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<document id="4ABF877E935132EE82C35B2D1EFE49F6" ID-DOI="10.5281/zenodo.200811" ID-GBIF-Dataset="76b2912f-8856-4c85-842d-c3312bc2650f" ID-ISSN="1175-5326" ID-Zenodo-Dep="200811" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460250427582" checkinUser="plazi" docAuthor="Mcdonald, Andrew T." docDate="2011" docId="E80287E9B11CFFFBCDB954FBFDF5FD59" docLanguage="en" docName="zt02783p068.pdf" docOrigin="Zootaxa 2783" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Planicoxa venenica DiCroce &amp; Carpenter 2001" docType="treatment" docVersion="7" lastPageNumber="66" masterDocId="143BFF91B110FFF5CD2E537DFFE3FF95" masterDocTitle="The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)" masterLastPageNumber="68" masterPageNumber="52" pageNumber="64" updateTime="1698242557538" updateUser="plazi">
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<mods:title id="C16037D7F3B5D155C15287554F36F89C">The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)</mods:title>
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<mods:namePart id="CE09E0DABBC247F49974FECB08912998">Mcdonald, Andrew T.</mods:namePart>
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<mods:date id="86AAC68CA0A92BF95A63C4AE1B936248">2011</mods:date>
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<treatment id="E80287E9B11CFFFBCDB954FBFDF5FD59" ID-DOI="http://doi.org/10.5281/zenodo.6185219" ID-GBIF-Taxon="119391373" ID-Zenodo-Dep="6185219" LSID="urn:lsid:plazi:treatment:E80287E9B11CFFFBCDB954FBFDF5FD59" httpUri="http://treatment.plazi.org/id/E80287E9B11CFFFBCDB954FBFDF5FD59" lastPageId="14" lastPageNumber="66" pageId="12" pageNumber="64">
<subSubSection id="28B16574B11CFFF9CDB954FBFD01F877" pageId="12" pageNumber="64" type="nomenclature">
<paragraph id="601436FFB11CFFF9CDB954FBFD01F835" blockId="12.[151,738,1926,1952]" box="[151,738,1926,1952]" pageId="12" pageNumber="64">
<heading id="3B5C8193B11CFFF9CDB954FBFD01F835" bold="true" box="[151,738,1926,1952]" fontSize="11" level="1" pageId="12" pageNumber="64" reason="1">
<taxonomicName id="A7AB4D7CB11CFFF9CDB954FBFD01F835" authority="DiCroce &amp; Carpenter, 2001" authorityName="DiCroce &amp; Carpenter" authorityYear="2001" box="[151,738,1926,1952]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11CFFF9CDB954FBFD01F835" bold="true" box="[151,738,1926,1952]" pageId="12" pageNumber="64">
<emphasis id="52DFEAEDB11CFFF9CDB954FBFE60F835" bold="true" box="[151,387,1926,1952]" italics="true" pageId="12" pageNumber="64">Planicoxa venenica</emphasis>
<bibRefCitation id="043A4B0EB11CFFF9CCA554FBFD01F835" author="DiCroce" box="[395,738,1926,1952]" pageId="12" pageNumber="64" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="601436FFB11CFFF9CDB954B0FD01F877" blockId="12.[151,738,1997,2019]" box="[151,738,1997,2019]" pageId="12" pageNumber="64">
<bibRefCitation id="043A4B0EB11CFFF9CDB954B0FF29F877" author="Carpenter" box="[151,202,1997,2018]" pageId="12" pageNumber="64" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">2001</bibRefCitation>
<taxonomicName id="A7AB4D7CB11CFFF9CDC754B0FD72F877" authority="DiCroce &amp; Carpenter" authorityName="DiCroce &amp; Carpenter" box="[233,657,1997,2019]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11CFFF9CDC754B0FE53F876" box="[233,432,1997,2019]" italics="true" pageId="12" pageNumber="64">Planicoxa venenica</emphasis>
DiCroce &amp; Carpenter
</taxonomicName>
, p. 190.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B11DFFFBCDB953E5FAE4FEAA" lastPageId="14" lastPageNumber="66" pageId="13" pageNumber="65" type="materials_examined">
<paragraph id="601436FFB11DFFF8CDB953E5FDD2FF3A" blockId="13.[151,1436,151,284]" box="[151,561,151,176]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDB953E5FEE8FF25" bold="true" box="[151,267,152,176]" pageId="13" pageNumber="65">
<typeStatus id="BF10885DB11DFFF8CDB953E5FEE5FF25" box="[151,262,152,176]" pageId="13" pageNumber="65" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="06BAAE3AB11DFFF8CC3C53E5FE8BFF3A" LSID="urn:lsid:biocol.org:col:34940" box="[274,360,152,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="13" pageNumber="65">DMNH</collectionCode>
42504, left ilium.
</paragraph>
<paragraph id="601436FFB11DFFF8CDEB53C1FE91FE89" blockId="13.[151,1436,151,284]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDEB53C1FEA7FF41" bold="true" box="[197,324,188,212]" pageId="13" pageNumber="65">
<typeStatus id="BF10885DB11DFFF8CDEB53C1FEDCFF41" box="[197,319,188,212]" pageId="13" pageNumber="65" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="06BAAE3AB11DFFF8CC6253C0FE41FF41" LSID="urn:lsid:biocol.org:col:34940" box="[332,418,189,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="13" pageNumber="65">DMNH</collectionCode>
42511, 42516, 42518, 42519, 42520, 42521, 42522, 42524, 42513, 42515, 42525, 42523, 42526, 42527, 42510, 42514, 42517, 42508, 42507, 42505, 40917, 40914, 40918, 42506, 42509, 42512 (
<bibRefCitation id="043A4B0EB11DFFF8C81553A2FE86FE89" author="DiCroce" pageId="13" pageNumber="65" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter 2001</bibRefCitation>
).
</paragraph>
<caption id="34D46677B11DFFF8CDB95732FA8EFB11" httpUri="https://zenodo.org/record/200822/files/figure.png" pageId="13" pageNumber="65" targetBox="[151,1436,327,1082]" targetPageId="13">
<paragraph id="601436FFB11DFFF8CDB95732FA8EFB11" blockId="13.[151,1436,1103,1156]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDB95732FEFDFBF0" bold="true" box="[151,286,1103,1126]" pageId="13" pageNumber="65">FIGURE 11.</emphasis>
Left (
<emphasis id="52DFEAEDB11DFFF8CC70572DFE93FBF0" bold="true" box="[350,368,1104,1125]" pageId="13" pageNumber="65">A</emphasis>
) and right (
<emphasis id="52DFEAEDB11DFFF8CCC7572DFE19FBF0" bold="true" box="[489,506,1104,1125]" pageId="13" pageNumber="65">B</emphasis>
, reversed) ilia of USNM 4753 (holotype of
<taxonomicName id="A7AB4D7CB11DFFF8CE945732FB59FBF0" box="[954,1210,1103,1125]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="65" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11DFFF8CE945732FB59FBF0" box="[954,1210,1103,1125]" italics="true" pageId="13" pageNumber="65">Osmakasaurus depressus</emphasis>
</taxonomicName>
) in lateral view, set to the same scale and with the notch between the preacetabular process and pubic peduncle lined up. Scale bar equals 10 cm.
</paragraph>
</caption>
<caption id="34D46677B11DFFF8CDB954C3FD1DF866" httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="13" pageNumber="65" targetBox="[151,1436,1206,1961]" targetPageId="13">
<paragraph id="601436FFB11DFFF8CDB954C3FD1DF866" blockId="13.[151,1437,1982,2035]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDB954C3FEC3F841" bold="true" box="[151,288,1982,2005]" pageId="13" pageNumber="65">FIGURE 12.</emphasis>
Left ilium of
<taxonomicName id="A7AB4D7CB11DFFF8CC9D54C2FD98F840" box="[435,635,1983,2005]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="65" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11DFFF8CC9D54C2FD98F840" box="[435,635,1983,2005]" italics="true" pageId="13" pageNumber="65">Planicoxa venenica</emphasis>
</taxonomicName>
(DMNH 42504, holotype) in (
<emphasis id="52DFEAEDB11DFFF8CE9754C2FC28F841" bold="true" box="[953,971,1983,2004]" pageId="13" pageNumber="65">A</emphasis>
) dorsal and (
<emphasis id="52DFEAEDB11DFFF8C97B54C2FB85F841" bold="true" box="[1109,1126,1983,2004]" pageId="13" pageNumber="65">B</emphasis>
) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm.
</paragraph>
</caption>
<paragraph id="601436FFB11EFFFBCDEB53EAFC47FF41" blockId="14.[151,1436,151,716]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB53EAFE24FF25" bold="true" box="[197,455,151,176]" pageId="14" pageNumber="66">Locality and horizon.</emphasis>
Tonys Bone Bed, near Moab, Grand County, Utah (
<bibRefCitation id="043A4B0EB11EFFFBC93153EAFABBFF3A" author="DiCroce" box="[1055,1368,151,175]" pageId="14" pageNumber="66" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter 2001</bibRefCitation>
); Poison Strip Member, Cedar Mountain Formation, lowermiddle Aptian.
</paragraph>
<paragraph id="601436FFB11EFFFBCDEB53A2FD04FF6D" blockId="14.[151,1436,151,716]" box="[197,743,223,248]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB53A2FE43FF6D" bold="true" box="[197,416,223,248]" pageId="14" pageNumber="66">Generic diagnosis.</emphasis>
<collectingCountry id="18BC766FB11EFFFBCC89539DFE25FF6D" box="[423,454,224,248]" name="American Samoa" pageId="14" pageNumber="66">As</collectingCountry>
for species by monotypy.
</paragraph>
<paragraph id="601436FFB11EFFFBCDEB527EFAE4FEAA" blockId="14.[151,1436,151,716]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB527EFE7DFE89" bold="true" box="[197,414,259,284]" pageId="14" pageNumber="66">Specific diagnosis.</emphasis>
Provisionally diagnostic basal styracosternan distinguished from
<taxonomicName id="A7AB4D7CB11EFFFBC9555279FA77FE89" box="[1147,1428,260,284]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11EFFFBC9555279FA77FE89" box="[1147,1428,260,284]" italics="true" pageId="14" pageNumber="66">Osmakasaurus depressus</emphasis>
</taxonomicName>
,
<taxonomicName id="A7AB4D7CB11EFFFBCDB95255FE6BFED5" box="[151,392,296,320]" class="Reptilia" genus="Cedrorestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="crichtoni">
<emphasis id="52DFEAEDB11EFFFBCDB95255FE6BFED5" box="[151,392,296,320]" italics="true" pageId="14" pageNumber="66">Cedrorestes crichtoni</emphasis>
</taxonomicName>
, and
<emphasis id="52DFEAEDB11EFFFBCCEB5255FD55FED5" box="[453,694,296,320]" italics="true" pageId="14" pageNumber="66">Iguanacolossus fortis</emphasis>
by the convex dorsal margin of the ilium (
<figureCitation id="F8902A7AB11EFFFBC9BA525AFB05FEAA" box="[1172,1254,295,319]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="28B16574B11EFFFBCDEB5231FDF5FD59" pageId="14" pageNumber="66" type="discussion">
<paragraph id="601436FFB11EFFFBCDEB5231FDF5FD59" blockId="14.[151,1436,151,716]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB5231FEDAFEF1" bold="true" box="[197,313,332,356]" pageId="14" pageNumber="66">Remarks.</emphasis>
The reassessment of USNM 4753 given above casts doubt upon the supposed horizontal postacetabular process of DMNH 42504, which was the primary feature used by
<bibRefCitation id="043A4B0EB11EFFFBCE995212FB1FFE12" author="DiCroce" box="[951,1276,367,391]" pageId="14" pageNumber="66" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter (2001)</bibRefCitation>
in diagnosing
<taxonomicName id="A7AB4D7CB11EFFFBCDB952E9FE96FE39" box="[151,373,404,428]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11EFFFBCDB952E9FE96FE39" box="[151,373,404,428]" italics="true" pageId="14" pageNumber="66">Planicoxa venenica</emphasis>
</taxonomicName>
. In dorsal view, the postacetabular process of DMNH 42504 is heavily fractured (
<figureCitation id="F8902A7AB11EFFFBC80B52E9FA9BFE39" box="[1317,1400,404,428]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
A). There are additional smaller cracks along the dorsal margin of the ilium visible in lateral view (
<figureCitation id="F8902A7AB11EFFFBC9C152CAFAA0FE5A" box="[1263,1347,439,463]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
B). It is likely that the fractures on the postacetabular process and dorsal margin are the results of deformation of the ilium, akin to what is proposed above for the left ilium of USNM 4753. If so, then the diagnosis of
<taxonomicName id="A7AB4D7CB11EFFFBC990517DFA7FFD8D" box="[1214,1436,512,536]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11EFFFBC990517DFA7FFD8D" box="[1214,1436,512,536]" italics="true" pageId="14" pageNumber="66">Planicoxa venenica</emphasis>
</taxonomicName>
loses its key feature. Nevertheless, pending recovery of additional iguanodont specimens from the Poison Strip Member,
<taxonomicName id="A7AB4D7CB11EFFFBCC2A5135FE01FDF5" box="[260,482,584,608]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11EFFFBCC2A5135FE01FDF5" box="[260,482,584,608]" italics="true" pageId="14" pageNumber="66">Planicoxa venenica</emphasis>
</taxonomicName>
is herein retained as a provisionally viable taxon, as its ilium can still be differentiated from all other basal styracosternan ilia known from the Early Cretaceous of North
<collectingCountry id="18BC766FB11EFFFBC9445111FB29FD11" box="[1130,1226,620,644]" name="United States of America" pageId="14" pageNumber="66">America</collectingCountry>
by its convex dorsal margin. The dorsal margin of the ilium is convex cranial to the distorted part of the bone, indicating that this is probably a real feature (
<figureCitation id="F8902A7AB11EFFFBCC8D51C9FE16FD59" box="[419,501,692,716]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
B).
</paragraph>
</subSubSection>
</treatment>
</document>