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<document id="50BF390A824376ED1BDC408DC0A6C70A" ID-DOI="10.11646/zootaxa.5506.1.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="13745886" ID-ZooBank="51FBDADB-CEFC-4A87-A6FA-CF5630D742AE" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1726052320197" checkinUser="plazi" docAuthor="Souza-Gonçalves, Igor &amp; Lopes-Andrade, Cristiano" docDate="2024" docId="630487D6814BFFD0F2E302BCFB955A18" docLanguage="en" docName="zootaxa.5506.1.10.pdf" docOrigin="Zootaxa 5506 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.1.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nipponocis palawanus Souza-Gonçalves &amp; Lopes-Andrade 2024, comb. nov." docType="treatment" docVersion="3" lastPageNumber="140" masterDocId="9F3DFFAE8148FFD3F274032FFF935D68" masterDocTitle="On the Ciidae (Coleoptera) described by Michio Chûjô deposited in the Natural History Museum of Denmark" masterLastPageNumber="144" masterPageNumber="137" pageNumber="140" updateTime="1726241149630" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="236B5BEDCF2344376920F85C15E66F5A">On the Ciidae (Coleoptera) described by Michio Chûjô deposited in the Natural History Museum of Denmark</mods:title>
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<mods:namePart id="8FF1B4D082D98033953216C1D72DB562">Souza-Gonçalves, Igor</mods:namePart>
<mods:affiliation id="20050AD5228D98C5EBD011FD76E111F3">Laboratório de Taxonomia de Insetos, Departamento de Agronomia / Fitossanidade, Universidade Federal Rural de Pernambuco, 52171 - 900, Recife, Pernambuco, Brazil &amp; Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
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<mods:namePart id="AE3796D518EEAAB861A60ABA954BC832">Lopes-Andrade, Cristiano</mods:namePart>
<mods:affiliation id="4B5D27F1536F1ACE8A58B6BC2C3E2BB8">Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
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<subSubSection id="A3B7654B814BFFD0F2E302BCFD795CC5" box="[151,746,402,429]" pageId="3" pageNumber="140" type="nomenclature">
<paragraph id="EB1236C0814BFFD0F2E302BCFD795CC5" blockId="3.[151,746,402,465]" box="[151,746,402,429]" pageId="3" pageNumber="140">
<heading id="B05A81AC814BFFD0F2E302BCFD795CC5" bold="true" box="[151,746,402,429]" fontSize="11" level="1" pageId="3" pageNumber="140" reason="1">
<taxonomicName id="2CAD4D43814BFFD0F2E302BCFDCF5CC4" authority="(Chujo, 1966)" authorityName="Souza-Gonçalves &amp; Lopes-Andrade" authorityYear="2024" baseAuthorityName="Chujo" baseAuthorityYear="1966" box="[151,604,402,429]" class="Insecta" family="Ciidae" genus="Nipponocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="palawanus" status="comb. nov.">
Nipponocis palawanus (
<bibRefCitation id="8F3C4B31814BFFD0F3CD02BDFDC75CC4" author="Chujo, M." box="[441,596,402,429]" pageId="3" pageNumber="140" pagination="521 - 532" refId="ref3237" refString="Chujo, M. (1966) Some Bostrihidae, Languriidae, and Ciidae (Col.) from the Philippine and Bismarck Islands collected by the Noona Dan Expedition. Entomologiske Meddelelser, 34, 521 - 532." type="journal article" year="1966">Chûjô, 1966</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C2EA57A9814BFFD0F01002BCFD795CC5" box="[612,746,403,429]" pageId="3" pageNumber="140" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A3B7654B814BFFD0F2E30298FE9C5CB9" box="[151,271,439,465]" pageId="3" pageNumber="140" type="description">
<paragraph id="EB1236C0814BFFD0F2E30298FE9C5CB9" blockId="3.[151,746,402,465]" box="[151,271,439,465]" pageId="3" pageNumber="140">
<figureCitation id="73962A45814BFFD0F2E30298FE9C5CB9" box="[151,271,439,465]" captionStart="FIGURE 3" captionStartId="3.[151,250,1447,1471]" captionTargetBox="[257,1322,555,1420]" captionTargetId="figure-107@3.[256,1331,555,1423]" captionTargetPageId="3" captionText="FIGURE 3. Nipponocis palawanus (Chûjô, 1966) comb. nov., holotype (A–D) from Brooke’s Point (Palawan, Philippines): A. Dorsal view. B. Lateral view. C. Ventral view. D. Labels. Scale bar: 1 mm (A–C)." figureDoi="http://doi.org/10.5281/zenodo.13745892" httpUri="https://zenodo.org/record/13745892/files/figure.png" pageId="3" pageNumber="140">Fig. 3A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A3B7654B814BFFD0F2E302D3FC925F7B" box="[151,769,507,531]" pageId="3" pageNumber="140" type="reference_group">
<paragraph id="EB1236C0814BFFD0F2E302D3FC925F7B" blockId="3.[151,769,507,531]" box="[151,769,507,531]" pageId="3" pageNumber="140">
<treatmentCitationGroup id="CBBD11EE814BFFD0F2E302D3FC925F7B" box="[151,769,507,531]" pageId="3" pageNumber="140">
<treatmentCitation id="6A0C10D1814BFFD0F2E302D3FD8E5F7B" author="Chujo, M." box="[151,541,507,531]" page="528" pageId="3" pageNumber="140" year="1966">
<taxonomicName id="2CAD4D43814BFFD0F2E302D3FD8E5F7B" authority="Chujo, 1966: 528" authorityName="Chujo" authorityPageNumber="528" authorityYear="1966" box="[151,541,507,531]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="palawanus" subGenus="Cis">
Cis (Cis) palawanus
<bibRefCitation id="8F3C4B31814BFFD0F31802D4FD8E5F7B" author="Chujo, M." box="[364,541,507,531]" pageId="3" pageNumber="140" pagination="521 - 532" refId="ref3237" refString="Chujo, M. (1966) Some Bostrihidae, Languriidae, and Ciidae (Col.) from the Philippine and Bismarck Islands collected by the Noona Dan Expedition. Entomologiske Meddelelser, 34, 521 - 532." type="journal article" year="1966">Chûjô, 1966: 528</bibRefCitation>
</taxonomicName>
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(original description).
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</paragraph>
</subSubSection>
<caption id="BFD26648814BFFD0F2E30688FC41588C" ID-DOI="http://doi.org/10.5281/zenodo.13745892" ID-Zenodo-Dep="13745892" httpUri="https://zenodo.org/record/13745892/files/figure.png" pageId="3" pageNumber="140" startId="3.[151,250,1447,1471]" targetBox="[257,1322,555,1420]" targetPageId="3" targetType="figure">
<paragraph id="EB1236C0814BFFD0F2E30688FC41588C" blockId="3.[151,1436,1447,1508]" pageId="3" pageNumber="140">
FIGURE 3.
<taxonomicName id="2CAD4D43814BFFD0F36C0687FD0158A8" authority="(Chujo, 1966)" authorityName="Souza-Gonçalves &amp; Lopes-Andrade" authorityYear="2024" baseAuthorityName="Chujo" baseAuthorityYear="1966" box="[280,658,1448,1472]" class="Insecta" family="Ciidae" genus="Nipponocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="palawanus" status="comb. nov.">
Nipponocis palawanus (
<bibRefCitation id="8F3C4B31814BFFD0F0780687FD1F58A8" author="Chujo, M." box="[524,652,1448,1472]" pageId="3" pageNumber="140" pagination="521 - 532" refId="ref3237" refString="Chujo, M. (1966) Some Bostrihidae, Languriidae, and Ciidae (Col.) from the Philippine and Bismarck Islands collected by the Noona Dan Expedition. Entomologiske Meddelelser, 34, 521 - 532." type="journal article" year="1966">Chûjô, 1966</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C2EA57A9814BFFD0F0EC0687FC9858A8" box="[664,779,1448,1472]" pageId="3" pageNumber="140" rank="species">comb. nov.</taxonomicNameLabel>
, holotype (A–D) from Brooke’s Point (Palawan, Philippines): A. Dorsal view. B. Lateral view. C. Ventral view. D. Labels. Scale bar: 1 mm (A–C).
</paragraph>
</caption>
<subSubSection id="A3B7654B814BFFD0F2E30519FB955A18" pageId="3" pageNumber="140" type="discussion">
<paragraph id="EB1236C0814BFFD0F2E30519FB955A18" blockId="3.[151,1437,1590,1904]" pageId="3" pageNumber="140">
Comments. From the Province of
<collectingRegion id="2969F822814BFFD0F05F0519FD1E5B38" box="[555,653,1590,1616]" country="Philippines" name="Palawan" pageId="3" pageNumber="140">Palawan</collectingRegion>
,
<collectingCountry id="93BA7650814BFFD0F0EE0519FC8F5B38" box="[666,796,1590,1616]" name="Philippines" pageId="3" pageNumber="140">Philippines</collectingCountry>
. This species is known only from the
<typeStatus id="34168862814BFFD0F6AE0519FAAF5B38" box="[1242,1340,1590,1616]" pageId="3" pageNumber="140" type="holotype">holotype</typeStatus>
and fits the limits of
<taxonomicName id="2CAD4D43814BFFD0F3450574FD405B1D" authority="Nobuchi &amp; Wada, 1955" authorityName="Nobuchi &amp; Wada" authorityYear="1955" box="[305,723,1626,1653]" class="Insecta" family="Ciidae" genus="Nipponocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">Nipponocis Nobuchi &amp; Wada, 1955</taxonomicName>
: body elongate, large, parallel-sided; 10-segmented antennae; dual pronotal and elytral punctation; all tibiae with simple outer apical angle; and males with anterocephalic edge with two tubercles and first abdominal ventrite without sex patch (
<bibRefCitation id="8F3C4B31814BFFD0F1EF058DFBD75BD5" author="Nobuchi, A." box="[923,1092,1698,1725]" pageId="3" pageNumber="140" pagination="53 - 58" refId="ref3686" refString="Nobuchi, A. (1955) Studies on the Ciid-Beetles from Japan. (I) With the Descriptions of a New Genus and Some New Species (Ciidae, Coleoptera). Entomological Review of Japan, 6 (7), 53 - 58, pl. 12." type="journal article" year="1955">Nobuchi 1955</bibRefCitation>
). It differs from other species in the genus, except for
<taxonomicName id="2CAD4D43814BFFD0F3C605E8FCCD5B89" authority="Nobuchi, 1959" authorityName="Nobuchi" authorityYear="1959" box="[434,862,1734,1761]" class="Insecta" family="Ciidae" genus="Nipponocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="ashuensis">
Nipponocis ashuensis
<bibRefCitation id="8F3C4B31814BFFD0F0C505E9FCCD5B89" author="Nobuchi, A." box="[689,862,1734,1761]" pageId="3" pageNumber="140" pagination="51 - 52" refId="ref3738" refString="Nobuchi, A. (1959) Two New Species of Ciid-Beetles. Entomological Review of Japan, 10 (2), 51 - 52." type="journal article" year="1959">Nobuchi, 1959</bibRefCitation>
</taxonomicName>
, in the emarginate anterior edge of the pronotum.
<taxonomicName id="2CAD4D43814BFFD0F2E305C4FE1F5A6C" authorityName="Nobuchi" authorityYear="1959" box="[151,396,1770,1796]" class="Insecta" family="Ciidae" genus="Nipponocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="ashuensis">Nipponocis ashuensis</taxonomicName>
differs from
<taxonomicName id="2CAD4D43814BFFD0F05705C4FD405A6C" authorityName="Chujo" authorityYear="1966" box="[547,723,1770,1796]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="palawanus">Nip. palawanus</taxonomicName>
in the dual dorsal vestiture, consisting of short decumbent and long erect setae. Besides
<taxonomicName id="2CAD4D43814BFFD0F3CE0420FDF35A40" authorityName="Nobuchi" authorityYear="1959" box="[442,608,1806,1832]" class="Insecta" family="Ciidae" genus="Nipponocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="ashuensis">Nip. ashuensis</taxonomicName>
and
<taxonomicName id="2CAD4D43814BFFD0F0EE0420FCDF5A40" authorityName="Chujo" authorityYear="1966" box="[666,844,1806,1832]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="palawanus">Nip. palawanus</taxonomicName>
,
<taxonomicName id="2CAD4D43814BFFD0F1280420FC4F5A40" authorityName="Nobuchi &amp; Wada" authorityYear="1955" box="[860,988,1807,1832]" class="Insecta" family="Ciidae" genus="Nipponocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">Nipponocis</taxonomicName>
includes three other described species: Nip. longisetosus
<bibRefCitation id="8F3C4B31814BFFD0F329041DFD965A25" author="Nobuchi, A." box="[349,517,1842,1869]" pageId="3" pageNumber="140" pagination="53 - 58" refId="ref3686" refString="Nobuchi, A. (1955) Studies on the Ciid-Beetles from Japan. (I) With the Descriptions of a New Genus and Some New Species (Ciidae, Coleoptera). Entomological Review of Japan, 6 (7), 53 - 58, pl. 12." type="journal article" year="1955">Nobuchi, 1955</bibRefCitation>
, Nip. magnus
<bibRefCitation id="8F3C4B31814BFFD0F0D6041DFCD45A25" author="Nobuchi, A." box="[674,839,1842,1869]" pageId="3" pageNumber="140" pagination="53 - 58" refId="ref3686" refString="Nobuchi, A. (1955) Studies on the Ciid-Beetles from Japan. (I) With the Descriptions of a New Genus and Some New Species (Ciidae, Coleoptera). Entomological Review of Japan, 6 (7), 53 - 58, pl. 12." type="journal article" year="1955">Nobuchi, 1955</bibRefCitation>
and Nip. unipunctatus
<bibRefCitation id="8F3C4B31814BFFD0F63C041DFAF45A25" author="Nakane, T. &amp; Nobuchi, A." box="[1096,1383,1842,1869]" pageId="3" pageNumber="140" pagination="55 - 56" refId="ref3632" refString="Nakane, T. &amp; Nobuchi, A. (1956) Entomological Results from the Scientific Survey of the Tokara Islands. II Coleoptera: Ciidae. Scientific Reports of the Saikyo University (Natural Science and Living Science), A Series, 2 (2), 55 - 56." type="journal article" year="1956">Nakane &amp; Nobuchi, 1956</bibRefCitation>
. The genus occurs in the Indo-Malayan transition zone and Chinese transition zone.
</paragraph>
</subSubSection>
</treatment>
</document>
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