collectionCode id="ED7FAE50FF836A58FCE3C87AFCE8F93B" box="[829,875,1656,1680]" country="Germany" lsid="urn:lsid:biocol.org:col:15866" name="Martin-Luther-Universität" pageId="26" pageNumber="144" type="Herbarium">HAL</collectionCode>
|
||
10.1); fingers moderately long, thin, lateral dermal fringes absent (
|
||
<figureCitation id="13552A10FF836A58FD88C899FD33F918" box="[598,688,1691,1715]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="26" pageNumber="144">Figure 5</figureCitation>
|
||
(d)), finger length formula I = II <IV <
|
||
<collectionCode id="ED7FAE50FF836A58FB9AC899FBD4F918" box="[1092,1111,1691,1715]" pageId="26" pageNumber="144">III</collectionCode>
|
||
(
|
||
<collectionCode id="ED7FAE50FF836A58FBBCC899FB00F918" box="[1122,1155,1691,1715]" pageId="26" pageNumber="144">FIL</collectionCode>
|
||
4.0,
|
||
<collectionCode id="ED7FAE50FF826A59FF11CE8CFF76FF0D" box="[207,245,142,166]" pageId="27" pageNumber="145">FIIL</collectionCode>
|
||
4.0,
|
||
<collectionCode id="ED7FAE50FF826A59FEF2CE8CFEDBFF0D" box="[300,344,142,166]" pageId="27" pageNumber="145">FIIIL</collectionCode>
|
||
7.0,
|
||
<collectionCode id="ED7FAE50FF826A59FE51CE8CFE3EFF0D" box="[399,445,142,166]" pageId="27" pageNumber="145">FIVL</collectionCode>
|
||
4.2); interdigital webbing, and subarticular and supernumerary tubercles absent; thenar and outer metacarpal tubercles weak; fingertips rounded, not expanded relative to digit widths, with small oval pads, terminal grooves absent on fingertips. Hindlimbs long, thin (
|
||
<figureCitation id="13552A10FF826A59FE19CEF4FDA2FEA5" box="[455,545,246,270]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="27" pageNumber="145">Figure 5</figureCitation>
|
||
(a
|
||
<emphasis id="B91AEA87FF826A59FDE8CEF4FDC0FEA5" box="[566,579,246,270]" italics="true" pageId="27" pageNumber="145">–</emphasis>
|
||
b)); thigh shorter than shank, and foot (TL 16.7,
|
||
<collectionCode id="ED7FAE50FF826A59FB87CEF4FB07FEA5" box="[1113,1156,246,270]" pageId="27" pageNumber="145">SHL</collectionCode>
|
||
18.3,
|
||
<collectionCode id="ED7FAE50FF826A59FF00CF1BFE8AFE9A" box="[222,265,281,305]" pageId="27" pageNumber="145">FOL</collectionCode>
|
||
17.1); toes long, rounded, distinct lateral dermal fringes absent (
|
||
<figureCitation id="13552A10FF826A59FC22CF1BFBD8FE9A" box="[1020,1115,281,305]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="27" pageNumber="145">Figure 5</figureCitation>
|
||
(e)), relative toe lengths I <II <
|
||
<collectionCode id="ED7FAE50FF826A59FE07CF3EFE6BFEFF" box="[473,488,316,340]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="27" pageNumber="145" type="Museum">V</collectionCode>
|
||
<
|
||
<collectionCode id="ED7FAE50FF826A59FDD4CF3EFD9FFEFF" box="[522,540,316,340]" pageId="27" pageNumber="145">III</collectionCode>
|
||
<IV; toe tips rounded, not dilated, with small circular pads, terminal groves absent on toe tips; webbing rudimentary; inner metatarsal tubercle indistinct; subarticular, supernumerary and outer metatarsal tubercles absent; ridge of callous tissue present on ventral surface of all toes.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF826A59FF1ECFC4FDFEFB87" blockId="27.[160,1156,142,1726]" pageId="27" pageNumber="145">
|
||
Skin of dorsal surfaces of body, limbs, and dorsal and lateral surfaces of head smooth (
|
||
<figureCitation id="13552A10FF826A59FF76CFEBFE86FDAA" box="[168,261,489,513]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="27" pageNumber="145">Figure 5</figureCitation>
|
||
(a)); tympanum smooth with borders weakly raised; ridge or tubercle on outer edge of upper eyelids absent; supratympanic ridge narrow anteriorly, expanding posterior to tympanum to become moderately large and glandular, extending from orbit in a straight line to upper tympanum border, broadly curving obliquely downward along posterior border of tympanum, terminating above forearm insertion on each side; flanks with numerous small to large pustular tubercles (
|
||
<figureCitation id="13552A10FF826A59FD04CC94FCBBFD05" box="[730,824,662,686]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="27" pageNumber="145">Figure 5</figureCitation>
|
||
(a)); dorsolateral ridges thick, well defined, extending from behind supratympanic ridge to above groin on each side; weak,
|
||
<emphasis id="B91AEA87FF826A59FF38CCD9FF68FD58" box="[230,235,731,755]" italics="true" pageId="27" pageNumber="145">‘</emphasis>
|
||
X
|
||
<emphasis id="B91AEA87FF826A59FF25CCD9FE82FD58" box="[251,257,731,755]" italics="true" pageId="27" pageNumber="145">’</emphasis>
|
||
-shaped ridge present on dorsum, its two sides extending posteriorly from above anterior border of tympanum and terminating beyond level of axilla on parietoscapular region, and posterior ends join to dorsolateral ridges on mid dorsum (
|
||
<figureCitation id="13552A10FF826A59FC65CD23FB9BFC92" box="[955,1048,801,825]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="27" pageNumber="145">Figure 5</figureCitation>
|
||
(a)); weak tubercles and short, obliquely transverse ridges present on dorsal surfaces of thighs, shanks and forearms. Gular region, chest, abdomen and ventral surfaces of limbs all smooth; pectoral glands medium-sized, flat, positioned on level of axilla (
|
||
<figureCitation id="13552A10FF826A59FC27CD8BFBDAFC0A" box="[1017,1113,905,929]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="27" pageNumber="145">Figure 5</figureCitation>
|
||
(b)); femoral glands moderately large, slightly raised, positioned equidistant from knee and cloaca on posterior surface of thighs; black dermal asperities restricted to tympanic regions, posterior upper eyelids and posterior-lateral portion of head near supratympanic ridges, absent from all remaining surfaces.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF826A5EFF1ECA34FC44FF47" blockId="27.[160,1156,142,1726]" lastBlockId="28.[160,1155,142,236]" lastPageId="28" lastPageNumber="146" pageId="27" pageNumber="145">
|
||
<emphasis id="B91AEA87FF826A59FF1ECA34FE30FBE5" box="[192,435,1078,1102]" italics="true" pageId="27" pageNumber="145">Colour: In preservation</emphasis>
|
||
(
|
||
<figureCitation id="13552A10FF826A59FE1ECA34FD98FBE5" box="[448,539,1078,1102]" captionStart="Figure 5" captionStartId="25.[159,224,1315,1338]" captionTargetBox="[161,1151,137,1284]" captionTargetId="figure-77@25.[161,1151,137,1284]" captionTargetPageId="25" captionText="Figure 5. Megophrys awuh sp. nov. adult male holotype (BNHS 6070: SVL 37.2 mm) in preservation, from Phek district, Nagaland state, India: (a). dorsal view; (b). ventral view; (c). profile view of head; (d). ventral view of hand; (e). ventral view of foot." figureDoi="http://doi.org/10.5281/zenodo.4608909" httpUri="https://zenodo.org/record/4608909/files/figure.png" pageId="27" pageNumber="145">Figure 5</figureCitation>
|
||
): Dorsal and lateral surfaces of head and body primarily light brownish-grey; solid dark brown triangular marking on dorsal surface of head between eyes; longitudinal brown stripe on mid-dorsal snout, extending from medial anterior border of triangular marking to tip of snout; dorsal ridges and flank tubercles light brownish-cream, some with small bordering dark brown spots and speckling; front of snout and lateral canthus rostralis dark brown; wide vertical dark brown bar below eyes; dark brown blotch extends from posterior canthus, through tympanum to posterior supratympanic ridge on each side; supratympanic ridges light yellowish-cream; dorsal surfaces of forelimbs and hindlimbs primarily light yellowish-grey; two dark brown blotches on anterior lateral surface of forearms; dorsal surface of outer three fingers with dark brown blotches; lateral surfaces of thighs and shanks with dark brown spots and blotches. Throat, chest and abdomen primarily pale grey, lightening to creamishyellow on lateral and posterior surfaces of abdomen and ventral surfaces of forelimbs, thighs and shanks; throat with faint dark brown longitudinal blotches, those most defined extending from rear axis of mandibles to base of forelimbs on each side, and third on midgular region extending onto anterior chest; dark brown blotches form broken stripe laterally along abdomen on both sides; area surrounding vent and posterior surfaces of thighs dark brown; ventral surfaces of tarsus and feet dark brown with contrasting light grey callous ridges on toes; light grey oval patch on region where inner metatarsal tubercle would be on other species; forelimbs and hands ventrally mottled creamishyellow and light greyish-brown, ventral surfaces of digits light grey; pectoral and femoral glands creamish-white.
|
||
<emphasis id="B91AEA87FF856A5EFE75CED6FE64FF47" box="[427,487,212,236]" italics="true" pageId="28" pageNumber="146">In life</emphasis>
|
||
: No images in life taken for this specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374651EFF856A5FFF7ECF15FE44FB50" lastPageId="29" lastPageNumber="147" pageId="28" pageNumber="146" type="description">
|
||
<paragraph id="8BD13695FF856A5EFF7ECF15FC2BFBDA" blockId="28.[160,1156,279,1137]" pageId="28" pageNumber="146">
|
||
<emphasis id="B91AEA87FF856A5EFF7ECF15FE96FE9A" bold="true" box="[160,277,279,305]" italics="true" pageId="28" pageNumber="146">Variation.</emphasis>
|
||
See
|
||
<tableCitation id="C6EC032EFF856A5EFE89CF1BFE2FFE9A" box="[343,428,281,305]" captionStart="Table 2" captionStartId="16.[133,189,446,469]" captionText="Table 2. Specimen morphometrics for all M. (X.) megacephala SG taxa and four paralectotypes of M. parva. Previously published measurements are included for M. megacephala from Mahony et al. (2011), and for M. ancrae and M. oropedion from Mahony et al. (2013). Refer to the Methods and materials section for" pageId="28" pageNumber="146">Table 2</tableCitation>
|
||
for morphometric variation within the specimen series of four adult males and two adult females.
|
||
<typeStatus id="54D58837FF856A5EFDE0CF3EFD28FEFF" box="[574,683,316,340]" pageId="28" pageNumber="146" type="paratype">Paratypes</typeStatus>
|
||
and the referred specimen resemble the
|
||
<typeStatus id="54D58837FF856A5EFF7ECF5DFE87FEDC" box="[160,260,351,375]" pageId="28" pageNumber="146" type="holotype">holotype</typeStatus>
|
||
for most morphological characters with some exceptions: finger length formula typically I = II <IV <III for four specimens, except for BNHS 6069 (I <II <IV <III) and SDBDU 2007.111 (I <II <IV <III); on SDBDU 2007.111, SDBDU 2007.079, and SDBDU 2007.080, vomerine ridges differ from type series by possessing small, weakly raised, hemispherical or ovoid posterior end of vomerine ridges, positioned between to slightly posterior to level of choanae; only SDBDU 2007.079 possesses small vomerine teeth, otherwise absent on all other specimens; posterior edge of tongue on SDBDU 2007.079 and SDBDU 2007.080 appears rounded rather than notched (which may be artefact of preservation); SDBDU 2007.079 and SDBDU 2007.080 (
|
||
<figureCitation id="13552A10FF856A5EFDBCCC76FD3DFD27" box="[610,702,628,652]" captionStart="Figure 6" captionStartId="26.[160,225,1399,1422]" captionTargetBox="[164,1152,155,1354]" captionTargetId="figure-52@26.[163,1153,154,1368]" captionTargetPageId="26" captionText="Figure 6. Megophrys awuh sp. nov. referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b–f). specimens in preservation from Dzüleke village, showing variation: (b–d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e–f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view." figureDoi="http://doi.org/10.5281/zenodo.4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="28" pageNumber="146">Figure 6</figureCitation>
|
||
(c,e)) have brown
|
||
<emphasis id="B91AEA87FF856A5EFC59CC76FC0EFD27" box="[903,909,628,652]" italics="true" pageId="28" pageNumber="146">‘</emphasis>
|
||
V
|
||
<emphasis id="B91AEA87FF856A5EFC42CC76FC21FD27" box="[924,930,628,652]" italics="true" pageId="28" pageNumber="146">’</emphasis>
|
||
-shaped marking on anterior dorsum surrounding parietoscapular ridges; SDBDU 2007.111 possesses small tubercle on outer upper eyelid, SDBDU 2007.079 and SDBDU 2007.080 have short ridge on upper eyelid (
|
||
<figureCitation id="13552A10FF856A5EFEBFCCDEFE3DFD5F" box="[353,446,732,756]" captionStart="Figure 6" captionStartId="26.[160,225,1399,1422]" captionTargetBox="[164,1152,155,1354]" captionTargetId="figure-52@26.[163,1153,154,1368]" captionTargetPageId="26" captionText="Figure 6. Megophrys awuh sp. nov. referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b–f). specimens in preservation from Dzüleke village, showing variation: (b–d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e–f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view." figureDoi="http://doi.org/10.5281/zenodo.4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="28" pageNumber="146">Figure 6</figureCitation>
|
||
(a)); parietoscapular and dorsal ridge configuration differs from
|
||
<typeStatus id="54D58837FF856A5EFF7ECCFDFE87FCBC" box="[160,260,767,791]" pageId="28" pageNumber="146" type="holotype">holotype</typeStatus>
|
||
as follows: SDBDU 2007.111 has faintly raised, longitudinal
|
||
<emphasis id="B91AEA87FF856A5EFC6FCCFCFC34FCBD" box="[945,951,766,790]" italics="true" pageId="28" pageNumber="146">‘</emphasis>
|
||
>-<
|
||
<emphasis id="B91AEA87FF856A5EFC00CCFCFC67FCBD" box="[990,996,766,790]" italics="true" pageId="28" pageNumber="146">’</emphasis>
|
||
configuration (
|
||
<figureCitation id="13552A10FF856A5EFF76CD23FE80FC92" box="[168,259,801,825]" captionStart="Figure 6" captionStartId="26.[160,225,1399,1422]" captionTargetBox="[164,1152,155,1354]" captionTargetId="figure-52@26.[163,1153,154,1368]" captionTargetPageId="26" captionText="Figure 6. Megophrys awuh sp. nov. referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b–f). specimens in preservation from Dzüleke village, showing variation: (b–d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e–f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view." figureDoi="http://doi.org/10.5281/zenodo.4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="28" pageNumber="146">Figure 6</figureCitation>
|
||
(a)), SDBDU 2007.079 and SDBDU 2007.080 possess only
|
||
<emphasis id="B91AEA87FF856A5EFCA0CD23FC07FC92" box="[894,900,801,825]" italics="true" pageId="28" pageNumber="146">‘</emphasis>
|
||
V
|
||
<emphasis id="B91AEA87FF856A5EFC4CCD23FC1BFC92" box="[914,920,801,825]" italics="true" pageId="28" pageNumber="146">’</emphasis>
|
||
-shaped parietoscapular ridges (
|
||
<figureCitation id="13552A10FF856A5EFEFACD46FEFCFCF7" box="[292,383,836,860]" captionStart="Figure 6" captionStartId="26.[160,225,1399,1422]" captionTargetBox="[164,1152,155,1354]" captionTargetId="figure-52@26.[163,1153,154,1368]" captionTargetPageId="26" captionText="Figure 6. Megophrys awuh sp. nov. referred specimens from Kohima district, Nagaland state, India: (a). adult male (SDBDU 2007.111: SVL 35.7 mm) in life from Jakhama village; (b–f). specimens in preservation from Dzüleke village, showing variation: (b–d). adult female (SDBDU 2007.080: SVL 48.0 mm): (b). profile view of head; (c). dorsolateral view; (d). ventral view; (e–f). adult male (SDBDU 2007.079: SVL 41.1 mm): (e). dorsolateral view; (f). ventral view." figureDoi="http://doi.org/10.5281/zenodo.4608911" httpUri="https://zenodo.org/record/4608911/files/figure.png" pageId="28" pageNumber="146">Figure 6</figureCitation>
|
||
(c,e)); BNHS 6069, BNHS 6071 and SDBDU 2007.080 have no observed dermal asperities, SDBDU 2007.111 has white dermal asperities found only on middorsum, increasing in density posteriorly to above vent, on SDBDU 2007.079, asperities only found on posterior surface of upper eyelids; SDBDU 2007.111, SDBDU 2007.079 and SDBDU 2007.080 are mid to light brown dorsally; on BNHS 6069 and BNHS 6071, triangular brown marking on dorsal surface of the head is faint, ventrolateral dark brown blotches are smaller and less dense, and do not form broken stripe laterally along abdomen on both sides, and ventral shanks and thighs with dark brown, dense speckling; BNHS 6069 has small dark brown spots spread over entire abdomen.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF856A5EFF7ECA9FFBE2FA01" blockId="28.[160,1156,1181,1450]" pageId="28" pageNumber="146">
|
||
<emphasis id="B91AEA87FF856A5EFF7ECA9FFE79FB1C" bold="true" box="[160,506,1181,1207]" italics="true" pageId="28" pageNumber="146">Secondary sexual characters.</emphasis>
|
||
<emphasis id="B91AEA87FF856A5EFDD7CA9DFDC4FB1C" box="[521,583,1183,1207]" italics="true" pageId="28" pageNumber="146">Males</emphasis>
|
||
: weakly raised nuptial pads covered with brown or black micro-asperities, covering most of dorsal surface of base of Finger I, narrowing distally, extending onto base of distal phalange on inner dorsal side; on Finger II, nuptial pad small, oval shaped, widest proximally, extending just onto basal phalange on inner dorsal side; external vocal sac indistinct; internal vocal slits present near rear of lower mandible; forearms slightly enlarged relative to upper forelimbs; fleshy projection above cloaca absent.
|
||
<emphasis id="B91AEA87FF856A5EFF2ACB6DFEC9FA2C" box="[244,330,1391,1415]" italics="true" pageId="28" pageNumber="146">Females</emphasis>
|
||
: nuptial pads, vocal slits, cloacal projection all absent; forearms not enlarged relative to upper forelimbs; both females contained moderately large, pigmentless ova.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF856A5FFF7ECBD7FD08FF47" blockId="28.[160,1157,1493,1727]" lastBlockId="29.[160,1156,142,1275]" lastPageId="29" lastPageNumber="147" pageId="28" pageNumber="146">
|
||
<emphasis id="B91AEA87FF856A5EFF7ECBD7FE6DFA44" bold="true" box="[160,494,1493,1519]" italics="true" pageId="28" pageNumber="146">Morphological comparisons.</emphasis>
|
||
<taxonomicName id="4C6E4D16FF856A5EFE20CBD5FD35FA44" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[510,694,1495,1519]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="28" pageNumber="146" phylum="Chordata" rank="species" species="awuh" status="sp. nov.">
|
||
<emphasis id="B91AEA87FF856A5EFE20CBD5FD35FA44" box="[510,694,1495,1519]" italics="true" pageId="28" pageNumber="146">Megophrys awuh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF856A5EFD64CBD5FC90FA44" box="[698,787,1495,1519]" pageId="28" pageNumber="146" rank="species">
|
||
<emphasis id="B91AEA87FF856A5EFD64CBD5FC8EFA44" bold="true" box="[698,781,1495,1519]" pageId="28" pageNumber="146">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(adult males,
|
||
<emphasis id="B91AEA87FF856A5EFC76CBD5FC3AFA44" box="[936,953,1495,1519]" italics="true" pageId="28" pageNumber="146">N</emphasis>
|
||
= 4; adult females,
|
||
<emphasis id="B91AEA87FF856A5EFF7ECBF8FF32F9B9" box="[160,177,1530,1554]" italics="true" pageId="28" pageNumber="146">N</emphasis>
|
||
= 2) differs from the following species that have not yet been assigned to a subgenus or species group through molecular analyses: from
|
||
<taxonomicName id="4C6E4D16FF856A5EFD6EC81FFC98F99E" authorityName="Mahony" authorityYear="2011" box="[688,795,1565,1589]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="28" pageNumber="146" phylum="Chordata" rank="species" species="damrei">
|
||
<emphasis id="B91AEA87FF856A5EFD6EC81FFC98F99E" box="[688,795,1565,1589]" italics="true" pageId="28" pageNumber="146">M. damrei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6E4D16FF856A5EFC91C81FFB8DF99E" authorityName="Tian and Sun" authorityYear="1995" box="[847,1038,1565,1589]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="28" pageNumber="146" phylum="Chordata" rank="species" species="shuichengensis">
|
||
<emphasis id="B91AEA87FF856A5EFC91C81FFB8DF99E" box="[847,1038,1565,1589]" italics="true" pageId="28" pageNumber="146">M. shuichengensis</emphasis>
|
||
</taxonomicName>
|
||
by smaller adult size, male SVL
|
||
<quantity id="4C969B70FF856A5EFEA2C83DFD9AF9FC" box="[380,537,1599,1623]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" metricValueMax="4.11" metricValueMin="3.57" pageId="28" pageNumber="146" unit="mm" value="38.4" valueMax="41.1" valueMin="35.7">
|
||
35.7
|
||
<emphasis id="B91AEA87FF856A5EFE74C83DFE34F9FC" box="[426,439,1599,1623]" italics="true" pageId="28" pageNumber="146">–</emphasis>
|
||
41.1 mm
|
||
</quantity>
|
||
, female SVL 43.7
|
||
<emphasis id="B91AEA87FF856A5EFD10C83DFD58F9FC" box="[718,731,1599,1623]" italics="true" pageId="28" pageNumber="146">–</emphasis>
|
||
48.0 mm (vs.
|
||
<taxonomicName id="4C6E4D16FF856A5EFCB5C83DFC55F9FC" authorityName="Mahony" authorityYear="2011" box="[875,982,1599,1623]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="28" pageNumber="146" phylum="Chordata" rank="species" species="damrei">
|
||
<emphasis id="B91AEA87FF856A5EFCB5C83DFC55F9FC" box="[875,982,1599,1623]" italics="true" pageId="28" pageNumber="146">M. damrei</emphasis>
|
||
</taxonomicName>
|
||
: adult male SVL
|
||
<quantity id="4C969B70FF856A5EFF7EC860FEBDF9D1" box="[160,318,1634,1658]" metricMagnitude="-2" metricUnit="m" metricValue="5.24" metricValueMax="5.71" metricValueMin="4.77" pageId="28" pageNumber="146" unit="mm" value="52.4" valueMax="57.1" valueMin="47.7">
|
||
47.7
|
||
<emphasis id="B91AEA87FF856A5EFF13C860FF59F9D1" box="[205,218,1634,1658]" italics="true" pageId="28" pageNumber="146">–</emphasis>
|
||
57.1 mm
|
||
</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF856A5EFE97C860FED9F9D1" box="[329,346,1634,1658]" italics="true" pageId="28" pageNumber="146">N</emphasis>
|
||
= 7, adult female SVL
|
||
<quantity id="4C969B70FF856A5EFD8BC860FD3BF9D1" box="[597,696,1634,1658]" metricMagnitude="-2" metricUnit="m" metricValue="6.909999999999999" pageId="28" pageNumber="146" unit="mm" value="69.1">69.1 mm</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF856A5EFD1DC860FD57F9D1" box="[707,724,1634,1658]" italics="true" pageId="28" pageNumber="146">N</emphasis>
|
||
= 1 [Mahony 2011;
|
||
<bibRefCitation id="EFFF4B64FF856A5EFC6AC860FBF4F9D1" author="Neang T & Chhin S & Meang M & Hun S" box="[948,1143,1634,1658]" pageId="28" pageNumber="146" pagination="66 - 72" refId="ref47315" refString="Neang T, Chhin S, Meang M, Hun S. 2013. Confirmation of three species of megophryid frogs (Amphibia: Megophryidae) from the Cardamom Mountains of Southwest Cambodia, with the rediscovery of a long lost species. Cambodian J Nat Hist. 2013 (2): 66 - 72." type="journal article" year="2013">Neang et al. 2013</bibRefCitation>
|
||
];
|
||
<taxonomicName id="4C6E4D16FF856A5EFF7EC887FEDCF936" authorityName="Tian and Sun" authorityYear="1995" box="[160,351,1669,1693]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="28" pageNumber="146" phylum="Chordata" rank="species" species="shuichengensis">
|
||
<emphasis id="B91AEA87FF856A5EFF7EC887FEDCF936" box="[160,351,1669,1693]" italics="true" pageId="28" pageNumber="146">M. shuichengensis</emphasis>
|
||
</taxonomicName>
|
||
: adult male SVL 102.0
|
||
<emphasis id="B91AEA87FF856A5EFD90C886FDD8F937" box="[590,603,1668,1692]" italics="true" pageId="28" pageNumber="146">–</emphasis>
|
||
<quantity id="4C969B70FF856A5EFD85C887FD49F936" box="[603,714,1669,1693]" metricMagnitude="-1" metricUnit="m" metricValue="1.183" pageId="28" pageNumber="146" unit="mm" value="118.3">118.3 mm</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF856A5EFD0DC887FD67F936" box="[723,740,1669,1693]" italics="true" pageId="28" pageNumber="146">N</emphasis>
|
||
= 8, adult female SVL
|
||
<quantity id="4C969B70FF856A5EFC08C887FB03F936" box="[982,1152,1668,1693]" metricMagnitude="-1" metricUnit="m" metricValue="1.077" metricValueMax="1.156" metricValueMin="0.998" pageId="28" pageNumber="146" unit="mm" value="107.69999999999999" valueMax="115.6" valueMin="99.8">
|
||
99.8
|
||
<emphasis id="B91AEA87FF856A5EFBDAC886FB92F937" box="[1028,1041,1668,1692]" italics="true" pageId="28" pageNumber="146">–</emphasis>
|
||
115.6 mm
|
||
</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF856A5EFF7EC8A5FF32F914" box="[160,177,1703,1727]" italics="true" pageId="28" pageNumber="146">N</emphasis>
|
||
= 7 [
|
||
<bibRefCitation id="EFFF4B64FF856A5EFF3BC8A5FE08F914" author="Tian Y & Gu X & Sun A" box="[229,395,1703,1727]" pageId="28" pageNumber="146" pagination="462 - 466" refId="ref48867" refString="Tian Y, Gu X, Sun A. 2000. A new species of Megophrys in China (Amphibia: Pelobatidae). Acta Zootaxon Sin. 25 (4): 462 - 466." type="journal article" year="2000">Tian et al. 2000</bibRefCitation>
|
||
]); from
|
||
<taxonomicName id="4C6E4D16FF856A5EFE00C8A5FD9EF914" authorityName="Yang, Wang and Wang" authorityYear="2018" box="[478,541,1703,1727]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="28" pageNumber="146" phylum="Chordata" rank="species" species="feii">
|
||
<emphasis id="B91AEA87FF856A5EFE00C8A5FD9EF914" box="[478,541,1703,1727]" italics="true" pageId="28" pageNumber="146">M. feii</emphasis>
|
||
</taxonomicName>
|
||
by larger adult size, male SVL
|
||
<quantity id="4C969B70FF856A5EFCBBC8A5FB81F914" box="[869,1026,1703,1727]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" metricValueMax="4.11" metricValueMin="3.57" pageId="28" pageNumber="146" unit="mm" value="38.4" valueMax="41.1" valueMin="35.7">
|
||
35.7
|
||
<emphasis id="B91AEA87FF856A5EFC4DC8A5FC23F914" box="[915,928,1703,1727]" italics="true" pageId="28" pageNumber="146">–</emphasis>
|
||
41.1 mm
|
||
</quantity>
|
||
, female SVL 43.7
|
||
<emphasis id="B91AEA87FF846A5FFF13CE8CFF59FF0D" box="[205,218,142,166]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
48.0 mm (vs. adult male SVL
|
||
<quantity id="4C969B70FF846A5FFDFDCE8CFD41FF0D" box="[547,706,142,166]" metricMagnitude="-2" metricUnit="m" metricValue="2.47" metricValueMax="2.51" metricValueMin="2.43" pageId="29" pageNumber="147" unit="mm" value="24.7" valueMax="25.1" valueMin="24.3">
|
||
24.3
|
||
<emphasis id="B91AEA87FF846A5FFD8ECE8CFDDEFF0D" box="[592,605,142,166]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
25.1 mm
|
||
</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF846A5FFD10CE8CFD5CFF0D" box="[718,735,142,166]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 4, adult female SVL
|
||
<quantity id="4C969B70FF846A5FFC3FCE8CFB03FF0D" box="[993,1152,142,166]" metricMagnitude="-2" metricUnit="m" metricValue="2.8549999999999995" metricValueMax="2.8899999999999997" metricValueMin="2.82" pageId="29" pageNumber="147" unit="mm" value="28.549999999999997" valueMax="28.9" valueMin="28.2">
|
||
28.2
|
||
<emphasis id="B91AEA87FF846A5FFBD0CE8CFB98FF0D" box="[1038,1051,142,166]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
28.9 mm
|
||
</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF846A5FFF7ECEB3FF32FF62" box="[160,177,177,201]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 2), lateral dermal fringes on toes absent (vs. present), and nuptial pads on fingers of males present (vs. absent) (
|
||
<bibRefCitation id="EFFF4B64FF846A5FFE15CED6FDFCFF47" author="Yang JH & Wang J & Wang YY" box="[459,639,212,236]" pageId="29" pageNumber="147" pagination="325 - 338" refId="ref49371" refString="Yang JH, Wang J, Wang YY. 2018. A new species of the genus Megophrys (Anura: Megophryidae) from Yunnan Province, China. Zootaxa. 4413 (2): 325 - 338. doi: 10.11646 / zootaxa. 4413.2.5." type="journal article" year="2018">Yang et al. 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF846A5FFF1ECEF4FBAEFE32" blockId="29.[160,1156,142,1275]" pageId="29" pageNumber="147">
|
||
<taxonomicName id="4C6E4D16FF846A5FFF1ECEF4FEF8FEA5" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[192,379,246,270]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="awuh" status="sp. nov.">
|
||
<emphasis id="B91AEA87FF846A5FFF1ECEF4FEF8FEA5" box="[192,379,246,270]" italics="true" pageId="29" pageNumber="147">Megophrys awuh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF846A5FFE5CCEF4FE5DFEA5" box="[386,478,246,270]" pageId="29" pageNumber="147" rank="species">
|
||
<emphasis id="B91AEA87FF846A5FFE5CCEF4FE5BFEA5" bold="true" box="[386,472,246,270]" pageId="29" pageNumber="147">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C6E4D16FF846A5FFDB2CEF4FD4FFEA5" baseAuthorityName="Boulenger" baseAuthorityYear="1893" box="[620,716,246,270]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" sensu="stricto" species="parva">
|
||
<emphasis id="B91AEA87FF846A5FFDB2CEF4FD4FFEA5" box="[620,716,246,270]" italics="true" pageId="29" pageNumber="147">M. parva</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF846A5FFD0DCEF4FD71FEA5" box="[723,754,246,270]" pageId="29" pageNumber="147" sensu="stricto">
|
||
<emphasis id="B91AEA87FF846A5FFD0DCEF4FD6FFEA5" box="[723,748,246,270]" italics="true" pageId="29" pageNumber="147">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by larger tympanum relative to eye length, TYD/ED 55.8
|
||
<emphasis id="B91AEA87FF846A5FFE5ECF1BFE0EFE9A" box="[384,397,281,305]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
72.7%,
|
||
<emphasis id="B91AEA87FF846A5FFE03CF1BFE6DFE9A" box="[477,494,281,305]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. TYD/ED 40.0
|
||
<emphasis id="B91AEA87FF846A5FFD0BCF1BFD61FE9A" box="[725,738,281,305]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
55.0%,
|
||
<emphasis id="B91AEA87FF846A5FFCEDCF1BFCC7FE9A" box="[819,836,281,305]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 4), longer relative forearm length, FAL/SVL 24.5
|
||
<emphasis id="B91AEA87FF846A5FFE59CF39FE17FEF8" box="[391,404,315,339]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
26.3%,
|
||
<emphasis id="B91AEA87FF846A5FFE3BCF3EFE75FEFF" box="[485,502,316,340]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. FAL/SVL 20.9
|
||
<emphasis id="B91AEA87FF846A5FFD36CF39FD76FEF8" box="[744,757,315,339]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
23.9%,
|
||
<emphasis id="B91AEA87FF846A5FFC98CF3EFCD4FEFF" box="[838,855,316,340]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 4), longer relative shank length, SHL/SVL 49.2
|
||
<emphasis id="B91AEA87FF846A5FFE4BCF5CFE21FEDD" box="[405,418,350,374]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
54.9%,
|
||
<emphasis id="B91AEA87FF846A5FFE27CF5CFD89FEDD" box="[505,522,350,374]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. SHL/SVL 42.8
|
||
<emphasis id="B91AEA87FF846A5FFCC2CF5CFCAAFEDD" box="[796,809,350,374]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
48.2%,
|
||
<emphasis id="B91AEA87FF846A5FFC5ECF5CFC12FEDD" box="[896,913,350,374]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 4), fingertips not expanded relative to finger width (vs. slightly expanded relative to finger width).
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF846A5FFF1ECFA6FE5EFB1D" blockId="29.[160,1156,142,1275]" pageId="29" pageNumber="147">
|
||
<taxonomicName id="4C6E4D16FF846A5FFF1ECFA6FEFBFE17" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[192,376,420,444]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="awuh" status="sp. nov.">
|
||
<emphasis id="B91AEA87FF846A5FFF1ECFA6FEFBFE17" box="[192,376,420,444]" italics="true" pageId="29" pageNumber="147">Megophrys awuh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF846A5FFEA1CFA6FE59FE17" box="[383,474,420,444]" pageId="29" pageNumber="147" rank="species">
|
||
<emphasis id="B91AEA87FF846A5FFEA1CFA6FE57FE17" bold="true" box="[383,468,420,444]" pageId="29" pageNumber="147">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from species molecularly assigned to the
|
||
<emphasis id="B91AEA87FF846A5FFC2BCFA6FB89FE17" box="[1013,1034,420,444]" italics="true" pageId="29" pageNumber="147">M</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E4D16FF846A5FFBC8CFA6FE99FE75" authority="SG" authorityName="Mahony, Sengupta, Kamei and Biju" authorityYear="2011" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="megacephala" subGenus="Xenophrys">
|
||
(
|
||
<emphasis id="B91AEA87FF846A5FFBC3CFA6FBA8FE17" box="[1053,1067,420,444]" italics="true" pageId="29" pageNumber="147">X</emphasis>
|
||
.)
|
||
<emphasis id="B91AEA87FF846A5FFBE0CFA6FF77FE75" italics="true" pageId="29" pageNumber="147">megacephala</emphasis>
|
||
SG
|
||
</taxonomicName>
|
||
based on the following combinations of characters: differs from
|
||
<taxonomicName id="4C6E4D16FF846A5FFC2ACFC4FBE3FE75" authorityName="Mahony, Teeling & Biju" authorityYear="2013" box="[1012,1120,454,478]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="ancrae">
|
||
<emphasis id="B91AEA87FF846A5FFC2ACFC4FBE3FE75" box="[1012,1120,454,478]" italics="true" pageId="29" pageNumber="147">
|
||
M.
|
||
<materialsCitation id="3B063CC8FF846A5FFBC9CFC4FBE3FE75" box="[1047,1120,454,478]" collectionCode="HW" pageId="29" pageNumber="147" specimenCode="HW 45.3, 49.6" specimenCount="1">ancrae</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
by finger and toe tips not expanded relative to digit width (vs. digit tips distinctly expanded terminally), small raised bump or short ridge on outer upper eyelids sometimes present (vs. short, sharp horn-like tubercle on outer upper eyelids present), circummarginal patch of demal asperities ventrally on mandible of males absent (vs. present on seven of eight males examined), and typically narrower snout, IFE/
|
||
<specimenCode id="DBC89EEEFF846A5FFD54CC71FD67FD20" box="[650,740,627,651]" collectionCode="HW" pageId="29" pageNumber="147">HW 45.3</specimenCode>
|
||
<emphasis id="B91AEA87FF846A5FFD3ACC71FD72FD20" box="[740,753,627,651]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
<specimenCode id="DBC89EEEFF846A5FFD2FCC71FCA2FD20" box="[753,801,627,651]" collectionCode="HW" pageId="29" pageNumber="147">49.6</specimenCode>
|
||
%,
|
||
<emphasis id="B91AEA87FF846A5FFCE1CC71FCD3FD20" box="[831,848,627,651]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. 49.6
|
||
<emphasis id="B91AEA87FF846A5FFC0BCC71FC61FD20" box="[981,994,627,651]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
57.1%,
|
||
<emphasis id="B91AEA87FF846A5FFBF1CC71FBC3FD20" box="[1071,1088,627,651]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 10); differs from
|
||
<taxonomicName id="4C6E4D16FF846A5FFEEACC94FE6EFD05" authorityName="Mahony, Sengupta, Kamei and Biju" authorityYear="2011" box="[308,493,662,686]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="megacephala">
|
||
<emphasis id="B91AEA87FF846A5FFEEACC94FE6EFD05" box="[308,493,662,686]" italics="true" pageId="29" pageNumber="147">
|
||
M.
|
||
<materialsCitation id="3B063CC8FF846A5FFE82CC94FE6EFD05" box="[348,493,662,686]" collectionCode="SVL, HW" pageId="29" pageNumber="147" specimenCode="SVL 31.5, 36.7, SVL 40.2, 45.1" specimenCount="1">megacephala</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
by smaller adult male size, SVL
|
||
<quantity id="4C969B70FF846A5FFCA4CC94FB9FFD05" box="[890,1052,662,686]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" metricValueMax="4.11" metricValueMin="3.57" pageId="29" pageNumber="147" unit="mm" value="38.4" valueMax="41.1" valueMin="35.7">
|
||
35.7
|
||
<emphasis id="B91AEA87FF846A5FFC79CC94FC37FD05" box="[935,948,662,686]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
41.1 mm
|
||
</quantity>
|
||
(vs. SVL
|
||
<quantity id="4C969B70FF846A5FFF7ECCBBFEC3FD7A" box="[160,320,697,721]" metricMagnitude="-2" metricUnit="m" metricValue="4.965" metricValueMax="5.34" metricValueMin="4.59" pageId="29" pageNumber="147" unit="mm" value="49.65" valueMax="53.4" valueMin="45.9">
|
||
45.9
|
||
<emphasis id="B91AEA87FF846A5FFF13CCBBFF59FD7A" box="[205,218,697,721]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
53.4 mm
|
||
</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF846A5FFE90CCBBFEDCFD7A" box="[334,351,697,721]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 7), and considerably narrower head, HW/
|
||
<specimenCode id="DBC89EEEFF846A5FFC8FCCBBFC37FD7A" box="[849,948,697,721]" collectionCode="SVL" pageId="29" pageNumber="147">SVL 31.5</specimenCode>
|
||
<emphasis id="B91AEA87FF846A5FFC6ACCBBFC42FD7A" box="[948,961,697,721]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
<specimenCode id="DBC89EEEFF846A5FFC1FCCBBFC72FD7A" box="[961,1009,697,721]" collectionCode="HW" pageId="29" pageNumber="147">36.7</specimenCode>
|
||
%,
|
||
<emphasis id="B91AEA87FF846A5FFBCDCCBBFBA7FD7A" box="[1043,1060,697,721]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. HW/
|
||
<specimenCode id="DBC89EEEFF846A5FFF13CCD9FEB3FD58" box="[205,304,731,755]" collectionCode="SVL" pageId="29" pageNumber="147">SVL 40.2</specimenCode>
|
||
<emphasis id="B91AEA87FF846A5FFEF1CCD9FEBFFD58" box="[303,316,731,755]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
<specimenCode id="DBC89EEEFF846A5FFEE2CCD9FEE9FD58" box="[316,362,731,755]" collectionCode="HW" pageId="29" pageNumber="147">45.1</specimenCode>
|
||
%,
|
||
<emphasis id="B91AEA87FF846A5FFE50CCD9FE1CFD58" box="[398,415,731,755]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 12); differs from
|
||
<taxonomicName id="4C6E4D16FF846A5FFD5ECCD9FC92FD58" authorityName="Mahony, Teeling and Biju" authorityYear="2013" box="[640,785,731,755]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="oropedion">
|
||
<emphasis id="B91AEA87FF846A5FFD5ECCD9FC92FD58" box="[640,785,731,755]" italics="true" pageId="29" pageNumber="147">
|
||
M.
|
||
<materialsCitation id="3B063CC8FF846A5FFD7ACCD9FC92FD58" box="[676,785,731,755]" collectionCode="SHL" pageId="29" pageNumber="147" specimenCode="SVL 49.2, 54.9, SVL 41.1, 48.4" specimenCount="11">oropedion</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
by slightly longer relative shank length, SHL/
|
||
<specimenCode id="DBC89EEEFF846A5FFEFBCCFCFE01FCBD" box="[293,386,766,790]" collectionCode="SVL" pageId="29" pageNumber="147">SVL 49.2</specimenCode>
|
||
<emphasis id="B91AEA87FF846A5FFE5FCCFCFE0DFCBD" box="[385,398,766,790]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
<specimenCode id="DBC89EEEFF846A5FFE50CCFCFE3DFCBD" box="[398,446,766,790]" collectionCode="SHL" pageId="29" pageNumber="147">54.9</specimenCode>
|
||
%,
|
||
<emphasis id="B91AEA87FF846A5FFE02CCFCFE6EFCBD" box="[476,493,766,790]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. SHL/
|
||
<specimenCode id="DBC89EEEFF846A5FFDA9CCFCFD56FCBD" box="[631,725,766,790]" collectionCode="SVL" pageId="29" pageNumber="147">SVL 41.1</specimenCode>
|
||
<emphasis id="B91AEA87FF846A5FFD0ACCFCFD62FCBD" box="[724,737,766,790]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
<specimenCode id="DBC89EEEFF846A5FFD3FCCFCFC92FCBD" box="[737,785,766,790]" collectionCode="SHL" pageId="29" pageNumber="147">48.4</specimenCode>
|
||
%,
|
||
<emphasis id="B91AEA87FF846A5FFCF1CCFCFCC3FCBD" box="[815,832,766,790]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 12), vomerine teeth usually absent, five of six specimens (vs. usually present, nine of
|
||
<specimenCount id="9D68FD1CFF846A5FFCC6CD23FC2DFC92" box="[792,942,801,825]" pageId="29" pageNumber="147" type="generic">11 specimens</specimenCount>
|
||
examined), dermal asperities, if present, sparsely distributed, may be found on tympanic region, mid to posterior dorsum, and posterior upper eyelids (vs. extensively distributed on most dorsal, and some ventral surfaces of head, body and limbs); differs from
|
||
<taxonomicName id="4C6E4D16FF846A5FFC9ACD8BFC49FC0A" baseAuthorityName="Mathew & Sen" baseAuthorityYear="2007" box="[836,970,905,929]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="serchhipii">
|
||
<emphasis id="B91AEA87FF846A5FFC9ACD8BFC49FC0A" box="[836,970,905,929]" italics="true" pageId="29" pageNumber="147">
|
||
M.
|
||
<materialsCitation id="3B063CC8FF846A5FFCB8CD8BFC49FC0A" box="[870,970,905,929]" collectionCode="FIIL, FIVL, FOL, SVL" pageId="29" pageNumber="147" specimenCode="SVL 44.3" specimenCount="1">serchhipii</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
by FIIL≤FIVL (vs. FIVL<FIIL), dorsolateral ridges typically extending for entire trunk length (vs. absent, or short if present, extending <40% trunk length), tubercles and short ridges on dorsum absent (vs. present, typically extensive), and typically shorter FOL/SVL (mean±SD) 49.2 ± 3.1%, range 46.0
|
||
<emphasis id="B91AEA87FF846A5FFF13CA11FF59FB80" box="[205,218,1043,1067]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
54.9%,
|
||
<emphasis id="B91AEA87FF846A5FFEF1CA11FEC3FB80" box="[303,320,1043,1067]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. FOL/
|
||
<specimenCode id="DBC89EEEFF846A5FFE3ACA11FDCBFB80" box="[484,584,1043,1067]" collectionCode="SVL" pageId="29" pageNumber="147">SVL 44.3</specimenCode>
|
||
± 2.2%, range 39.5
|
||
<emphasis id="B91AEA87FF846A5FFCEECA11FCBEFB80" box="[816,829,1043,1067]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
48.7%,
|
||
<emphasis id="B91AEA87FF846A5FFC4FCA11FC21FB80" box="[913,930,1043,1067]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 32); differs from
|
||
<taxonomicName id="4C6E4D16FF846A5FFF7ECA34FEE5FBE5" baseAuthorityName="Mathew & Sen" baseAuthorityYear="2007" box="[160,358,1078,1102]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="29" pageNumber="147" phylum="Chordata" rank="species" species="zunhebotoensis">
|
||
<emphasis id="B91AEA87FF846A5FFF7ECA34FEE5FBE5" box="[160,358,1078,1102]" italics="true" pageId="29" pageNumber="147">
|
||
M.
|
||
<materialsCitation id="3B063CC8FF846A5FFF1FCA34FEE5FBE5" box="[193,358,1078,1102]" collectionCode="SVL, TYD" pageId="29" pageNumber="147" specimenCode="ED 50.5" specimenCount="1">zunhebotoensis</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
by larger adult male size, SVL
|
||
<quantity id="4C969B70FF846A5FFD69CA34FCD0FBE5" box="[695,851,1078,1102]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" metricValueMax="4.11" metricValueMin="3.57" pageId="29" pageNumber="147" unit="mm" value="38.4" valueMax="41.1" valueMin="35.7">
|
||
35.7
|
||
<emphasis id="B91AEA87FF846A5FFD3BCA34FD71FBE5" box="[741,754,1078,1102]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
41.1 mm
|
||
</quantity>
|
||
(vs.
|
||
<quantity id="4C969B70FF846A5FFC5BCA34FBA1FBE5" box="[901,1058,1078,1102]" metricMagnitude="-2" metricUnit="m" metricValue="3.1149999999999998" metricValueMax="3.3899999999999997" metricValueMin="2.84" pageId="29" pageNumber="147" unit="mm" value="31.15" valueMax="33.9" valueMin="28.4">
|
||
28.4
|
||
<emphasis id="B91AEA87FF846A5FFC6CCA34FC3CFBE5" box="[946,959,1078,1102]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
33.9 mm
|
||
</quantity>
|
||
,
|
||
<emphasis id="B91AEA87FF846A5FFBF3CA34FBBDFBE5" box="[1069,1086,1078,1102]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 22), typically larger tympanum diameter relative to eye diameter ratio, TYD/ED (mean±SD) 60.1 ± 2.8%, range 55.7
|
||
<emphasis id="B91AEA87FF846A5FFE7DCA79FE33FB38" box="[419,432,1147,1171]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
63.6%,
|
||
<emphasis id="B91AEA87FF846A5FFDDECA79FD92FB38" box="[512,529,1147,1171]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 6 (vs. TYD/
|
||
<specimenCode id="DBC89EEEFF846A5FFD79CA79FD79FB38" box="[679,762,1147,1171]" collectionCode="ED" pageId="29" pageNumber="147">ED 50.5</specimenCode>
|
||
± 3.9%, range 42.9
|
||
<emphasis id="B91AEA87FF846A5FFC11CA79FC5FFB38" box="[975,988,1147,1171]" italics="true" pageId="29" pageNumber="147">–</emphasis>
|
||
58.5%,
|
||
<emphasis id="B91AEA87FF846A5FFBF2CA79FBBEFB38" box="[1068,1085,1147,1171]" italics="true" pageId="29" pageNumber="147">N</emphasis>
|
||
= 26), tympanum oval (vs. circular).
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF846A5FFF1ECAC3FE44FB50" blockId="29.[160,1156,142,1275]" pageId="29" pageNumber="147">For a comparison with the two new species described below, see the comparison sections for those species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374651EFF846A5FFF7ECB2FFDADF9A0" pageId="29" pageNumber="147" type="etymology">
|
||
<paragraph id="8BD13695FF846A5FFF7ECB2FFCB6FA26" blockId="29.[160,1155,1325,1421]" pageId="29" pageNumber="147">
|
||
<emphasis id="B91AEA87FF846A5FFF7ECB2FFEA0FAEC" bold="true" box="[160,291,1325,1351]" italics="true" pageId="29" pageNumber="147">Etymology.</emphasis>
|
||
The specific epithet
|
||
<emphasis id="B91AEA87FF846A5FFDCACB2DFDDEFAEC" box="[532,605,1327,1351]" italics="true" pageId="29" pageNumber="147">‘awuh’</emphasis>
|
||
is treated as a noun in apposition taken from the Pochury language, meaning
|
||
<emphasis id="B91AEA87FF846A5FFE3DCB50FE6BFAC1" box="[483,488,1362,1386]" italics="true" pageId="29" pageNumber="147">‘</emphasis>
|
||
frog
|
||
<emphasis id="B91AEA87FF846A5FFDC8CB50FD98FAC1" box="[534,539,1362,1386]" italics="true" pageId="29" pageNumber="147">’</emphasis>
|
||
. The Pochury tribe are the native inhabitants of Meluri circle in
|
||
<collectingRegion id="49AAF877FF846A5FFF20CB77FE29FA26" box="[254,426,1397,1421]" country="India" name="Nagaland" pageId="29" pageNumber="147">Nagaland state</collectingRegion>
|
||
, within which the
|
||
<typeStatus id="54D58837FF846A5FFDA4CB77FD2FFA26" box="[634,684,1397,1421]" pageId="29" pageNumber="147">type</typeStatus>
|
||
locality lies.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF846A5FFF7ECBCDFDADF9A0" blockId="29.[160,1155,1487,1547]" pageId="29" pageNumber="147">
|
||
<emphasis id="B91AEA87FF846A5FFF7ECBCDFE5FFA42" bold="true" box="[160,476,1487,1513]" italics="true" pageId="29" pageNumber="147">Suggested common name.</emphasis>
|
||
We suggest
|
||
<emphasis id="B91AEA87FF846A5FFDA8CBD2FDFFFA43" box="[630,636,1488,1512]" italics="true" pageId="29" pageNumber="147">‘</emphasis>
|
||
Naga Hills Horned Frog
|
||
<emphasis id="B91AEA87FF846A5FFC5DCBD2FC0AFA43" box="[899,905,1488,1512]" italics="true" pageId="29" pageNumber="147">’</emphasis>
|
||
for this species based on its currently known distribution.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374651EFF846A5CFF7EC83FFB92FCF7" lastPageId="30" lastPageNumber="148" pageId="29" pageNumber="147" type="distribution">
|
||
<paragraph id="8BD13695FF846A5CFF7EC83FFE74FF47" blockId="29.[160,1156,1597,1727]" lastBlockId="30.[160,1156,142,236]" lastPageId="30" lastPageNumber="148" pageId="29" pageNumber="147">
|
||
<emphasis id="B91AEA87FF846A5FFF7EC83FFEB0F9FC" bold="true" box="[160,307,1597,1623]" italics="true" pageId="29" pageNumber="147">Distribution.</emphasis>
|
||
Known only from the vicinity of Kohima and Star Lake in southern and eastern Nagaland state, respectively, at elevations between 1650 and
|
||
<quantity id="4C969B70FF846A5FFC70C860FB80F9D1" box="[942,1027,1634,1658]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="29" pageNumber="147" unit="m" value="2220.0">2220 m</quantity>
|
||
asl. (
|
||
<figureCitation id="13552A10FF846A5FFBE0C860FF2EF936" captionStart="Figure 2" captionStartId="11.[160,225,1528,1551]" captionTargetBox="[165,1152,1053,1494]" captionTargetId="figure-391@11.[163,1153,1050,1497]" captionTargetPageId="11" captionText="Figure 2. Topographic distribution map of the Megophrys (X.) megacephala SG taxa: Left. All taxa from Northeast India and surrounding countries: diamonds represent type localities; circles represent referred localities based on examined specimens; circles with central dot represent identifications based on literature where sufficient evidence was available (see Remarks sections); Right. Inset rectangle enlarged to show the distributions of species from more densely sampled southern Nagaland and northern Manipur states, Northeast India. Blue lines represent major rivers; grey lines represent international borders." figureDoi="http://doi.org/10.5281/zenodo.4608903" httpUri="https://zenodo.org/record/4608903/files/figure.png" pageId="29" pageNumber="147">Figure 2</figureCitation>
|
||
). Considering the wide elevational range inhabited, the geographic distribution of this species is expected to be more extensive than currently reported. The proximity of the collection localities relative to political borders indicates that
|
||
<taxonomicName id="4C6E4D16FF876A5CFCE0CE8CFC74FF0D" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[830,1015,142,166]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="30" pageNumber="148" phylum="Chordata" rank="species" species="awuh" status="sp. nov.">
|
||
<emphasis id="B91AEA87FF876A5CFCE0CE8CFC74FF0D" box="[830,1015,142,166]" italics="true" pageId="30" pageNumber="148">Megophrys awuh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF876A5CFC23CE8CFBD4FF0D" box="[1021,1111,142,166]" pageId="30" pageNumber="148" rank="species">
|
||
<emphasis id="B91AEA87FF876A5CFC23CE8CFBD2FF0D" bold="true" box="[1021,1105,142,166]" pageId="30" pageNumber="148">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
will most likely be found in neighbouring Manipur state (NEI) and
|
||
<collectingRegion id="49AAF877FF876A5CFCB5CEB3FB99FF62" box="[875,1050,177,201]" country="Myanmar" name="Sagaing" pageId="30" pageNumber="148">Sagaing Region</collectingRegion>
|
||
(western
|
||
<collectingCountry id="F3797605FF876A5CFF7ECED6FE8AFF47" box="[160,265,212,236]" name="Myanmar" pageId="30" pageNumber="148">Myanmar</collectingCountry>
|
||
) at similar elevations.
|
||
</paragraph>
|
||
<paragraph id="8BD13695FF876A5CFF7ECF15FB92FCF7" blockId="30.[160,1156,279,860]" pageId="30" pageNumber="148">
|
||
<emphasis id="B91AEA87FF876A5CFF7ECF15FE74FE9A" bold="true" box="[160,503,279,305]" italics="true" pageId="30" pageNumber="148">Habitat and natural history.</emphasis>
|
||
All males were calling when collected, and both females contained mature ova indicating that the breeding season includes at least the mid monsoon months of June and July. The type specimens were collected from a primary forest with lush undergrowth, patches of closed canopy, hill streams, and where there were not any significant signs of human activities except perhaps occasional and regulated hunting by the local Pochurys. All specimens were collected during daylight hours (12:00
|
||
<emphasis id="B91AEA87FF876A5CFC5DCFC4FC13FE75" box="[899,912,454,478]" italics="true" pageId="30" pageNumber="148">–</emphasis>
|
||
16:00 h). The
|
||
<typeStatus id="54D58837FF876A5CFBFDCFC4FB07FE75" box="[1059,1156,454,478]" pageId="30" pageNumber="148" type="holotype">holotype</typeStatus>
|
||
was found calling from within grasses on the edge of a spring about a metre deep and three metres diameter; the male
|
||
<typeStatus id="54D58837FF876A5CFE12CC0EFDAFFD8F" box="[460,556,524,548]" pageId="30" pageNumber="148" type="paratype">paratype</typeStatus>
|
||
was found calling beneath grasses half submerged in a puddle in the forest. The female
|
||
<typeStatus id="54D58837FF876A5CFDD3CC2CFDEEFDED" box="[525,621,558,582]" pageId="30" pageNumber="148" type="paratype">paratype</typeStatus>
|
||
was collected from the edge of the same pond as the
|
||
<typeStatus id="54D58837FF876A5CFF13CC53FEB3FDC2" box="[205,304,593,617]" pageId="30" pageNumber="148" type="holotype">holotype</typeStatus>
|
||
. The referred specimen SDBDU 2007.111 was collected from ground level in secondary forest adjacent to highway NH 39 on a night with light rain, about
|
||
<quantity id="4C969B70FF876A5CFC08CC76FB8FFD27" box="[982,1036,628,652]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="148" unit="m" value="10.0">10 m</quantity>
|
||
away from a seasonal stream. SDBDU 2007.079 and SDBDU 2007.080 were collected at ~20:00 h from vegetation within
|
||
<quantity id="4C969B70FF876A5CFEB4CCBBFE27FD7A" box="[362,420,697,721]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="148" unit="m" value="10.0">10 m</quantity>
|
||
of the river Dzütsürü in a mature secondary forest. The Aradurah Forest tissue sample was collected from a juvenile (SVL ~
|
||
<quantity id="4C969B70FF876A5CFCD0CCDEFCDFFD5F" box="[782,860,732,756]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="148" unit="mm" value="12.0">12 mm</quantity>
|
||
) caught on a rainy night at ~19:30 h on the ground under a bush. The bush was situated about four metres away from a seasonal hill stream. A few males were heard calling but possibly due to the moderately heavy rain the calls were not prolonged enough for the individuals to be located.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374651EFF876A5CFF7ECD85FD4DFBE5" pageId="30" pageNumber="148" type="discussion">
|
||
<paragraph id="8BD13695FF876A5CFF7ECD85FD4DFBE5" blockId="30.[160,1156,903,1102]" pageId="30" pageNumber="148">
|
||
<emphasis id="B91AEA87FF876A5CFF7ECD85FE8FFC0A" bold="true" box="[160,268,903,929]" italics="true" pageId="30" pageNumber="148">Remarks.</emphasis>
|
||
Molecular phylogenetic analyses consistently resolved
|
||
<taxonomicName id="4C6E4D16FF876A5CFC49CD8BFBD5FC0A" authority="Mahony & Kamei & Teeling & Biju, 2020" authorityName="Mahony & Kamei & Teeling & Biju" authorityYear="2020" box="[919,1110,905,929]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="30" pageNumber="148" phylum="Chordata" rank="species" species="awuh" status="sp. nov.">
|
||
<emphasis id="B91AEA87FF876A5CFC49CD8BFBD5FC0A" box="[919,1110,905,929]" italics="true" pageId="30" pageNumber="148">Megophrys awuh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FCFF876A5CFBBFCD8BFF50FC6F" pageId="30" pageNumber="148" rank="species">
|
||
<emphasis id="B91AEA87FF876A5CFBBFCD8BFF4EFC6F" bold="true" pageId="30" pageNumber="148">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(adult male SVL
|
||
<quantity id="4C969B70FF876A5CFE4CCDAEFDB1FC6F" box="[402,562,940,964]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" metricValueMax="4.11" metricValueMin="3.57" pageId="30" pageNumber="148" unit="mm" value="38.4" valueMax="41.1" valueMin="35.7">
|
||
35.7
|
||
<emphasis id="B91AEA87FF876A5CFE1FCDAEFE4DFC6F" box="[449,462,940,964]" italics="true" pageId="30" pageNumber="148">–</emphasis>
|
||
41.1 mm
|
||
</quantity>
|
||
) as the sister taxon of the smaller
|
||
<taxonomicName id="4C6E4D16FF876A5CFC67CDAEFB00FC6F" baseAuthorityName="Mathew & Sen" baseAuthorityYear="2007" box="[953,1155,940,964]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="30" pageNumber="148" phylum="Chordata" rank="species" species="zunhebotoensis">
|
||
<emphasis id="B91AEA87FF876A5CFC67CDAEFB00FC6F" box="[953,1155,940,964]" italics="true" pageId="30" pageNumber="148">M. zunhebotoensis</emphasis>
|
||
</taxonomicName>
|
||
(male SVL
|
||
<quantity id="4C969B70FF876A5CFEC7CDCDFE3AFC4C" box="[281,441,974,999]" metricMagnitude="-2" metricUnit="m" metricValue="3.1149999999999998" metricValueMax="3.3899999999999997" metricValueMin="2.84" pageId="30" pageNumber="148" unit="mm" value="31.15" valueMax="33.9" valueMin="28.4">
|
||
28.4
|
||
<emphasis id="B91AEA87FF876A5CFE99CDCCFED7FC4D" box="[327,340,974,998]" italics="true" pageId="30" pageNumber="148">–</emphasis>
|
||
33.9 mm
|
||
</quantity>
|
||
) in all analyses that included mtDNA (
|
||
<figureCitation id="13552A10FF876A5CFCABCDCDFB9FFC4C" box="[885,1052,975,999]" captionStart="Figure 1" captionStartId="10.[160,225,1616,1639]" captionTargetBox="[259,1053,625,1575]" captionTargetId="figure-124@10.[252,1060,616,1633]" captionTargetPageId="10" captionText="Figure 1. ML phylogenetic tree of the subgenus Megophrys (Xenophrys) based on the concatenated mt +nuDNA alignment. Bootstrap values ≥95% are represented by a black spot; 80–95% represented by a grey spot; <80% not given. Numbers next to species names correspond with specimen numbers in Table 1." figureDoi="http://doi.org/10.5281/zenodo.4608900" httpUri="https://zenodo.org/record/4608900/files/figure.png" pageId="30" pageNumber="148">Figures 1 & S1</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFFF4B64FF876A5CFBF6CDCDFE91FBA2" author="Mahony S & Foley NC & Biju SD & Teeling EC" pageId="30" pageNumber="148" pagination="744 - 771" refId="ref46521" refString="Mahony S, Foley NC, Biju SD, Teeling EC. 2017. Evolutionary history of the Asian Horned Frogs (Megophryinae): integrative approaches to timetree dating in the absence of a fossil record. Mol Biol Evol. 34 (4): 744 - 771 + S. I. 33. doi: 10.1093 / molbev / msw 267." type="journal article" year="2017">Mahony et al. 2017</bibRefCitation>
|
||
, figs. 1 & 3 [as
|
||
<emphasis id="B91AEA87FF876A5CFE6CCDF3FE4EFBA2" box="[434,461,1009,1033]" italics="true" pageId="30" pageNumber="148">‘M</emphasis>
|
||
. sp. [2 & 3]
|
||
<emphasis id="B91AEA87FF876A5CFD9ACDF3FDCAFBA2" box="[580,585,1009,1033]" italics="true" pageId="30" pageNumber="148">’</emphasis>
|
||
];
|
||
<bibRefCitation id="EFFF4B64FF876A5CFD83CDF3FCB3FBA2" author="Mahony S & Kamei RG & Teeling EC & Biju SD" box="[605,816,1009,1033]" pageId="30" pageNumber="148" pagination="1 - 96" refId="ref46587" refString="Mahony S, Kamei RG, Teeling EC, Biju SD. 2018. Cryptic diversity within the Megophrys major species group (Amphibia: Megophryidae) of the Asian horned frogs: phylogenetic perspectives and a taxonomic revision of South Asian taxa, with descriptions of four new species. Zootaxa. 4523 (1): 1 - 96. doi: 10.11646 / zootaxa. 4523.1.1." type="journal article" year="2018">Mahony et al. 2018</bibRefCitation>
|
||
, fig. 4 [as
|
||
<emphasis id="B91AEA87FF876A5CFC42CDF3FC35FBA2" box="[924,950,1009,1033]" italics="true" pageId="30" pageNumber="148">‘M</emphasis>
|
||
. sp. 5
|
||
<emphasis id="B91AEA87FF876A5CFC2BCDF3FC79FBA2" box="[1013,1018,1009,1033]" italics="true" pageId="30" pageNumber="148">’</emphasis>
|
||
]). Molecular dating analyses suggested that these sister taxa diverged from their most recent common ancestor at ca. 3.8 MY (
|
||
<bibRefCitation id="EFFF4B64FF876A5CFE7ACA34FDFEFBE5" author="Mahony S & Foley NC & Biju SD & Teeling EC" box="[420,637,1078,1102]" pageId="30" pageNumber="148" pagination="744 - 771" refId="ref46521" refString="Mahony S, Foley NC, Biju SD, Teeling EC. 2017. Evolutionary history of the Asian Horned Frogs (Megophryinae): integrative approaches to timetree dating in the absence of a fossil record. Mol Biol Evol. 34 (4): 744 - 771 + S. I. 33. doi: 10.1093 / molbev / msw 267." type="journal article" year="2017">Mahony et al. 2017</bibRefCitation>
|
||
, fig. 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |