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<document id="30A2C3278D01DC70103CC16F268B49AE" ID-DOI="10.11646/zootaxa.3990.2.5" ID-GBIF-Dataset="b2824eb9-5792-47f2-b7be-7b7b326685e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="239690" ID-ZooBank="35D06588-6A54-46BF-BBD8-B93C063E86FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268048733" checkinUser="plazi" docAuthor="Yang, Jian-Huan, Rao, Ding-Qi &amp; Wang, Ying-Yong" docDate="2015" docId="03A89849F37FFFBDFF2BFB70F2B4064A" docLanguage="en" docName="zt03990p258.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liuixalus feii Yang, Rao &amp; Wang, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="255" masterDocId="FF91E031F37BFFB5FFBCFFBAF2460528" masterDocTitle="A new species of the genus Liuixalus (Anura: Rhacophoridae) from southern China" masterLastPageNumber="258" masterPageNumber="247" pageNumber="251" updateTime="1698610944727" updateUser="plazi">
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<mods:title id="DC4F938435775D6C440C29D1BBC4206F">A new species of the genus Liuixalus (Anura: Rhacophoridae) from southern China</mods:title>
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<mods:namePart id="6A751C2AB7F5AD48FE82BFC939B8652D">Yang, Jian-Huan</mods:namePart>
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<mods:namePart id="D989B2A13EF35D83AB54AC6E93816576">Rao, Ding-Qi</mods:namePart>
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<mods:namePart id="1D80318FC0E936FA824BC18AD464E4CE">Wang, Ying-Yong</mods:namePart>
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<mods:date id="EFF1DBCE3FE8AD8C92AF518C32DA0793">2015</mods:date>
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<treatment id="03A89849F37FFFBDFF2BFB70F2B4064A" ID-DOI="http://doi.org/10.5281/zenodo.5612867" ID-GBIF-Taxon="119638421" ID-Zenodo-Dep="5612867" LSID="urn:lsid:plazi:treatment:03A89849F37FFFBDFF2BFB70F2B4064A" httpUri="http://treatment.plazi.org/id/03A89849F37FFFBDFF2BFB70F2B4064A" lastPageId="8" lastPageNumber="255" pageId="4" pageNumber="251">
<subSubSection id="C31B7AD4F37FFFB1FF2BFB70F3D801CD" box="[151,414,1226,1253]" pageId="4" pageNumber="251" type="nomenclature">
<paragraph id="8BBE295FF37FFFB1FF2BFB70F3D801CD" blockId="4.[151,414,1226,1253]" box="[151,414,1226,1253]" pageId="4" pageNumber="251">
<heading id="D0F69E33F37FFFB1FF2BFB70F3D801CD" bold="true" box="[151,414,1226,1253]" fontSize="11" level="1" pageId="4" pageNumber="251" reason="1">
<emphasis id="B975F54DF37FFFB1FF2BFB70F3D801CD" bold="true" box="[151,414,1226,1253]" pageId="4" pageNumber="251">
<taxonomicName id="4C0152DCF37FFFB1FF2BFB70F37F01CC" ID-CoL="3VNX5" box="[151,313,1226,1252]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="251" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF37FFFB1FF2BFB70F37F01CC" bold="true" box="[151,313,1226,1252]" italics="true" pageId="4" pageNumber="251">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F37FFFB1FEFCFB71F3D801CD" box="[320,414,1227,1253]" pageId="4" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31B7AD4F37FFFB1FF2BFAABF1F5032A" pageId="4" pageNumber="251" type="materials_examined">
<paragraph id="8BBE295FF37FFFB1FF2BFAABF37E005A" blockId="4.[151,1437,1297,2006]" pageId="4" pageNumber="251">
<emphasis id="B975F54DF37FFFB1FF2BFAABF34D0002" bold="true" box="[151,267,1297,1322]" pageId="4" pageNumber="251">
<typeStatus id="54BA97FDF37FFFB1FF2BFAABF3400002" box="[151,262,1297,1322]" pageId="4" pageNumber="251" type="holotype">Holotype</typeStatus>
.
</emphasis>
SYS a002389, adult male, from Heishiding Nature Reserve, Fengkai County, Guangdong Province,
<collectingCountry id="F31669CFF37FFFB1FF2BFA8FF29C0066" box="[151,218,1333,1358]" name="China" pageId="4" pageNumber="251">China</collectingCountry>
(
<geoCoordinate id="EE354F98F37FFFB1FF57FA8CF3C80065" box="[235,398,1334,1358]" direction="north" orientation="latitude" pageId="4" pageNumber="251" precision="1" value="23.45289">23°27'10.4'' N</geoCoordinate>
,
<geoCoordinate id="EE354F98F37FFFB1FE27FA8FF0000065" box="[411,582,1333,1358]" direction="east" orientation="longitude" pageId="4" pageNumber="251" precision="1" value="111.88761">111°53'15.4'' E</geoCoordinate>
,
<quantity id="4CF984BAF37FFFB1FDEFFA8FF0DA0065" box="[595,668,1333,1358]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="251" unit="m" value="550.0">550 m</quantity>
a.s.l.;
<figureCitation id="133A35DAF37FFFB1FD54FA8FF16B0066" box="[744,813,1333,1358]" captionStart="FIGURE 2" captionStartId="4.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@4.[151,1436,193,1049]" captionTargetPageId="4" captionText="FIGURE 2. Map showing the distribution of of Liuixalus species and populations included for phylogenetic analyses in this study. 1: Wuzhishan NR, Hainan; 2: Diaoluoshan NR, Hainan; 3: Cat Ba Island, Vietnam; 4: Mt. Shiwan, Guangxi; 5: Heishiding NR, Guangdong; 6: Hong Kong." httpUri="https://zenodo.org/record/239692/files/figure.png" pageId="4" pageNumber="251">Fig. 2</figureCitation>
), collected by Yu-Long Li and Jian Zhao at night on
<date id="FFBF0F9FF37FFFB1FA32FA8FF372005A" pageId="4" pageNumber="251" value="2013-10-06">6 October 2013</date>
.
</paragraph>
<paragraph id="8BBE295FF37FFFB1FF7BFAC7F1F5032A" blockId="4.[151,1437,1297,2006]" pageId="4" pageNumber="251">
<emphasis id="B975F54DF37FFFB1FF7BFAC7F30000BE" bold="true" box="[199,326,1405,1430]" pageId="4" pageNumber="251">
<typeStatus id="54BA97FDF37FFFB1FF7BFAC7F30700BE" box="[199,321,1405,1430]" pageId="4" pageNumber="251" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Four adult males and two adult females from the same locality as
<typeStatus id="54BA97FDF37FFFB1FBF9FAC7F6ED00BE" box="[1093,1195,1405,1430]" pageId="4" pageNumber="251" type="holotype">holotype</typeStatus>
: SYS a000454, adult female, collected by Jian-Huan Yang on
<date id="FFBF0F9FF37FFFB1FDD4FA18F1420092" box="[616,772,1442,1466]" pageId="4" pageNumber="251" value="2009-05-29">29 May 2009</date>
; SYS a002158, adult male, collected by YLL and JZ on
<date id="FFBF0F9FF37FFFB1FA32FA18F34A00F6" pageId="4" pageNumber="251" value="2013-06-01">1 June 2013</date>
; SYS a002388, adult female, and a002390, adult male, same data as
<typeStatus id="54BA97FDF37FFFB1FBBBFA7FF62B00F6" box="[1031,1133,1477,1502]" pageId="4" pageNumber="251" type="holotype">holotype</typeStatus>
; SYS a0030493050, adult males, collected by Ying-Yong Wang and Zu-Yao Liu on
<date id="FFBF0F9FF37FFFB1FCA7FA53F1E9032A" box="[795,943,1513,1538]" pageId="4" pageNumber="251" value="2014-07-15">15 July 2014</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B7AD4F37FFFB1FF7BF9B7F65703BA" pageId="4" pageNumber="251" type="etymology">
<paragraph id="8BBE295FF37FFFB1FF7BF9B7F65703BA" blockId="4.[151,1437,1297,2006]" pageId="4" pageNumber="251">
<emphasis id="B975F54DF37FFFB1FF7BF9B7F308030E" bold="true" box="[199,334,1549,1574]" pageId="4" pageNumber="251">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="4C0152DCF37FFFB1FDF9F9B4F02F030E" box="[581,617,1550,1574]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="251" phylum="Chordata" rank="species" species="feii">
<emphasis id="B975F54DF37FFFB1FDF9F9B4F02F030E" box="[581,617,1550,1574]" italics="true" pageId="4" pageNumber="251">feii</emphasis>
</taxonomicName>
” is a patronymic noun in the genitive singular; derived from the name of Professor Liang Fei of the Chengdu Institute of Biology, Chinese Academy of Sciences,
<collectingCountry id="F31669CFF37FFFB1FB30F98BF6970362" box="[1164,1233,1585,1610]" name="China" pageId="4" pageNumber="251">China</collectingCountry>
, in recognition of his long-term great contribution to the advancement of amphibian research in
<collectingCountry id="F31669CFF37FFFB1FBAAF9EFF61B0346" box="[1046,1117,1621,1646]" name="China" pageId="4" pageNumber="251">China</collectingCountry>
. For the common name, we suggest “Feis Small Tree Frog” in English, “Fei Shi Xiao Shu Wa” in Chinese.
</paragraph>
</subSubSection>
<subSubSection id="C31B7AD4F37FFFB0FF7BF927F00B05FD" lastPageId="5" lastPageNumber="252" pageId="4" pageNumber="251" type="diagnosis">
<paragraph id="8BBE295FF37FFFB0FF7BF927F00B05FD" blockId="4.[151,1437,1297,2006]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="252" pageId="4" pageNumber="251">
<emphasis id="B975F54DF37FFFB1FF7BF927F307039E" bold="true" box="[199,321,1693,1718]" pageId="4" pageNumber="251">Diagnosis.</emphasis>
The new species is assigned to the genus
<taxonomicName id="4C0152DCF37FFFB1FC97F927F1D2039D" box="[811,916,1693,1717]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="251" phylum="Chordata" rank="genus">
<emphasis id="B975F54DF37FFFB1FC97F927F1D2039D" box="[811,916,1693,1717]" italics="true" pageId="4" pageNumber="251">Liuixalus</emphasis>
</taxonomicName>
by presenting the following characters: body size small; tympanum well distinct and about half the eye diameter; fingers without webbing; toes weakly webbed; hindlimb relatively long, tibio-tarsal articulation reaching the naris or loreal; discs on the digits well developed, discs of toes slightly smaller than disks of fingers; dorsum of body with a X-shaped marking (
<bibRefCitation id="EF9054AEF37FFFB1FB75F8B3F7D7020A" author="Milto" box="[1225,1425,1801,1826]" pageId="4" pageNumber="251" refString="Milto, K. D., Poyarkov, N. A., Orlov, N. L. &amp; Nguyen, T. T. (2013) Two New Rhacophorid Frogs from Cat Ba Island, Gulf of Tonkin (Hai Phong Province, Vietnam). Russian Journal of Herpetology, 20 (4), 287 - 300." type="journal article" year="2013">
Milto
<emphasis id="B975F54DF37FFFB1FAAEF8B0F702020A" box="[1298,1348,1802,1826]" italics="true" pageId="4" pageNumber="251">et al</emphasis>
. 2013
</bibRefCitation>
).
<taxonomicName id="4C0152DCF37FFFB1FF2BF897F36B026E" box="[151,301,1837,1862]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="251" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF37FFFB1FF2BF897F36B026E" box="[151,301,1837,1862]" italics="true" pageId="4" pageNumber="251">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F37FFFB1FE8BF897F3D2026E" box="[311,404,1837,1862]" pageId="4" pageNumber="251" rank="species">
<emphasis id="B975F54DF37FFFB1FE8BF897F3D2026E" bold="true" box="[311,404,1837,1862]" pageId="4" pageNumber="251">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from its congeners by a combination of following characters: small size (SVL
<quantity id="4CF984BAF37FFFB1FF5DF8EBF3CF0241" box="[225,393,1873,1898]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" metricValueMax="1.76" metricValueMin="1.6199999999999999" pageId="4" pageNumber="251" unit="mm" value="16.9" valueMax="17.6" valueMin="16.2">16.217.6 mm</quantity>
in adult males and 18.0
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in adult females); snout obtusely pointed; tympanum distinct, about half size of eye diameter; nostril closer to eye than to the tip of snout; fingers free of webbing; toe III longer than toe V; toes weakly webbed; tibiotarsal articulation reaching the naris or loreal; dorsal skin smooth and scattered with fine granulars; a subtle longitudinal median ridge present on dorsum; weak skin folds present on dorsal surface of body and thighs; supratympanic fold distinct and curved; ventral surface dull white with more or less irregular dark spots; iris bicolored.
</paragraph>
</subSubSection>
<caption id="DF7E79D7F37EFFB0FF2BF911F1DD03C8" httpUri="https://zenodo.org/record/239693/files/figure.png" pageId="5" pageNumber="252" targetBox="[151,1436,265,1685]" targetPageId="5">
<paragraph id="8BBE295FF37EFFB0FF2BF911F1DD03C8" blockId="5.[151,1435,1707,1760]" pageId="5" pageNumber="252">
<emphasis id="B975F54DF37EFFB0FF2BF911F35503E8" bold="true" box="[151,275,1707,1729]" pageId="5" pageNumber="252">FIGURE 3.</emphasis>
Holotype SYS a002389 of
<taxonomicName id="4C0152DCF37EFFB0FD92F911F0F203E9" box="[558,692,1707,1729]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="252" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF37EFFB0FD92F911F0F203E9" box="[558,692,1707,1729]" italics="true" pageId="5" pageNumber="252">Liuixalus feii</emphasis>
</taxonomicName>
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<emphasis id="B975F54DF37EFFB0FD07F916F14D03E9" bold="true" box="[699,779,1708,1729]" pageId="5" pageNumber="252">sp. nov.</emphasis>
</taxonomicNameLabel>
in preservative. A: dorsal view; B: ventral view; C: lateral view of head and body; D: ventral view of right hand; E: ventral view of left foot.
</paragraph>
</caption>
<subSubSection id="C31B7AD4F37EFFB2FF7BF8BDF04205DF" lastPageId="7" lastPageNumber="254" pageId="5" pageNumber="252" type="description">
<paragraph id="8BBE295FF37EFFB3FF7BF8BDF71505D0" blockId="5.[151,1436,1799,2004]" lastBlockId="6.[151,1437,151,789]" lastPageId="6" lastPageNumber="253" pageId="5" pageNumber="252">
<emphasis id="B975F54DF37EFFB0FF7BF8BDF3AD0208" bold="true" box="[199,491,1799,1824]" pageId="5" pageNumber="252">
Description of
<typeStatus id="54BA97FDF37EFFB0FEC1F8BDF3A00208" box="[381,486,1799,1824]" pageId="5" pageNumber="252" type="holotype">holotype</typeStatus>
.
</emphasis>
SYS a002389, adult male, body size small (SVL
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), head length slightly greater than head width (HL
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; HW
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); snout obtusely pointed in dorsal view and profile, projecting beyond margin of the lower jaw; interorbital region flat; canthus rostralis distinct and rounded; loreal region slightly oblique and concave; nostrils oval, slightly protuberant, slightly closer to eye than to tip of snout; interorbital distance slightly greater than internasal distance; pupil horizontal; tympanum distinct, 48.0% of eye diameter, distant from eye by half of its own diameter (ratio of TED/TD 0.5); tympanic rim weakly elevated relative to skin of temporal region; interorbital distance wider than the upper eyelid; pineal ocellus absent; vomerine teeth absent; choanae rounded, at margins of mouth roof; tongue oblong, attached anteriorly and notched posteriorly; supratympanic fold distinct and somewhat curved, extending from behind eye to shoulder basis.
</paragraph>
<paragraph id="8BBE295FF37DFFB3FF7BFEBEF39704F8" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="253">
Forelimbs moderately robust, relative length of fingers I &lt;II &lt;IV &lt;III (
<figureCitation id="133A35DAF37DFFB3FC50FEBEF6690434" box="[1004,1071,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1707,1729]" captionTargetBox="[151,1436,265,1685]" captionTargetId="figure@5.[151,1436,265,1686]" captionTargetPageId="5" captionText="FIGURE 3. Holotype SYS a 002389 of Liuixalus feii sp. nov. in preservative. A: dorsal view; B: ventral view; C: lateral view of head and body; D: ventral view of right hand; E: ventral view of left foot." httpUri="https://zenodo.org/record/239693/files/figure.png" pageId="6" pageNumber="253">Fig. 3</figureCitation>
D); tips of fingers II, III, IV with well-developed disks with distinct circummarginal grooves, disks relatively wide compared to finger width (disk of finger III 216% wider than width of finger III), third finger disk width 68.4% of tympanum diameter; tip of finger I round and without distinct disk; fingers free of webbing; subarticular tubercles prominent, rounded, formula 1, 1, 2, 2; outer palmar tubercle weakly divided into two; accessory palmar tubercles indistinct; no nuptial pads observed on dorsal surface of fingers.
</paragraph>
<paragraph id="8BBE295FF37DFFB3FF7BFE66F33C0780" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="253">
Hindlimbs long and slender, 171% of SVL; tibiotarsal articulation reaching the naris when adpressed along the body; tibia length 54% of SVL; relative length of toes I &lt;II&lt;V &lt;III &lt;IV (
<figureCitation id="133A35DAF37DFFB3FBA6FE45F6260730" box="[1050,1120,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1707,1729]" captionTargetBox="[151,1436,265,1685]" captionTargetId="figure@5.[151,1436,265,1686]" captionTargetPageId="5" captionText="FIGURE 3. Holotype SYS a 002389 of Liuixalus feii sp. nov. in preservative. A: dorsal view; B: ventral view; C: lateral view of head and body; D: ventral view of right hand; E: ventral view of left foot." httpUri="https://zenodo.org/record/239693/files/figure.png" pageId="6" pageNumber="253">Fig. 3</figureCitation>
E); tips of toes with poorly developed disks with distinct circummarginal grooves, disks smaller than those of fingers; toe weakly webbed, webbing formula
<date id="FFBF0F9FF37DFFB3FEE1FDF2F3D10748" box="[349,407,583,608]" pageId="6" pageNumber="253">I 22</date>
½
<date id="FFBF0F9FF37DFFB3FE0EFDF2F3B30748" box="[434,501,583,608]" pageId="6" pageNumber="253">II 23</date>
¼
<date id="FFBF0F9FF37DFFB3FDACFDF2F0060748" box="[528,576,583,608]" pageId="6" pageNumber="253">III 2</date>
4
<date id="FFBF0F9FF37DFFB3FDC4FDF2F0EF0748" box="[632,681,583,608]" pageId="6" pageNumber="253">IV 3</date>
1¾ V; subarticular tubercles distinct, rounded, formula 1, 1, 2, 3, 2; inner metatarsal tubercle low, oval, distinct, approximately
<quantity id="4CF984BAF37DFFB3FCD4FDD6F18407AC" box="[872,962,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="253" unit="mm" value="0.8">0.8 mm</quantity>
in length; outer metatarsal tubercle very small and indistinct.
</paragraph>
<paragraph id="8BBE295FF37DFFB3FF7BFD0EF6AA063D" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="253">Dorsal surface of head and body smooth and scattered with fine granules, those on flanks and eyelids somewhat prominent; a subtle longitudinal median ridge present on dorsum; weak skin folds present on dorsal surface of body and thigh; ventral surface of thighs and abdomen with flat granules, throat smooth.</paragraph>
<caption id="DF7E79D7F37DFFB3FF2BF9BDF60B0313" httpUri="https://zenodo.org/record/239694/files/figure.png" pageId="6" pageNumber="253" targetBox="[151,1436,861,1503]" targetPageId="6">
<paragraph id="8BBE295FF37DFFB3FF2BF9BDF60B0313" blockId="6.[151,1437,1543,1595]" pageId="6" pageNumber="253">
<emphasis id="B975F54DF37DFFB3FF2BF9BDF3520334" bold="true" box="[151,276,1543,1565]" pageId="6" pageNumber="253">FIGURE 4.</emphasis>
Paratypes specimens of
<taxonomicName id="4C0152DCF37DFFB3FDABF9BDF0D80335" box="[535,670,1543,1565]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="253" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF37DFFB3FDABF9BDF0D80335" box="[535,670,1543,1565]" italics="true" pageId="6" pageNumber="253">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F37DFFB3FD1BF9BDF0BF0334" box="[679,761,1543,1564]" pageId="6" pageNumber="253" rank="species">
<emphasis id="B975F54DF37DFFB3FD1BF9BDF0BF0334" bold="true" box="[679,761,1543,1564]" pageId="6" pageNumber="253">sp. nov.</emphasis>
</taxonomicNameLabel>
in preservative, note the variation in dorsal and ventral patterns: SYS a002158, SYS a002388, SYS a002390, SYS a003049, SYS a003050 (from left to right).
</paragraph>
</caption>
<paragraph id="8BBE295FF37DFFB3FF7BF9D9F13F027C" blockId="6.[151,1437,1635,2020]" pageId="6" pageNumber="253">
<emphasis id="B975F54DF37DFFB3FF7BF9D9F0E10354" bold="true" box="[199,679,1635,1660]" pageId="6" pageNumber="253">
Coloration of
<typeStatus id="54BA97FDF37DFFB3FECBF9D9F39B0354" box="[375,477,1635,1660]" pageId="6" pageNumber="253" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dorsal surface pale brown with a dark X-shaped marking from behind the eyes, consisting of two crooked dark lines along body not in contact medially; a dark interorbital narrow bar between the eyes extending to the upper eyelids; dark transverse bars present on dorsal surface of fore and hind limbs: two on the lower arm, four on the thigh, three on the tibia; ventral surface dirty-white, scattered with few irregular dark spots on chest and venter; lateral of head dark brown, supratympanic fold distinctly darker; lower lip pale white with conspicuous dark bars; a remarkable dark blotch present on the anterior portion of shoulder joint; iris bicolored, upper third pale white, lower two third dark brown.
</paragraph>
<paragraph id="8BBE295FF37DFFB2FF7BF8E5F04205DF" blockId="6.[151,1437,1635,2020]" lastBlockId="7.[151,1436,151,248]" lastPageId="7" lastPageNumber="254" pageId="6" pageNumber="253">
<emphasis id="B975F54DF37DFFB3FF7BF8E5F37D0250" bold="true" box="[199,315,1887,1912]" pageId="6" pageNumber="253">Variation.</emphasis>
All six
<typeStatus id="54BA97FDF37DFFB3FE1DF8DAF04A0250" box="[417,524,1888,1912]" pageId="6" pageNumber="253" type="paratype">paratypes</typeStatus>
match the overall characters of the
<typeStatus id="54BA97FDF37DFFB3FC0FF8E5F6530250" box="[947,1045,1887,1912]" pageId="6" pageNumber="253" type="holotype">holotype</typeStatus>
(measurements of
<typeStatus id="54BA97FDF37DFFB3FB48F8DAF7620250" box="[1268,1316,1888,1912]" pageId="6" pageNumber="253">type</typeStatus>
series see
<tableCitation id="C6831CE4F37DFFB3FF2BF839F2AD02B4" box="[151,235,1923,1948]" captionStart="TABLE 3" captionStartId="7.[151,239,292,315]" captionTargetBox="[159,1410,342,1552]" captionTargetPageId="7" captionText="TABLE 3. Measurements (mm) of type specimens of Liuixalus feii sp. nov.. Abbreviations defined in text." httpUri="http://table.plazi.org/id/DF7E79D7F37CFFB2FF2BFE9EF74A0413" pageId="6" pageNumber="253" tableUuid="DF7E79D7F37CFFB2FF2BFE9EF74A0413">Table 3</tableCitation>
). All
<typeStatus id="54BA97FDF37DFFB3FE90F83EF31A02B4" box="[300,348,1924,1948]" pageId="6" pageNumber="253">type</typeStatus>
specimens have a dark X-shaped marking on dorsum composed of two crooked dark lines, these lines are usually separated but are in contact medially in SYS a002158; the dark spots on the ventral surface are somewhat distinct in SYS a002388, 2390, 3049, 3050 (see
<figureCitation id="133A35DAF37DFFB3FCEBF871F18202CC" box="[855,964,1995,2020]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="5.[151,250,1707,1729]" captionStartId-1="6.[151,250,1543,1565]" captionStartId-2="8.[151,250,1850,1872]" captionTargetBox-0="[151,1436,265,1685]" captionTargetBox-1="[151,1436,861,1503]" captionTargetBox-2="[151,1436,943,1806]" captionTargetId-0="figure@5.[151,1436,265,1686]" captionTargetId-1="figure@6.[151,1436,861,1504]" captionTargetId-2="figure@8.[151,1436,943,1806]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="FIGURE 3. Holotype SYS a 002389 of Liuixalus feii sp. nov. in preservative. A: dorsal view; B: ventral view; C: lateral view of head and body; D: ventral view of right hand; E: ventral view of left foot." captionText-1="FIGURE 4. Paratypes specimens of Liuixalus feii sp. nov. in preservative, note the variation in dorsal and ventral patterns: SYS a 002158, SYS a 002388, SYS a 002390, SYS a 003049, SYS a 003050 (from left to right)." captionText-2="FIGURE 5. Liuixalus feii sp. nov. from its type locality, showing the color variation in life. A: SYS a 003049, adult male; B: SYS a 002158, adult male; C: adult female, photographed on 2 May 2010, not collected; D: adult male, photographed on 29 May 2009, not collected." httpUri-0="https://zenodo.org/record/239693/files/figure.png" httpUri-1="https://zenodo.org/record/239694/files/figure.png" httpUri-2="https://zenodo.org/record/239695/files/figure.png" pageId="6" pageNumber="253">Figs. 35</figureCitation>
for coloration variation). No significant differences between males and females were revealed, however females are slightly larger than males (SVL 18.0
<quantity id="4CF984BAF37CFFB2FF2BFF06F34605FC" box="[151,256,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="7" pageNumber="254" unit="mm" value="18.6">18.6 mm</quantity>
versus
<quantity id="4CF984BAF37CFFB2FEE2FF06F04C05FC" box="[350,522,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" metricValueMax="1.76" metricValueMin="1.6199999999999999" pageId="7" pageNumber="254" unit="mm" value="16.9" valueMax="17.6" valueMin="16.2">16.217.6 mm</quantity>
). Of the five males, only SYS a003050 possesses distinct nuptial pad on the posterodorsal surface of finger I.
</paragraph>
</subSubSection>
<caption id="DF7E79D7F37CFFB2FF2BFE9EF74A0413" ID-Table-UUID="DF7E79D7F37CFFB2FF2BFE9EF74A0413" box="[151,1292,291,315]" httpUri="http://table.plazi.org/id/DF7E79D7F37CFFB2FF2BFE9EF74A0413" pageId="7" pageNumber="254" targetBox="[159,1410,342,1552]" targetIsTable="true" targetPageId="7">
<paragraph id="8BBE295FF37CFFB2FF2BFE9EF74A0413" blockId="7.[151,1292,291,315]" box="[151,1292,291,315]" pageId="7" pageNumber="254">
<emphasis id="B975F54DF37CFFB2FF2BFE9EF34F0413" bold="true" box="[151,265,292,315]" pageId="7" pageNumber="254">TABLE 3.</emphasis>
Measurements (mm) of type specimens of
<taxonomicName id="4C0152DCF37CFFB2FD64FE99F1220413" box="[728,868,291,315]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="254" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF37CFFB2FD64FE99F1220413" box="[728,868,291,315]" italics="true" pageId="7" pageNumber="254">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F37CFFB2FCD7FE9EF1860413" box="[875,960,292,315]" pageId="7" pageNumber="254" rank="species">
<emphasis id="B975F54DF37CFFB2FCD7FE9EF1860413" bold="true" box="[875,960,292,315]" pageId="7" pageNumber="254">sp. nov.</emphasis>
</taxonomicNameLabel>
. Abbreviations defined in text.
</paragraph>
</caption>
<paragraph id="8BBE295FF37CFFB2FE97FEECF7330338" pageId="7" pageNumber="254">
<table id="F901DBFFF37C004AFF23FEECF7C40338" box="[159,1410,342,1552]" gridcols="9" gridrows="10" pageId="7" pageNumber="254">
<tr id="35312B1DF37C004AFF23FEECF7C4049C" box="[159,1410,342,436]" gridrow="0" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FEECF3C4049C" box="[159,386,342,436]" gridcol="0" gridrow="0" pageId="7" pageNumber="254">SYS a002158 sex Male</th>
<th id="76E04261F37C004AFE1FFEECF3BD049C" box="[419,507,342,436]" gridcol="1" gridrow="0" pageId="7" pageNumber="254">SYS a002389 Male</th>
<th id="76E04261F37C004AFDA0FEECF032049C" box="[540,628,342,436]" gridcol="2" gridrow="0" pageId="7" pageNumber="254">SYS a002390 Male</th>
<th id="76E04261F37C004AFD29FEECF0AD049C" box="[661,747,342,436]" gridcol="3" gridrow="0" pageId="7" pageNumber="254">SYS a003049 Male</th>
<th id="76E04261F37C004AFCB2FEECF123049C" box="[782,869,342,436]" gridcol="4" gridrow="0" pageId="7" pageNumber="254">SYS a003050 Male</th>
<th id="76E04261F37C004AFC3AFEECF1BD049C" box="[902,1019,342,436]" gridcol="5" gridrow="0" pageId="7" pageNumber="254">Mean ± SD (N=5)</th>
<th id="76E04261F37C004AFBA0FEECF632049C" box="[1052,1140,342,436]" gridcol="6" gridrow="0" pageId="7" pageNumber="254">SYS a000454 Female</th>
<th id="76E04261F37C004AFB29FEECF6AA049C" box="[1173,1260,342,436]" gridcol="7" gridrow="0" pageId="7" pageNumber="254">SYS a002388 Female</th>
<th id="76E04261F37C004AFAB2FEECF7C4049C" box="[1294,1410,342,436]" gridcol="8" gridrow="0" pageId="7" pageNumber="254">Mean ± SD (N=2)</th>
</tr>
<tr id="35312B1DF37C004AFF23FE72F7C40718" box="[159,1410,456,560]" gridrow="1" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FE72F3C40718" box="[159,386,456,560]" gridcol="0" gridrow="1" pageId="7" pageNumber="254">SVL 16.3 HL 6.2 HW 6.1</th>
<td id="76E04261F37C004AFE1FFE72F3BD0718" box="[419,507,456,560]" gridcol="1" gridrow="1" pageId="7" pageNumber="254">16.8 6.6 5.7</td>
<td id="76E04261F37C004AFDA0FE72F0320718" box="[540,628,456,560]" gridcol="2" gridrow="1" pageId="7" pageNumber="254">17.0 6.6 6.0</td>
<td id="76E04261F37C004AFD29FE72F0AD0718" box="[661,747,456,560]" gridcol="3" gridrow="1" pageId="7" pageNumber="254">16.2 6.2 5.6</td>
<td id="76E04261F37C004AFCB2FE72F1230718" box="[782,869,456,560]" gridcol="4" gridrow="1" pageId="7" pageNumber="254">17.6 6.8 6.3</td>
<td id="76E04261F37C004AFC3AFE72F1BD0718" box="[902,1019,456,560]" gridcol="5" gridrow="1" pageId="7" pageNumber="254">16.78±0.57 6.48±0.27 5.94±0.29</td>
<td id="76E04261F37C004AFBA0FE72F6320718" box="[1052,1140,456,560]" gridcol="6" gridrow="1" pageId="7" pageNumber="254">18.7 7.2 6.2</td>
<td id="76E04261F37C004AFB29FE72F6AA0718" box="[1173,1260,456,560]" gridcol="7" gridrow="1" pageId="7" pageNumber="254">18.0 6.7 6.2</td>
<td id="76E04261F37C004AFAB2FE72F7C40718" box="[1294,1410,456,560]" gridcol="8" gridrow="1" pageId="7" pageNumber="254">18.35±0.49 6.95±0.35 6.20±0.00</td>
</tr>
<tr id="35312B1DF37C004AFF23FDFEF7C40784" box="[159,1410,580,684]" gridrow="2" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FDFEF3C40784" box="[159,386,580,684]" gridcol="0" gridrow="2" pageId="7" pageNumber="254">SL 2.4 IND 1.9 IOD 2.3</th>
<td id="76E04261F37C004AFE1FFDFEF3BD0784" box="[419,507,580,684]" gridcol="1" gridrow="2" pageId="7" pageNumber="254">2.7 2.0 2.1</td>
<td id="76E04261F37C004AFDA0FDFEF0320784" box="[540,628,580,684]" gridcol="2" gridrow="2" pageId="7" pageNumber="254">2.6 2.0 2.0</td>
<td id="76E04261F37C004AFD29FDFEF0AD0784" box="[661,747,580,684]" gridcol="3" gridrow="2" pageId="7" pageNumber="254">2.4 2.0 2.1</td>
<td id="76E04261F37C004AFCB2FDFEF1230784" box="[782,869,580,684]" gridcol="4" gridrow="2" pageId="7" pageNumber="254">2.9 2.2 2.4</td>
<td id="76E04261F37C004AFC3AFDFEF1BD0784" box="[902,1019,580,684]" gridcol="5" gridrow="2" pageId="7" pageNumber="254">2.60±0.21 2.02±0.11 2.18±0.16</td>
<td id="76E04261F37C004AFBA0FDFEF6320784" box="[1052,1140,580,684]" gridcol="6" gridrow="2" pageId="7" pageNumber="254">2.9 2.3 2.5</td>
<td id="76E04261F37C004AFB29FDFEF6AA0784" box="[1173,1260,580,684]" gridcol="7" gridrow="2" pageId="7" pageNumber="254">2.7 2.1 2.3</td>
<td id="76E04261F37C004AFAB2FDFEF7C40784" box="[1294,1410,580,684]" gridcol="8" gridrow="2" pageId="7" pageNumber="254">2.80±0.14 2.20±0.14 2.40±0.14</td>
</tr>
<tr id="35312B1DF37C004AFF23FD7AF7C40600" box="[159,1410,704,808]" gridrow="3" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FD7AF3C40600" box="[159,386,704,808]" gridcol="0" gridrow="3" pageId="7" pageNumber="254">UEW 1.2 ED 2.0 TD 0.9</th>
<td id="76E04261F37C004AFE1FFD7AF3BD0600" box="[419,507,704,808]" gridcol="1" gridrow="3" pageId="7" pageNumber="254">1.4 2.1 1.0</td>
<td id="76E04261F37C004AFDA0FD7AF0320600" box="[540,628,704,808]" gridcol="2" gridrow="3" pageId="7" pageNumber="254">1.3 2.2 0.9</td>
<td id="76E04261F37C004AFD29FD7AF0AD0600" box="[661,747,704,808]" gridcol="3" gridrow="3" pageId="7" pageNumber="254">1.1 2.2 0.9</td>
<td id="76E04261F37C004AFCB2FD7AF1230600" box="[782,869,704,808]" gridcol="4" gridrow="3" pageId="7" pageNumber="254">1.1 2.3 1.0</td>
<td id="76E04261F37C004AFC3AFD7AF1BD0600" box="[902,1019,704,808]" gridcol="5" gridrow="3" pageId="7" pageNumber="254">1.22±0.13 2.16±0.11 0.94±0.05</td>
<td id="76E04261F37C004AFBA0FD7AF6320600" box="[1052,1140,704,808]" gridcol="6" gridrow="3" pageId="7" pageNumber="254">1.3 2.5 1.0</td>
<td id="76E04261F37C004AFB29FD7AF6AA0600" box="[1173,1260,704,808]" gridcol="7" gridrow="3" pageId="7" pageNumber="254">1.1 2.2 0.9</td>
<td id="76E04261F37C004AFAB2FD7AF7C40600" box="[1294,1410,704,808]" gridcol="8" gridrow="3" pageId="7" pageNumber="254">1.20±0.14 2.35±0.21 0.95±0.07</td>
</tr>
<tr id="35312B1DF37C004AFF23FC86F7C4068C" box="[159,1410,828,932]" gridrow="4" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FC86F3C4068C" box="[159,386,828,932]" gridcol="0" gridrow="4" pageId="7" pageNumber="254">TED 0.6 LAHL 7.9 HAL 3.9</th>
<td id="76E04261F37C004AFE1FFC86F3BD068C" box="[419,507,828,932]" gridcol="1" gridrow="4" pageId="7" pageNumber="254">0.5 8.0 4.2</td>
<td id="76E04261F37C004AFDA0FC86F032068C" box="[540,628,828,932]" gridcol="2" gridrow="4" pageId="7" pageNumber="254">0.6 7.4 4.3</td>
<td id="76E04261F37C004AFD29FC86F0AD068C" box="[661,747,828,932]" gridcol="3" gridrow="4" pageId="7" pageNumber="254">0.6 7.8 4.1</td>
<td id="76E04261F37C004AFCB2FC86F123068C" box="[782,869,828,932]" gridcol="4" gridrow="4" pageId="7" pageNumber="254">0.6 8.1 4.2</td>
<td id="76E04261F37C004AFC3AFC86F1BD068C" box="[902,1019,828,932]" gridcol="5" gridrow="4" pageId="7" pageNumber="254">0.58±0.04 7.84±0.27 4.14±0.15</td>
<td id="76E04261F37C004AFBA0FC86F632068C" box="[1052,1140,828,932]" gridcol="6" gridrow="4" pageId="7" pageNumber="254">0.7 7.4 3.9</td>
<td id="76E04261F37C004AFB29FC86F6AA068C" box="[1173,1260,828,932]" gridcol="7" gridrow="4" pageId="7" pageNumber="254">0.6 8.1 4.3</td>
<td id="76E04261F37C004AFAB2FC86F7C4068C" box="[1294,1410,828,932]" gridcol="8" gridrow="4" pageId="7" pageNumber="254">0.65±0.07 7.75±0.49 4.10±0.28</td>
</tr>
<tr id="35312B1DF37C004AFF23FC02F7C40108" box="[159,1410,952,1056]" gridrow="5" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FC02F3C40108" box="[159,386,952,1056]" gridcol="0" gridrow="5" pageId="7" pageNumber="254">HLL 26.9 TIB 8.3 LFT 11.9</th>
<td id="76E04261F37C004AFE1FFC02F3BD0108" box="[419,507,952,1056]" gridcol="1" gridrow="5" pageId="7" pageNumber="254">28.7 9.1 12.6</td>
<td id="76E04261F37C004AFDA0FC02F0320108" box="[540,628,952,1056]" gridcol="2" gridrow="5" pageId="7" pageNumber="254">27.0 9.0 12.1</td>
<td id="76E04261F37C004AFD29FC02F0AD0108" box="[661,747,952,1056]" gridcol="3" gridrow="5" pageId="7" pageNumber="254">27.8 8.7 11.9</td>
<td id="76E04261F37C004AFCB2FC02F1230108" box="[782,869,952,1056]" gridcol="4" gridrow="5" pageId="7" pageNumber="254">28.3 9.0 12.0</td>
<td id="76E04261F37C004AFC3AFC02F1BD0108" box="[902,1019,952,1056]" gridcol="5" gridrow="5" pageId="7" pageNumber="254">27.74±0.79 8.82±0.33 12.10±0.29</td>
<td id="76E04261F37C004AFBA0FC02F6320108" box="[1052,1140,952,1056]" gridcol="6" gridrow="5" pageId="7" pageNumber="254">28.7 9.4 12.1</td>
<td id="76E04261F37C004AFB29FC02F6AA0108" box="[1173,1260,952,1056]" gridcol="7" gridrow="5" pageId="7" pageNumber="254">29.9 9.6 13</td>
<td id="76E04261F37C004AFAB2FC02F7C40108" box="[1294,1410,952,1056]" gridcol="8" gridrow="5" pageId="7" pageNumber="254">29.30±0.85 9.50±0.14 12.55±0.64</td>
</tr>
<tr id="35312B1DF37C004AFF23FB8EF7C401B4" box="[159,1410,1076,1180]" gridrow="6" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FB8EF3C401B4" box="[159,386,1076,1180]" gridcol="0" gridrow="6" pageId="7" pageNumber="254">FT 7.3 FDW III 0.5 TDW IV 0.5</th>
<td id="76E04261F37C004AFE1FFB8EF3BD01B4" box="[419,507,1076,1180]" gridcol="1" gridrow="6" pageId="7" pageNumber="254">7.8 0.7 0.5</td>
<td id="76E04261F37C004AFDA0FB8EF03201B4" box="[540,628,1076,1180]" gridcol="2" gridrow="6" pageId="7" pageNumber="254">7.2 0.5 0.5</td>
<td id="76E04261F37C004AFD29FB8EF0AD01B4" box="[661,747,1076,1180]" gridcol="3" gridrow="6" pageId="7" pageNumber="254">7.3 0.5 0.5</td>
<td id="76E04261F37C004AFCB2FB8EF12301B4" box="[782,869,1076,1180]" gridcol="4" gridrow="6" pageId="7" pageNumber="254">7.7 0.7 0.6</td>
<td id="76E04261F37C004AFC3AFB8EF1BD01B4" box="[902,1019,1076,1180]" gridcol="5" gridrow="6" pageId="7" pageNumber="254">7.46±0.27 0.58±0.11 0.52±0.04</td>
<td id="76E04261F37C004AFBA0FB8EF63201B4" box="[1052,1140,1076,1180]" gridcol="6" gridrow="6" pageId="7" pageNumber="254">7.8 0.5 0.5</td>
<td id="76E04261F37C004AFB29FB8EF6AA01B4" box="[1173,1260,1076,1180]" gridcol="7" gridrow="6" pageId="7" pageNumber="254">7.6 0.6 0.5</td>
<td id="76E04261F37C004AFAB2FB8EF7C401B4" box="[1294,1410,1076,1180]" gridcol="8" gridrow="6" pageId="7" pageNumber="254">7.70±0.14 0.55±0.07 0.50±0.00</td>
</tr>
<tr id="35312B1DF37C004AFF23FB0AF7C40030" box="[159,1410,1200,1304]" gridrow="7" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FB0AF3C40030" box="[159,386,1200,1304]" gridcol="0" gridrow="7" pageId="7" pageNumber="254">HL/SVL 0.38 HL/HW 1.02 SL/SVL 0.15</th>
<td id="76E04261F37C004AFE1FFB0AF3BD0030" box="[419,507,1200,1304]" gridcol="1" gridrow="7" pageId="7" pageNumber="254">0.39 1.16 0.16</td>
<td id="76E04261F37C004AFDA0FB0AF0320030" box="[540,628,1200,1304]" gridcol="2" gridrow="7" pageId="7" pageNumber="254">0.39 1.10 0.15</td>
<td id="76E04261F37C004AFD29FB0AF0AD0030" box="[661,747,1200,1304]" gridcol="3" gridrow="7" pageId="7" pageNumber="254">0.38 1.11 0.15</td>
<td id="76E04261F37C004AFCB2FB0AF1230030" box="[782,869,1200,1304]" gridcol="4" gridrow="7" pageId="7" pageNumber="254">0.39 1.08 0.16</td>
<td id="76E04261F37C004AFC3AFB0AF1BD0030" box="[902,1019,1200,1304]" gridcol="5" gridrow="7" pageId="7" pageNumber="254">0.39±0.00 1.09±0.05 0.15±0.01</td>
<td id="76E04261F37C004AFBA0FB0AF6320030" box="[1052,1140,1200,1304]" gridcol="6" gridrow="7" pageId="7" pageNumber="254">0.39 1.16 0.16</td>
<td id="76E04261F37C004AFB29FB0AF6AA0030" box="[1173,1260,1200,1304]" gridcol="7" gridrow="7" pageId="7" pageNumber="254">0.37 1.08 0.15</td>
<td id="76E04261F37C004AFAB2FB0AF7C40030" box="[1294,1410,1200,1304]" gridcol="8" gridrow="7" pageId="7" pageNumber="254">0.38±0.01 1.12±0.06 0.15±0.00</td>
</tr>
<tr id="35312B1DF37C004AFF23FA96F7C400BC" box="[159,1410,1324,1428]" gridrow="8" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FA96F3C400BC" box="[159,386,1324,1428]" gridcol="0" gridrow="8" pageId="7" pageNumber="254">ED/SL 0.83 TD/ED 0.45 TED/TD 0.67</th>
<td id="76E04261F37C004AFE1FFA96F3BD00BC" box="[419,507,1324,1428]" gridcol="1" gridrow="8" pageId="7" pageNumber="254">0.78 0.48 0.50</td>
<td id="76E04261F37C004AFDA0FA96F03200BC" box="[540,628,1324,1428]" gridcol="2" gridrow="8" pageId="7" pageNumber="254">0.85 0.41 0.67</td>
<td id="76E04261F37C004AFD29FA96F0AD00BC" box="[661,747,1324,1428]" gridcol="3" gridrow="8" pageId="7" pageNumber="254">0.92 0.41 0.67</td>
<td id="76E04261F37C004AFCB2FA96F12300BC" box="[782,869,1324,1428]" gridcol="4" gridrow="8" pageId="7" pageNumber="254">0.79 0.43 0.60</td>
<td id="76E04261F37C004AFC3AFA96F1BD00BC" box="[902,1019,1324,1428]" gridcol="5" gridrow="8" pageId="7" pageNumber="254">0.83±0.05 0.44±0.03 0.62±0.07</td>
<td id="76E04261F37C004AFBA0FA96F63200BC" box="[1052,1140,1324,1428]" gridcol="6" gridrow="8" pageId="7" pageNumber="254">0.86 0.40 0.70</td>
<td id="76E04261F37C004AFB29FA96F6AA00BC" box="[1173,1260,1324,1428]" gridcol="7" gridrow="8" pageId="7" pageNumber="254">0.81 0.41 0.67</td>
<td id="76E04261F37C004AFAB2FA96F7C400BC" box="[1294,1410,1324,1428]" gridcol="8" gridrow="8" pageId="7" pageNumber="254">0.84±0.03 0.40±0.01 0.68±0.02</td>
</tr>
<tr id="35312B1DF37C004AFF23FA12F7C40338" box="[159,1410,1448,1552]" gridrow="9" pageId="7" pageNumber="254">
<th id="76E04261F37C004AFF23FA12F3C40338" box="[159,386,1448,1552]" gridcol="0" gridrow="9" pageId="7" pageNumber="254">LAHL/SVL 0.48 HLL/SVL 1.65 TIB/SVL 0.51</th>
<td id="76E04261F37C004AFE1FFA12F3BD0338" box="[419,507,1448,1552]" gridcol="1" gridrow="9" pageId="7" pageNumber="254">0.48 1.71 0.54</td>
<td id="76E04261F37C004AFDA0FA12F0320338" box="[540,628,1448,1552]" gridcol="2" gridrow="9" pageId="7" pageNumber="254">0.44 1.59 0.53</td>
<td id="76E04261F37C004AFD29FA12F0AD0338" box="[661,747,1448,1552]" gridcol="3" gridrow="9" pageId="7" pageNumber="254">0.48 1.72 0.54</td>
<td id="76E04261F37C004AFCB2FA12F1230338" box="[782,869,1448,1552]" gridcol="4" gridrow="9" pageId="7" pageNumber="254">0.46 1.61 0.51</td>
<td id="76E04261F37C004AFC3AFA12F1BD0338" box="[902,1019,1448,1552]" gridcol="5" gridrow="9" pageId="7" pageNumber="254">0.47±0.02 1.65±0.06 0.53±0.01</td>
<td id="76E04261F37C004AFBA0FA12F6320338" box="[1052,1140,1448,1552]" gridcol="6" gridrow="9" pageId="7" pageNumber="254">0.40 1.53 0.50</td>
<td id="76E04261F37C004AFB29FA12F6AA0338" box="[1173,1260,1448,1552]" gridcol="7" gridrow="9" pageId="7" pageNumber="254">0.45 1.66 0.53</td>
<td id="76E04261F37C004AFAB2FA12F7C40338" box="[1294,1410,1448,1552]" gridcol="8" gridrow="9" pageId="7" pageNumber="254">0.42±0.04 1.60±0.09 0.52±0.02</td>
</tr>
</table>
</paragraph>
<subSubSection id="C31B7AD4F37CFFBDFF7BF9EBF2B4064A" lastPageId="8" lastPageNumber="255" pageId="7" pageNumber="254" type="distribution">
<paragraph id="8BBE295FF37CFFB2FF7BF9EBF7760399" blockId="7.[151,1436,1617,2002]" pageId="7" pageNumber="254">
<emphasis id="B975F54DF37CFFB2FF7BF9EBF0230342" bold="true" box="[199,613,1617,1642]" pageId="7" pageNumber="254">Distribution and natural history.</emphasis>
The new species,
<taxonomicName id="4C0152DCF37CFFB2FCEFF9EBF1A80341" box="[851,1006,1617,1641]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="254" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF37CFFB2FCEFF9EBF1A80341" box="[851,1006,1617,1641]" italics="true" pageId="7" pageNumber="254">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F37CFFB2FC41F9EBF6180342" box="[1021,1118,1617,1642]" pageId="7" pageNumber="254" rank="species">
<emphasis id="B975F54DF37CFFB2FC41F9EBF6180342" bold="true" box="[1021,1118,1617,1642]" pageId="7" pageNumber="254">sp. nov.</emphasis>
</taxonomicNameLabel>
, is currently known from Heishiding Nature Reserve, Fengkai County, Guangdong Province,
<collectingCountry id="F31669CFF37CFFB2FC26F9CFF19B03A6" box="[922,989,1653,1678]" name="China" pageId="7" pageNumber="254">China</collectingCountry>
(
<figureCitation id="133A35DAF37CFFB2FC51F9CFF67203A6" box="[1005,1076,1653,1678]" captionStart="FIGURE 2" captionStartId="4.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@4.[151,1436,193,1049]" captionTargetPageId="4" captionText="FIGURE 2. Map showing the distribution of of Liuixalus species and populations included for phylogenetic analyses in this study. 1: Wuzhishan NR, Hainan; 2: Diaoluoshan NR, Hainan; 3: Cat Ba Island, Vietnam; 4: Mt. Shiwan, Guangxi; 5: Heishiding NR, Guangdong; 6: Hong Kong." httpUri="https://zenodo.org/record/239692/files/figure.png" pageId="7" pageNumber="254">Fig. 2</figureCitation>
). All individuals recorded were found on leaf-litter on the forest floor in well-preserved primary forest at elevations between
<quantity id="4CF984BAF37CFFB2FB16F923F76A0399" box="[1194,1324,1689,1714]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="8.0" metricValueMin="3.5" pageId="7" pageNumber="254" unit="m" value="575.0" valueMax="800.0" valueMin="350.0">
350
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</quantity>
.
</paragraph>
<paragraph id="8BBE295FF37CFFB2FF7BF904F1B202FA" blockId="7.[151,1436,1617,2002]" pageId="7" pageNumber="254">
In its
<typeStatus id="54BA97FDF37CFFB2FEBAF904F37003FE" box="[262,310,1726,1750]" pageId="7" pageNumber="254">type</typeStatus>
locality, calls of males of
<taxonomicName id="4C0152DCF37CFFB2FDE0F907F0A903FD" box="[604,751,1725,1749]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="254" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF37CFFB2FDE0F907F0A903FD" box="[604,751,1725,1749]" italics="true" pageId="7" pageNumber="254">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F37CFFB2FD49F907F10803FE" box="[757,846,1725,1750]" pageId="7" pageNumber="254" rank="species">
<emphasis id="B975F54DF37CFFB2FD49F907F10803FE" bold="true" box="[757,846,1725,1750]" pageId="7" pageNumber="254">sp. nov.</emphasis>
</taxonomicNameLabel>
have been heard from April to October. A small tree hole in the roots of a large tree (the hole opening approximately
<quantity id="4CF984BAF37CFFB2FCD7F958F1FC03D1" box="[875,954,1762,1786]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="254" unit="mm" value="28.0">28 mm</quantity>
wide,
<quantity id="4CF984BAF37CFFB2FBB9F95BF61303D1" box="[1029,1109,1761,1786]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="7" pageNumber="254" unit="mm" value="64.0">64 mm</quantity>
high and
<quantity id="4CF984BAF37CFFB2FB78F958F75503D1" box="[1220,1299,1762,1786]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="7" pageNumber="254" unit="mm" value="78.0">78 mm</quantity>
deep; water depth inside the hole about
<quantity id="4CF984BAF37CFFB2FE72F8BCF0670236" box="[462,545,1798,1822]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="254" unit="mm" value="20.0">20 mm</quantity>
),
<quantity id="4CF984BAF37CFFB2FD89F8BCF03C0235" box="[565,634,1798,1822]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="254" unit="cm" value="10.0">10 cm</quantity>
above the ground, was found containing eight free-swimming tadpoles on
<date id="FFBF0F9FF37CFFB2FF05F890F305026A" box="[185,323,1834,1858]" pageId="7" pageNumber="254" value="2010-05-02">2 May 2010</date>
, as well as an adult male calling nearby. We assume these to belong to the new species, but have not confirmed it. All individuals of the new species were found on the forest floor and not closed to streams or pools; and no individual was recorded along the streams, ponds or other kinds of wetlands during our surveys in the nature reserve. Therefore, the new species appears to be a forest-dependent species and is likely a phytotelmbreeder, breeding in water accumulated in tree or bamboo holes in the forest.
</paragraph>
<paragraph id="8BBE295FF373FFBDFF7BFF2DF2B4064A" blockId="8.[151,1436,151,866]" pageId="8" pageNumber="255">
<emphasis id="B975F54DF373FFBDFF7BFF2DF32B0598" bold="true" box="[199,365,151,176]" pageId="8" pageNumber="255">Comparisons.</emphasis>
<taxonomicName id="4C0152DCF373FFBDFECAFF22F04D0598" box="[374,523,152,176]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFECAFF22F04D0598" box="[374,523,152,176]" italics="true" pageId="8" pageNumber="255">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFDA8FF2DF0370598" box="[532,625,151,176]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFDA8FF2DF0370598" bold="true" box="[532,625,151,176]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C0152DCF373FFBDFCB0FF23F1D20598" box="[780,916,152,176]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="calcarius">
<emphasis id="B975F54DF373FFBDFCB0FF23F1D20598" box="[780,916,152,176]" italics="true" pageId="8" pageNumber="255">L. calcarius</emphasis>
</taxonomicName>
by having nostril closer to eye than to tip of snout (reverse condition in
<taxonomicName id="4C0152DCF373FFBDFE71FF07F01205FC" box="[461,596,188,212]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="calcarius">
<emphasis id="B975F54DF373FFBDFE71FF07F01205FC" box="[461,596,188,212]" italics="true" pageId="8" pageNumber="255">L. calcarius</emphasis>
</taxonomicName>
); tibiotarsal articulation reaching the naris (versus beyond the tip of snout in
<taxonomicName id="4C0152DCF373FFBDFF08FF5BF37C05D0" box="[180,314,224,248]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="calcarius">
<emphasis id="B975F54DF373FFBDFF08FF5BF37C05D0" box="[180,314,224,248]" italics="true" pageId="8" pageNumber="255">L. calcarius</emphasis>
</taxonomicName>
); toe III longer than toe V (versus toe III length equal to toe V in
<taxonomicName id="4C0152DCF373FFBDFB9DFF5BF6E105D0" box="[1057,1191,224,248]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="calcarius">
<emphasis id="B975F54DF373FFBDFB9DFF5BF6E105D0" box="[1057,1191,224,248]" italics="true" pageId="8" pageNumber="255">L. calcarius</emphasis>
</taxonomicName>
); throat of males dull white (versus dark colored in males in
<taxonomicName id="4C0152DCF373FFBDFDE9FEBFF09A0434" box="[597,732,260,284]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="calcarius">
<emphasis id="B975F54DF373FFBDFDE9FEBFF09A0434" box="[597,732,260,284]" italics="true" pageId="8" pageNumber="255">L. calcarius</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0152DCF373FFBDFD4AFEBEF1CD0434" box="[758,907,260,284]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFD4AFEBEF1CD0434" box="[758,907,260,284]" italics="true" pageId="8" pageNumber="255">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFC2FFEBEF1A80435" box="[915,1006,260,285]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFC2FFEBEF1A80435" bold="true" box="[915,1006,260,285]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C0152DCF373FFBDFB3BFEBFF74B0434" box="[1159,1293,260,284]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B975F54DF373FFBDFB3BFEBFF74B0434" box="[1159,1293,260,284]" italics="true" pageId="8" pageNumber="255">L. hainanus</emphasis>
</taxonomicName>
by having a relatively smaller tympanum in males (mean ratio of TD/SVL
<quantity id="4CF984BAF373FFBDFCF3FE9DF1EC0468" box="[847,938,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.4224" pageId="8" pageNumber="255" unit="in" value="0.056">0.056 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFC0CFE93F1B70468" box="[944,1009,296,320]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFC0CFE93F1B70468" box="[944,1009,296,320]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFC44FE9DF6170468" box="[1016,1105,295,320]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFC44FE9DF6170468" bold="true" box="[1016,1105,295,320]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFB19FE92F7470468" box="[1189,1281,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.0574" pageId="8" pageNumber="255" unit="in" value="0.081">0.081 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFABBFE93F7CC0468" box="[1287,1418,296,320]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B975F54DF373FFBDFABBFE93F7CC0468" box="[1287,1418,296,320]" italics="true" pageId="8" pageNumber="255">L. hainanus</emphasis>
</taxonomicName>
); relatively shorter hindlimbs in males (mean ratio of HLL/SVL
<quantity id="4CF984BAF373FFBDFCE3FEF6F1EB044D" box="[863,941,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="4.191" pageId="8" pageNumber="255" unit="in" value="1.65">1.65 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFC08FEF7F1B1044C" box="[948,1015,332,356]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFC08FEF7F1B1044C" box="[948,1015,332,356]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFC42FEF6F61F044D" box="[1022,1113,332,357]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFC42FEF6F61F044D" bold="true" box="[1022,1113,332,357]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFB0CFEF6F6B8044D" box="[1200,1278,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="4.2672" pageId="8" pageNumber="255" unit="in" value="1.68">1.68 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFABAFEF7F7CC044C" box="[1286,1418,332,356]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B975F54DF373FFBDFABAFEF7F7CC044C" box="[1286,1418,332,356]" italics="true" pageId="8" pageNumber="255">L. hainanus</emphasis>
</taxonomicName>
); relatively shorter tibias in males (mean ratio of TIB/SVL
<quantity id="4CF984BAF373FFBDFC83FED5F1D704A0" box="[831,913,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.3462" pageId="8" pageNumber="255" unit="in" value="0.53">0.53 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFC27FECBF1A604A0" box="[923,992,368,392]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFC27FECBF1A604A0" box="[923,992,368,392]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFC57FED5F60F04A0" box="[1003,1097,367,392]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFC57FED5F60F04A0" bold="true" box="[1003,1097,367,392]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFB1AFED5F6BE04A0" box="[1190,1272,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.6256" pageId="8" pageNumber="255" unit="in" value="0.64">0.64 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFABFFECBF7CC04A0" box="[1283,1418,368,392]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B975F54DF373FFBDFABFFECBF7CC04A0" box="[1283,1418,368,392]" italics="true" pageId="8" pageNumber="255">L. hainanus</emphasis>
</taxonomicName>
); tibiotarsal articulation reaching the naris (versus beyond the tip of snout in
<taxonomicName id="4C0152DCF373FFBDFBB6FE2FF6D40484" box="[1034,1170,404,428]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B975F54DF373FFBDFBB6FE2FF6D40484" box="[1034,1170,404,428]" italics="true" pageId="8" pageNumber="255">L. hainanus</emphasis>
</taxonomicName>
); the absence of beige speckling on the dorsum (versus present in
<taxonomicName id="4C0152DCF373FFBDFD31FE03F15204F8" box="[653,788,440,464]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B975F54DF373FFBDFD31FE03F15204F8" box="[653,788,440,464]" italics="true" pageId="8" pageNumber="255">L. hainanus</emphasis>
</taxonomicName>
,
<figureCitation id="133A35DAF373FFBDFC99FE0DF12D04F8" box="[805,875,439,464]" captionStart="FIGURE 6. A" captionStartId="9.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1494]" captionTargetId="figure@9.[151,1436,193,1495]" captionTargetPageId="9" captionText="FIGURE 6. A: a male individual of Liuixalus feii sp. nov., photographed on 2 May 2010, not collected; B: L. hainanus from Diaoluoshan, NR Hainan (type locality); C: L. ocellatus from Wuzhishan NR, Hainan (type locality); D: L. romeri from Hong Kong (type locality); E: paratype of L. calcarius from Cat Ba Island, Vietnam; F: L. cf. romeri from Mt. Shiwan, Guangxi." httpUri="https://zenodo.org/record/239696/files/figure.png" pageId="8" pageNumber="255">Fig. 6</figureCitation>
); the distinct difference in eye color pattern (iris bicolor in
<taxonomicName id="4C0152DCF373FFBDFEB1FE67F31604DC" box="[269,336,476,500]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFEB1FE67F31604DC" box="[269,336,476,500]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFEE5FE66F3F204DD" box="[345,436,476,501]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFEE5FE66F3F204DD" bold="true" box="[345,436,476,501]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus iris yellowish overall with distinct small dark spots above and below pupil in
<taxonomicName id="4C0152DCF373FFBDFA39FE67F2B80730" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B975F54DF373FFBDFA39FE67F2B80730" italics="true" pageId="8" pageNumber="255">L. hainanus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0152DCF373FFBDFEA9FDBAF3EF0730" box="[277,425,512,536]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFEA9FDBAF3EF0730" box="[277,425,512,536]" italics="true" pageId="8" pageNumber="255">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFE0DFE45F04D0730" box="[433,523,511,536]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFE0DFE45F04D0730" bold="true" box="[433,523,511,536]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C0152DCF373FFBDFD1DFDBBF1630730" box="[673,805,512,536]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B975F54DF373FFBDFD1DFDBBF1630730" box="[673,805,512,536]" italics="true" pageId="8" pageNumber="255">L. ocellatus</emphasis>
</taxonomicName>
by having relatively smaller tympanum in males (mean ratio of TD/SVL
<quantity id="4CF984BAF373FFBDFEE0FD9EF3FF0715" box="[348,441,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.4224" pageId="8" pageNumber="255" unit="in" value="0.056">0.056 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFE7CFD9FF0450714" box="[448,515,548,572]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFE7CFD9FF0450714" box="[448,515,548,572]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFDB7FD9EF0230715" box="[523,613,548,573]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFDB7FD9EF0230715" bold="true" box="[523,613,548,573]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFD01FD9EF15C0715" box="[701,794,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.9811999999999999" pageId="8" pageNumber="255" unit="in" value="0.078">0.078 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFC9EFD9FF1E30714" box="[802,933,548,572]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B975F54DF373FFBDFC9EFD9FF1E30714" box="[802,933,548,572]" italics="true" pageId="8" pageNumber="255">L. ocellatus</emphasis>
</taxonomicName>
); relatively shorter snout (mean ratio of SL/ SVL
<quantity id="4CF984BAF373FFBDFF6FFDFDF3640748" box="[211,290,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.81" pageId="8" pageNumber="255" unit="in" value="0.15">0.15 in</quantity>
<emphasis id="B975F54DF373FFBDFE95FDF3F3060748" box="[297,320,585,608]" italics="true" pageId="8" pageNumber="255">L.</emphasis>
<taxonomicName id="4C0152DCF373FFBDFEF4FDFDF32B0748" box="[328,365,583,608]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">feii</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFEC8FDFDF3890748" box="[372,463,583,608]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFEC8FDFDF3890748" bold="true" box="[372,463,583,608]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFD99FDF2F0320748" box="[549,628,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="4.572" pageId="8" pageNumber="255" unit="in" value="0.18">0.18 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFDC0FDF3F0B90748" box="[636,767,584,608]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B975F54DF373FFBDFDC0FDF3F0B90748" box="[636,767,584,608]" italics="true" pageId="8" pageNumber="255">L. ocellatus</emphasis>
</taxonomicName>
); relatively longer hindlimbs in males (mean ratio of HLL/ SVL
<quantity id="4CF984BAF373FFBDFF6FFDD4F36707AF" box="[211,289,622,647]" metricMagnitude="-2" metricUnit="m" metricValue="4.2164" pageId="8" pageNumber="255" unit="in" value="1.66">1.66 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFE94FDCAF32C07AF" box="[296,362,623,647]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFE94FDCAF32C07AF" box="[296,362,623,647]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFECDFDD4F38D07AF" box="[369,459,622,647]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFECDFDD4F38D07AF" bold="true" box="[369,459,622,647]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFD9CFDD4F02907AF" box="[544,623,622,647]" metricMagnitude="-2" metricUnit="m" metricValue="4.1402" pageId="8" pageNumber="255" unit="in" value="1.63">1.63 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFDCAFDCAF0BF07AF" box="[630,761,623,647]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B975F54DF373FFBDFDCAFDCAF0BF07AF" box="[630,761,623,647]" italics="true" pageId="8" pageNumber="255">L. ocellatus</emphasis>
</taxonomicName>
); tip of 1st finger not enlarged and without disk (versus with tip of 1st finger with distinct disk in
<taxonomicName id="4C0152DCF373FFBDFD95FD2CF0ED0785" box="[553,683,661,685]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B975F54DF373FFBDFD95FD2CF0ED0785" box="[553,683,661,685]" italics="true" pageId="8" pageNumber="255">L. ocellatus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0152DCF373FFBDFD7FFD2FF1100786" box="[707,854,661,686]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFD7FFD2FF1100786" box="[707,854,661,686]" italics="true" pageId="8" pageNumber="255">Liuixalus feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFCE1FD2FF1F10786" box="[861,951,661,686]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFCE1FD2FF1F10786" bold="true" box="[861,951,661,686]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C0152DCF373FFBDFBF0FD2CF6F30785" box="[1100,1205,662,685]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="romeri">
<emphasis id="B975F54DF373FFBDFBF0FD2CF6F30785" box="[1100,1205,662,685]" italics="true" pageId="8" pageNumber="255">L. romeri</emphasis>
</taxonomicName>
by having relatively smaller tympanum in males (mean ratio of TD/SVL
<quantity id="4CF984BAF373FFBDFD41FD03F11B07FA" box="[765,861,697,722]" metricMagnitude="-3" metricUnit="m" metricValue="1.4224" pageId="8" pageNumber="255" unit="in" value="0.056">0.056 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFCDAFD00F1ED07F9" box="[870,939,697,721]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFCDAFD00F1ED07F9" box="[870,939,697,721]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFC09FD03F65407FA" box="[949,1042,697,722]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFC09FD03F65407FA" bold="true" box="[949,1042,697,722]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFBD2FD00F68807FA" box="[1134,1230,697,722]" metricMagnitude="-3" metricUnit="m" metricValue="2.2352" pageId="8" pageNumber="255" unit="in" value="0.088">0.088 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFB64FD00F70207F9" box="[1240,1348,698,721]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="romeri">
<emphasis id="B975F54DF373FFBDFB64FD00F70207F9" box="[1240,1348,698,721]" italics="true" pageId="8" pageNumber="255">L. romeri</emphasis>
</taxonomicName>
); snout longer than eye diameter in males (mean ratio of ED/SL
<quantity id="4CF984BAF373FFBDFCB3FD67F11B07DE" box="[783,861,733,758]" metricMagnitude="-2" metricUnit="m" metricValue="2.159" pageId="8" pageNumber="255" unit="in" value="0.85">0.85 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFCDFFD64F1E207DE" box="[867,932,734,758]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="feii" status="sp. nov.">
<emphasis id="B975F54DF373FFBDFCDFFD64F1E207DE" box="[867,932,734,758]" italics="true" pageId="8" pageNumber="255">L. feii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2464836F373FFBDFC16FD67F64507DE" box="[938,1027,733,758]" pageId="8" pageNumber="255" rank="species">
<emphasis id="B975F54DF373FFBDFC16FD67F64507DE" bold="true" box="[938,1027,733,758]" pageId="8" pageNumber="255">sp. nov.</emphasis>
</taxonomicNameLabel>
versus
<quantity id="4CF984BAF373FFBDFBE4FD64F6E307DE" box="[1112,1189,733,758]" metricMagnitude="-2" metricUnit="m" metricValue="2.7178" pageId="8" pageNumber="255" unit="in" value="1.07">1.07 in</quantity>
<taxonomicName id="4C0152DCF373FFBDFB10FD64F75207DD" box="[1196,1300,734,757]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="romeri">
<emphasis id="B975F54DF373FFBDFB10FD64F75207DD" box="[1196,1300,734,757]" italics="true" pageId="8" pageNumber="255">L. romeri</emphasis>
</taxonomicName>
); tibiotarsal articulation reaching the naris (versus reaching the tip of snout in
<taxonomicName id="4C0152DCF373FFBDFC18FCB8F6570631" box="[932,1041,770,793]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="romeri">
<emphasis id="B975F54DF373FFBDFC18FCB8F6570631" box="[932,1041,770,793]" italics="true" pageId="8" pageNumber="255">L. romeri</emphasis>
</taxonomicName>
); fingers free of webbing (versus weakly webbing present between fingers III and V in
<taxonomicName id="4C0152DCF373FFBDFD51FC9CF1130615" box="[749,853,806,829]" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="romeri">
<emphasis id="B975F54DF373FFBDFD51FC9CF1130615" box="[749,853,806,829]" italics="true" pageId="8" pageNumber="255">L. romeri</emphasis>
</taxonomicName>
); toes weakly webbed (versus toes 1/3 webbed in
<taxonomicName id="4C0152DCF373FFBDFA39FC9CF2A70649" class="Amphibia" family="Rhacophoridae" genus="Liuixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="255" phylum="Chordata" rank="species" species="romeri">
<emphasis id="B975F54DF373FFBDFA39FC9CF2A70649" italics="true" pageId="8" pageNumber="255">L. romeri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>