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<document id="4C7311F2F6B1F786CBA0F6832EE66FD3" ID-CLB-Dataset="6394" ID-DOI="10.5252/g2014n3a3" ID-GBIF-Dataset="d28ddaf8-3b50-46bb-89a4-0e325e35b705" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538531" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613150579605" checkinUser="felipe" docAuthor="Sanjuan, Josep &amp; Martín-Closas, Carles" docDate="2014" docId="D25987DDFFF3FFACFF2AED262B3B8CAB" docLanguage="en" docName="Geodiversitas.36.3.385-420.pdf" docOrigin="Geodiversitas 36 (3)" docSource="http://dx.doi.org/10.5252/g2014n3a3" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Harrisichara vasiformis" docType="treatment" docVersion="8" lastPageNumber="407" masterDocId="2E60FFA5FFE5FFBAFFD1E84D2F748863" masterDocTitle="Taxonomy and palaeobiogeography of charophytes from the Upper Eocene- Lower Oligocene of the Eastern Ebro Basin (Catalonia, NE Spain)" masterLastPageNumber="420" masterPageNumber="385" pageNumber="407" updateTime="1698929445887" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="FD90A05F4FFAE5EB90A8218092DDE308">Taxonomy and palaeobiogeography of charophytes from the Upper Eocene- Lower Oligocene of the Eastern Ebro Basin (Catalonia, NE Spain)</mods:title>
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<treatment id="D25987DDFFF3FFACFF2AED262B3B8CAB" ID-DOI="http://doi.org/10.5281/zenodo.4836384" ID-GBIF-Taxon="182403514" ID-Zenodo-Dep="4836384" LSID="urn:lsid:plazi:treatment:D25987DDFFF3FFACFF2AED262B3B8CAB" httpUri="http://treatment.plazi.org/id/D25987DDFFF3FFACFF2AED262B3B8CAB" lastPageNumber="407" pageId="22" pageNumber="407">
<subSubSection id="12EA6540FFF3FFACFF2AED262D0D8DC5" pageId="22" pageNumber="407" type="nomenclature">
<paragraph id="5A4F36CBFFF3FFACFF2AED262D0D8DC5" blockId="22.[126,633,1387,1477]" pageId="22" pageNumber="407">
<heading id="010781A7FFF3FFACFF2AED262E898DE6" box="[251,509,1387,1414]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="407" reason="2">
<taxonomicName id="9DF04D48FFF3FFACFF2AED262E898DE6" authority="Feist-Castel, 1977" box="[251,509,1387,1414]" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="vasiformis">
<emphasis id="6884EAD9FFF3FFACFF2AED262E898DE6" bold="true" box="[251,509,1387,1414]" italics="true" pageId="22" pageNumber="407">Harrisichara vasiformis</emphasis>
</taxonomicName>
</heading>
form
<emphasis id="6884EAD9FFF3FFACFF6AEDC62EDA8DC5" bold="true" box="[187,430,1419,1446]" italics="true" pageId="22" pageNumber="407">vasiformis-tuberculata</emphasis>
Feist-Castel, 1977
</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF3FFACFEEDEDE62ECE8DA6" box="[316,442,1451,1477]" pageId="22" pageNumber="407" type="description">
<paragraph id="5A4F36CBFFF3FFACFEEDEDE62ECE8DA6" blockId="22.[126,633,1387,1477]" box="[316,442,1451,1477]" pageId="22" pageNumber="407">
(
<figureCitation id="C2CB2A4EFFF3FFACFE95EDE62EF78DA6" box="[324,387,1451,1477]" captionStart="FIG" captionStartId="24.[113,124,1521,1539]" captionTargetBox="[156,1191,221,1464]" captionTargetId="figure-205@24.[130,750,752,1292]" captionTargetPageId="24" captionText="FIG. 9. — Charophytes from Sant Boi, Artés and lateral south-western Formations, Ebro Basin: A-C, Harrisichara vasiformis form vasiformis-tuberculata Feist-Castel,1977, Serrat Rodó,Sobremunt road and Cal Carreter sections; A, apical view, no. 80113 MGSCB sample SBR-1; B, lateral view, no. 80114 MGSCB, sample SBR-1; C, basal view, no.80115 MGSCB, sample SBR-1;D-I, Harrisichara lineata Grambast,1957,El Perers section;D, apical view, no. 80116 MGSCB,sample PE-1; E, lateral view, no. 80117 MGSCB,sample PE-1; F, basal view, no. 80118 MGSCB, sample PE-1; G, lateral view, no. 80119 MGSCB, sample PE-1; H, lateral view, no. 80120 MGSCB, sample PE-1; I, lateral view, no. 80121 MGSCB, sample PE-1; J-L, Harrisichara tuberculata (Lyell, 1826), Oristà, Santa Maria dOló, Torre Casanova, Moià, Santpedor, Rubió and Sarral sections and Solivella outcrop; J, apical view, no. 80122 MGSCB, sample MO-1; K, lateral view, no. 80123 MGSCB, sample SO-2; L, basal view, no. 80124 MGSCB, sample MO-1. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4538547" httpUri="https://zenodo.org/record/4538547/files/figure.png" pageId="22" pageNumber="407">Fig. 9</figureCitation>
A-C)
</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF3FFACFFA0EDA62E498E7D" pageId="22" pageNumber="407" type="reference_group">
<paragraph id="5A4F36CBFFF3FFACFFA0EDA62E498E7D" blockId="22.[113,647,1515,1566]" pageId="22" pageNumber="407">
<taxonomicName id="9DF04D48FFF3FFACFFA0EDA62FED8E7E" authority="Feist-Castel, 1977 b: 152" authorityName="Feist-Castel" authorityPageNumber="152" authorityYear="1977" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="vasiformis-tuberculata">
<emphasis id="6884EAD9FFF3FFACFFA0EDA62EC98E60" box="[113,445,1515,1539]" italics="true" pageId="22" pageNumber="407">Harrisichara vasiformis-tuberculata</emphasis>
Feist-Castel, 1977b: 152
</taxonomicName>
, pl. 21, figs 2, 3.
</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF3FFACFFA0EE712C9489BE" pageId="22" pageNumber="407" type="distribution">
<paragraph id="5A4F36CBFFF3FFACFFA0EE712C9489BE" blockId="22.[113,650,1596,1699]" lastBlockId="22.[675,1210,215,477]" pageId="22" pageNumber="407">
DISTRIBUTION. —
<taxonomicName id="9DF04D48FFF3FFACFEF5EE712E8D8E37" box="[292,505,1596,1620]" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="vasiformis">
<emphasis id="6884EAD9FFF3FFACFEF5EE712E8D8E37" box="[292,505,1596,1620]" italics="true" pageId="22" pageNumber="407">Harrisichara vasiformis</emphasis>
</taxonomicName>
form
<taxonomicName id="9DF04D48FFF3FFACFDE1EE712F858E0D" authorityName="Feist-Castel" authorityYear="1977" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="vasiformis-tuberculata">
<emphasis id="6884EAD9FFF3FFACFDE1EE712F858E0D" italics="true" pageId="22" pageNumber="407">vasiformis-tuberculata</emphasis>
</taxonomicName>
occurs in the
<collectingRegion id="9834F829FFF3FFACFEACEE1B2E9E8E0D" box="[381,490,1622,1646]" country="United Kingdom" name="Hampshire" pageId="22" pageNumber="407">Hampshire</collectingRegion>
Basin,
<collectingCountry id="22E7765BFFF3FFACFDE5EE1B2DF28E0D" box="[564,646,1622,1646]" name="United Kingdom" pageId="22" pageNumber="407">England</collectingCountry>
(Feist-Castel 1977b). In
<collectingCountry id="22E7765BFFF3FFACFEB9EE3C2ED88EEB" box="[360,428,1649,1672]" name="France" pageId="22" pageNumber="407">France</collectingCountry>
, it occurs in the Paris (
<bibRefCitation id="3E614B3AFFF3FFACFFA8EEC62F8A8EC0" author="RIVELINE J." box="[121,254,1675,1699]" pageId="22" pageNumber="407" refId="ref18744" refString="RIVELINE J. 1986. - Les Charophytes du Paleogene et du Miocene inferieur d'Europe Occidentale. Cahiers de Paleontologie, Centre National de la Recherche Scientifique, 227 p." type="book" year="1986">Riveline 1986</bibRefCitation>
) and Provence (Feist-Castel 1977a) basins. It has also been reported from Transylvania (
<collectingCountry id="22E7765BFFF3FFACFB86E89A2BC7888C" box="[1111,1203,215,239]" name="Romania" pageId="22" pageNumber="407">Romania</collectingCountry>
) by
<bibRefCitation id="3E614B3AFFF3FFACFD10E8BC2CFB896A" author="BACIU C. &amp; FEIST M." box="[705,911,241,265]" pageId="22" pageNumber="407" pagination="27 - 29" refId="ref16692" refString="BACIU C. &amp; FEIST M. 1999. - Les charophytes oligocenes du nord-ouest de la Transylvanie (Roumanie). Acta Palaeontologica Romana 2: 27 - 29." type="journal article" year="1999">Baciu &amp; Feist (1999)</bibRefCitation>
. In Spain, this morphotype has been documented by Martín-Closas &amp; Ramos (2005) in the Island of Mallorca and by
<bibRefCitation id="3E614B3AFFF3FFACFC1DE96B2B3E895D" author="CHOI S. J." box="[972,1098,294,318]" pageId="22" pageNumber="407" pagination="1 - 67" refId="ref16878" refString="CHOI S. J. 1989. - Les Charophytes du Bassin Potassique Catalan (Nord-Est de l'Espagne) a la limite Eocene-Oligocene. Paleobiologie continentale 26: 1 - 67." type="journal article" year="1989">Choi (1989)</bibRefCitation>
,
<bibRefCitation id="3E614B3AFFF3FFACFB84E96B2D93893A" author="FEIST M. &amp; ANADON P. &amp; CABRERA L. &amp; CHOI S. J. &amp; BO F. &amp; SAEZ M." pageId="22" pageNumber="407" pagination="1 - 32" refId="ref17135" refString="FEIST M., ANADON P., CABRERA L., CHOI S. J., COLOM- BO F. &amp; SAEZ M. 1994. - Upper Eocene-Lowermost Miocene charophyte in the Ebro Basin (Spain). Contribution to the charophyte biozonation in Western Europe. Newsletter Stratigraphy 30: 1 - 32." type="journal article" year="1994">
Feist
<emphasis id="6884EAD9FFF3FFACFB58E96A2BCD895D" box="[1161,1209,294,318]" italics="true" pageId="22" pageNumber="407">et al.</emphasis>
(1994)
</bibRefCitation>
and Sanjuan
<emphasis id="6884EAD9FFF3FFACFCBFE90C2CEA893A" box="[878,926,321,345]" italics="true" pageId="22" pageNumber="407">et al.</emphasis>
(2012) in the eastern part of the Ebro Basin. In the present study, it has been recorded in Sant Boi de Lluçanès, Sobremunt, Sant Bartomeu del Grau and
<taxonomicName id="9DF04D48FFF3FFACFCD1E9DD2C3189CB" authority="Sanjuan &amp; Martín-Closas, 2014" authorityName="Sanjuan &amp; Martín-Closas" authorityYear="2014" box="[768,837,400,424]" pageId="22" pageNumber="389" rank="section" section="Maians" status="section">Maians</taxonomicName>
(
<tableCitation id="17720370FFF3FFACFC9EE9DD2CD589CB" box="[847,929,400,424]" captionStart="TABLE" captionStartId="8.[559,568,219,237]" captionTargetPageId="8" captionText="TABLE 2. — Continuation." httpUri="http://table.plazi.org/id/0E8F6643FFEDFFB2FDFEE8962D8A888E" pageId="22" pageNumber="407" tableUuid="0E8F6643FFEDFFB2FDFEE8962D8A888E">Tables 2</tableCitation>
, 3). Sanjuan &amp; Martín-Closas (2012) showed that in the Ebro Basin, this species grew solely in brackish environments.
</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF3FFACFD74E9B82B408BC8" pageId="22" pageNumber="407" type="description">
<paragraph id="5A4F36CBFFF3FFACFD74E9B82C418A6E" blockId="22.[674,1212,501,940]" box="[677,821,501,527]" pageId="22" pageNumber="407">DESCRIPTION</paragraph>
<paragraph id="5A4F36CBFFF3FFACFD74EA582B408BC8" blockId="22.[674,1212,501,940]" pageId="22" pageNumber="407">
Gyrogonites medium to large,
<quantity id="9D089B2EFFF3FFACFC27EA592BF48A4C" box="[1014,1152,532,559]" metricMagnitude="-1" metricUnit="m" metricValue="6.75" metricValueMax="7.56" metricValueMin="5.9399999999999995" pageId="22" pageNumber="407" unit="mm" value="675.0" valueMax="756.0" valueMin="594.0">594-756 µm</quantity>
high (mean
<quantity id="9D089B2EFFF3FFACFD27EA792C268A2C" box="[758,850,564,591]" metricMagnitude="-1" metricUnit="m" metricValue="6.55" pageId="22" pageNumber="407" unit="mm" value="655.0">655 µm</quantity>
), and
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wide (mean
<quantity id="9D089B2EFFF3FFACFD76EA192C748A0C" box="[679,768,596,623]" metricMagnitude="-1" metricUnit="m" metricValue="6.1899999999999995" pageId="22" pageNumber="407" unit="mm" value="619.0">619 µm</quantity>
), ovoidal or sub-ovoidal in shape with an isopolarity index of 96-116 (mean 106). Nine to eleven (frequently ten) convolutions visible laterally. Spiral cells flat or concave,
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in width. Ŋe ornamentation consists of regularly spaced elongated tubercles, which may be joined by a fine central crest. Apex flat or rounded, generally without any periapical modification in the spiral cells, but with comma-shaped tubercles in a few specimens. Base rounded, conical or elongated in a small column. Basal pore pentagonal,
<quantity id="9D089B2EFFF3FFACFC4EEBDC2C928BC8" box="[927,998,913,940]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="22" pageNumber="407" unit="mm" value="54.0">54 µm</quantity>
across.
</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF3FFACFD74EB9C2B3B8CAB" pageId="22" pageNumber="407" type="discussion">
<paragraph id="5A4F36CBFFF3FFACFD74EB9C2C7D8B8A" blockId="22.[674,1209,977,1224]" box="[677,777,977,1003]" pageId="22" pageNumber="407">REMARKS</paragraph>
<paragraph id="5A4F36CBFFF3FFACFD74EBBD2B3B8CAB" blockId="22.[674,1209,977,1224]" pageId="22" pageNumber="407">
Ŋis taxon was proposed by Feist-Castel (1977) as an informal name to characterise the gradual change from
<taxonomicName id="9DF04D48FFF3FFACFD0EEC7D2C138C2A" box="[735,871,1071,1097]" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="vasiformis">
<emphasis id="6884EAD9FFF3FFACFD0EEC7D2C138C2A" box="[735,871,1071,1097]" italics="true" pageId="22" pageNumber="407">H. vasiformis</emphasis>
</taxonomicName>
to
<taxonomicName id="9DF04D48FFF3FFACFC59EC7D2B698C2A" authorityName="Grambast" authorityYear="1957" baseAuthorityName="Lyell" baseAuthorityYear="1826" box="[904,1053,1071,1097]" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="tuberculata">
<emphasis id="6884EAD9FFF3FFACFC59EC7D2B698C2A" box="[904,1053,1071,1097]" italics="true" pageId="22" pageNumber="407">H. tuberculata</emphasis>
</taxonomicName>
, rather than as a formal species. Later
<bibRefCitation id="3E614B3AFFF3FFACFC4CEC022B298C0A" author="SILLE N. P. &amp; COLLINSON M. E. &amp; KUCERA M. &amp; HOOKER J. J." box="[925,1117,1103,1129]" pageId="22" pageNumber="407" pagination="153 - 173" refId="ref19215" refString="SILLE N. P., COLLINSON M. E., KUCERA M. &amp; HOOKER J. J. 2004. - Evolution within the charophyte genus Harrisichara, late Palaeogene, southern England; environmental and biostratigraphic implications. Palaeogeography, Palaeoecology, Palaeoclimatology 208: 153 - 173." type="journal article" year="2004">
Sille
<emphasis id="6884EAD9FFF3FFACFC05EC1D2B7D8C0A" box="[980,1033,1103,1129]" italics="true" pageId="22" pageNumber="407">et al.</emphasis>
(2004)
</bibRefCitation>
, showed by Eigenshape analysis that these morphotypes are within the variation of
<taxonomicName id="9DF04D48FFF3FFACFC77ECC22B418CCB" box="[934,1077,1166,1192]" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="vasiformis">
<emphasis id="6884EAD9FFF3FFACFC77ECC22B418CCB" box="[934,1077,1166,1192]" italics="true" pageId="22" pageNumber="407">H. vasiformis</emphasis>
</taxonomicName>
rather than being intermediate with
<taxonomicName id="9DF04D48FFF3FFACFC61ECE22B3C8CAB" authorityName="Grambast" authorityYear="1957" baseAuthorityName="Lyell" baseAuthorityYear="1826" box="[944,1096,1198,1224]" class="Charophyceae" family="Characeae" genus="Harrisichara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="22" pageNumber="407" phylum="Charophyta" rank="species" species="tuberculata">
<emphasis id="6884EAD9FFF3FFACFC61ECE22B3C8CAB" box="[944,1096,1198,1224]" italics="true" pageId="22" pageNumber="407">H. tuberculata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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