treatments-xml/data/A3/2A/40/A32A4091456D5D4AB4B6F4DAFE56D085.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1059.70500" ID-GBIF-Dataset="d4f8c0f3-8fa8-4fee-a0ca-0dfd62042eef" ID-PMC="PMC8426311" ID-Pensoft-Pub="1313-2970-1059-23" ID-Pensoft-UUID="1C29C34AB9E65131AF19934F8050B6CC" ID-PubMed="34566445" ID-ZooBank="EA7AA73800D84437BA70940A5060DD89" ModsDocID="1313-2970-1059-23" checkinTime="1630544416668" checkinUser="pensoft" docAuthor="Scheffrahn, Rudolf H." docDate="2021" docId="A32A4091456D5D4AB4B6F4DAFE56D085" docLanguage="en" docName="ZooKeys 1059: 23-33" docOrigin="ZooKeys 1059" docPubDate="2021-09-01" docSource="http://dx.doi.org/10.3897/zookeys.1059.70500" docTitle="Glyptotermes hickmani Scheffrahn 2021, sp. nov." docType="treatment" docUuid="99B790A9-DF50-49E9-9A75-F0D67FB8F3FA" docUuidSource="ZooBank" docVersion="5" id="1C29C34AB9E65131AF19934F8050B6CC" lastPageNumber="23" masterDocId="1C29C34AB9E65131AF19934F8050B6CC" masterDocTitle="The termite genus Glyptotermes (Isoptera, Kalotermitidae) from Paraguay" masterLastPageNumber="33" masterPageNumber="23" pageNumber="23" updateTime="1668150894801" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The termite genus Glyptotermes (Isoptera, Kalotermitidae) from Paraguay</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scheffrahn, Rudolf H.</mods:namePart>
<mods:affiliation>University of Florida, Fort Lauderdale Research &amp; Education Center, 3205 College Avenue, Davie, Florida 33314 USA</mods:affiliation>
<mods:nameIdentifier type="email">rhsc@ufl.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-09-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1059</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>23</mods:start>
<mods:end>33</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1059.70500</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1059.70500</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1059-23</mods:identifier>
<mods:identifier type="ZooBank">EA7AA73800D84437BA70940A5060DD89</mods:identifier>
<mods:identifier type="Pensoft-UUID">1C29C34AB9E65131AF19934F8050B6CC</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="185459893" LSID="urn:lsid:zoobank.org:act:99B790A9-DF50-49E9-9A75-F0D67FB8F3FA" httpUri="http://treatment.plazi.org/id/A32A4091456D5D4AB4B6F4DAFE56D085" lastPageNumber="23" pageId="0" pageNumber="23">
<subSubSection pageId="0" pageNumber="23" type="nomenclature">
<paragraph pageId="0" pageNumber="23">
<taxonomicName LSID="http://zoobank.org/99B790A9-DF50-49E9-9A75-F0D67FB8F3FA" authority="Scheffrahn, 2021" authorityName="Scheffrahn" authorityYear="2021" class="Insecta" family="Kalotermitidae" genus="Glyptotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyptotermes hickmani" order="Blattodea" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="hickmani" status="sp. nov.">Glyptotermes hickmani</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="23">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="description">
<paragraph pageId="0" pageNumber="23">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal (top) and lateral views of the Glyptotermes hickmani sp. nov. female dealate." figureDoi="10.3897/zookeys.1059.70500.figure2" httpUri="https://binary.pensoft.net/fig/584167" pageId="0" pageNumber="23">Figs 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal (top) (B), lateral / oblique, and ventral views of the Glyptotermes hickmani sp. nov. soldier head capsule and pronotum. Arrow = frontal horn." figureDoi="10.3897/zookeys.1059.70500.figure3" httpUri="https://binary.pensoft.net/fig/584168" pageId="0" pageNumber="23">, 3</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live habitus of Glyptotermes hickmani sp. nov. imagos A and soldier B and G. canellae (Mueller) soldier C and imagos D." figureDoi="10.3897/zookeys.1059.70500.figure6" httpUri="https://binary.pensoft.net/fig/584171" pageId="0" pageNumber="23">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="diagnosis">
<paragraph pageId="0" pageNumber="23">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="23">
Among the ten South American 
<taxonomicName class="Insecta" family="Kalotermitidae" genus="Glyptotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyptotermes" order="Blattodea" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="23">Glyptotermes</emphasis>
</taxonomicName>
species for which the soldier is known, the 
<taxonomicName lsidName="G. hickmani" pageId="0" pageNumber="23" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">G. hickmani</emphasis>
</taxonomicName>
sp. nov. soldier is closest in head width to 
<taxonomicName lsidName="G. guianensis" pageId="0" pageNumber="23" rank="species" species="guianensis">
<emphasis italics="true" pageId="0" pageNumber="23">G. guianensis</emphasis>
</taxonomicName>
(Emerson, 1925), but in lateral view, 
<taxonomicName lsidName="G. guianensis" pageId="0" pageNumber="23" rank="species" species="guianensis">
<emphasis italics="true" pageId="0" pageNumber="23">G. guianensis</emphasis>
</taxonomicName>
has angular frontal horns compared to the rounded frontal horns of 
<taxonomicName lsidName="G. hickmani" pageId="0" pageNumber="23" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">G. hickmani</emphasis>
</taxonomicName>
. The frontal horns of the 
<taxonomicName lsidName="G. hickmani" pageId="0" pageNumber="23" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">G. hickmani</emphasis>
</taxonomicName>
soldier are diagnostic.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="type locality">
<paragraph pageId="0" pageNumber="23">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="23">Paraguay, Nueva Colombia.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="materials_examined">
<paragraph pageId="0" pageNumber="23">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="23">
<materialsCitation collectingDate="2012-06-05" collectorName="R. Scheffrahn, J. Chase, R. Hickman, J. Krecek, J. Mangold, A. Mullins, Davie Florida" country="Paraguay" elevation="115" latitude="-25.1747" location="Nueva" longLatPrecision="7" longitude="-57.2876" municipality="University of Florida Termite Collection" specimenCount="1" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">
<typeStatus>Holotype</typeStatus>
</emphasis>
soldier: 
<collectingCountry name="Paraguay">Paraguay</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:A32A4091456D5D4AB4B6F4DAFE56D085:19ED69EDA5AF1F053E7E1CE219BA7F8C" country="Paraguay" latitude="-25.1747" longLatPrecision="7" longitude="-57.2876" municipality="University of Florida Termite Collection" name="Nueva">Nueva</location>
<collectingCountry name="Colombia">Colombia</collectingCountry>
(
<geoCoordinate degrees="25.1747" direction="south" orientation="latitude" precision="5" value="-25.1747">-25.1747</geoCoordinate>
, 
<geoCoordinate degrees="57.2876" direction="west" orientation="longitude" precision="5" value="-57.2876">-57.2876</geoCoordinate>
), elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">115 m</elevation>
</quantity>
, 
<collectingDate value="2012-06-05">5JUN2012</collectingDate>
, 
<collectorName>R. Scheffrahn</collectorName>
et al. (
<collectorName>R. Scheffrahn</collectorName>
, 
<collectorName>J. Chase</collectorName>
, 
<collectorName>R. Hickman</collectorName>
, 
<collectorName>
J. 
<normalizedToken originalValue="Křeček">Krecek</normalizedToken>
</collectorName>
, 
<collectorName>J. Mangold</collectorName>
, 
<collectorName>A. Mullins</collectorName>
), 
<collectingMunicipality>University of Florida Termite Collection</collectingMunicipality>
(UFTC), 
<collectorName>Davie Florida</collectorName>
, no. PA1265
</materialsCitation>
. 
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="R. Scheffrahn &amp; J. Chase &amp; R. Hickman &amp; J. Krecek &amp; J. Mangold &amp; A. Mullins &amp; Davie Florida" country="Paraguay" elevation="115" latitude="-25.1747" location="Nueva" longLatPrecision="7" longitude="-57.2876" municipality="University of Florida Termite Collection" specimenCount="one" specimenCount-queen="2" specimenCount-soldier="1" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="23">
<emphasis italics="true" pageId="0" pageNumber="23">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
Five additional soldiers, 
<specimenCount type="queen">one queen</specimenCount>
, pseudergates/nymphs, same colony sample as 
<typeStatus>holotype</typeStatus>
. One additional colony from type locality (same data), 
<specimenCount type="soldier">four soldiers</specimenCount>
, one king, 
<specimenCount type="queen">one queen</specimenCount>
, and pseudergates/nymphs (PA1264)
</materialsCitation>
. 
<materialsCitation collectingDate="2012-05-29" collectorName="R. Scheffrahn" country="Paraguay" elevation="293" latitude="-22.6788" location="Cerra Cora" longLatPrecision="7" longitude="-55.995" specimenCount="two" specimenCount-soldier="1" typeStatus="paratype">
<collectingCountry name="Paraguay">Paraguay</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:A32A4091456D5D4AB4B6F4DAFE56D085:80ACB37036531A1FD1D37D1DF17B9ECB" country="Paraguay" latitude="-22.6788" longLatPrecision="7" longitude="-55.995" name="Cerra Cora">Cerra Cora</location>
(
<geoCoordinate degrees="22.6788" direction="south" orientation="latitude" precision="5" value="-22.6788">-22.6788</geoCoordinate>
, 
<geoCoordinate degrees="55.9950" direction="west" orientation="longitude" precision="5" value="-55.995">-55.9950</geoCoordinate>
), elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.93" unit="m" value="293.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.93" unit="m" value="293.0">293 m</elevation>
</quantity>
, 
<collectingDate value="2012-05-29">29MAY2012</collectingDate>
, 
<collectorName>R. Scheffrahn</collectorName>
et al., UFTC no. PA326 containing 
<specimenCount type="soldier">two soldiers</specimenCount>
and pseudergates/nymphs
</materialsCitation>
. 
<materialsCitation collectingDate="2012-05-29" collectorName="R. Scheffrahn" country="Paraguay" elevation="328" latitude="-22.56" location="Pedro Juan Caballero" longLatPrecision="7" longitude="-56.3006" specimenCount="one" specimenCount-soldier="1" typeStatus="paratype">
<collectingCountry name="Paraguay">Paraguay</collectingCountry>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.8" unit="km" value="58.0">58 km</quantity>
W of 
<location LSID="urn:lsid:plazi:treatment:A32A4091456D5D4AB4B6F4DAFE56D085:833201BD476B903420E8713102705684" country="Paraguay" latitude="-22.56" longLatPrecision="7" longitude="-56.3006" name="Pedro Juan Caballero">Pedro Juan Caballero</location>
(
<geoCoordinate degrees="22.5600" direction="south" orientation="latitude" precision="5" value="-22.56">-22.5600</geoCoordinate>
, 
<geoCoordinate degrees="56.3006" direction="west" orientation="longitude" precision="5" value="-56.3006">-56.3006</geoCoordinate>
), elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.28" unit="m" value="328.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.28" unit="m" value="328.0">328 m</elevation>
</quantity>
, 
<collectingDate value="2012-05-29">29MAY2012</collectingDate>
, 
<collectorName>R. Scheffrahn</collectorName>
et al., UFTC no. PA446 containing 
<specimenCount type="soldier">one soldier</specimenCount>
and 12 pseudergates/nymphs
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="description">
<paragraph pageId="0" pageNumber="23">Description.</paragraph>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">Dealated Imago</emphasis>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal (top) and lateral views of the Glyptotermes hickmani sp. nov. female dealate." figureDoi="10.3897/zookeys.1059.70500.figure2" httpUri="https://binary.pensoft.net/fig/584167" pageId="0" pageNumber="23">2</figureCitation>
, Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (mm) of Glyptotermes hickmani sp. nov. and G. canellae (Mueller) imagos." httpUri="http://table.plazi.org/id/9A3A1945C477EFF524F1AA0899B5F50D" pageId="0" pageNumber="23" tableUuid="9A3A1945C477EFF524F1AA0899B5F50D">1</tableCitation>
). Head and pronotum castaneus brown (almost black in live habitus, Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live habitus of Glyptotermes hickmani sp. nov. imagos A and soldier B and G. canellae (Mueller) soldier C and imagos D." figureDoi="10.3897/zookeys.1059.70500.figure6" httpUri="https://binary.pensoft.net/fig/584171" pageId="0" pageNumber="23">6A</figureCitation>
). Postclypeus hyaline; labrum light orange-brown. Fore wing scales concolorous with pronotum. Femora slightly lighter than pronotum; tibiae concolorous with labrum. In lateral view, vertex of head with 20-24 scattered setae of medium length. In dorsal view, pronotum with about 20 setae of variable length along each lateral margin; in lateral view, with setae in line with anterior and posterior margins; a few in middle. Eyes small, dark gray, occupying less than one-third the distance between vertex and genal margins; ellipsoid with rectate margin at antennal socket. Ocelli small, orange, elliptical; separated from eye by their narrow width. Antennae article formula 2 &lt;3&gt; 4 = 5. Pronotum wider than long, about as wide as head capsule without eyes; anterior and posterior margins with slight concavity, sides slightly convex. Arolia present.
</paragraph>
<caption ID-Table-UUID="9A3A1945C477EFF524F1AA0899B5F50D" httpUri="http://table.plazi.org/id/9A3A1945C477EFF524F1AA0899B5F50D" pageId="0" pageNumber="23" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Table 1.</emphasis>
Measurements (mm) of 
<taxonomicName authorityName="Scheffrahn" authorityYear="2021" class="Insecta" family="Kalotermitidae" genus="Glyptotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyptotermes hickmani" order="Blattodea" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">Glyptotermes hickmani</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="G. canellae" pageId="0" pageNumber="23" rank="species" species="canellae">
<emphasis italics="true" pageId="0" pageNumber="23">G. canellae</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
) imagos.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="23">
<table pageId="0" pageNumber="23">
<tr pageId="0" pageNumber="23">
<th colspan="1" pageId="0" pageNumber="23" rowspan="3">Characters</th>
<th colspan="2" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName lsidName="G. hickmani" pageId="0" pageNumber="23" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">G. hickmani</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName lsidName="G. canellae" pageId="0" pageNumber="23" rank="species" species="canellae">
<emphasis italics="true" pageId="0" pageNumber="23">G. canellae</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="23">
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">(n = 4)</th>
<th colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">(n = 12)</th>
<th colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</th>
</tr>
<tr pageId="0" pageNumber="23">
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">mean</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">range</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">mean</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">range</th>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of head to tip of labrum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.35-1.44</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.30-1.40</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of head to side base of mandibles</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.64</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.54-0.79</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.64</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.58-0.79</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Width of head</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.11-1.19</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.14-1.19</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Diameter of eye</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.26</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.25-0.30</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.27</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.25-0.28</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Eye from lower margin</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.27</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.23-0.33</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.18</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.16-0.25</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of ocellus</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.1</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.09-0.11</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.1</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.09-0.11</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Median length of pronotum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.62</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.58-0.65</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.57</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.53-0.60</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Maximum length of pronotum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.71</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.70-0.74</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.65-0.77</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Width of pronotum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.09-1.26</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.06</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.00-1.11</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of hind tibia</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.91</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.86-0.96</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.87</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.79-0.91</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of fore wing from suture</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">6.57</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">6.24-6.80</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Width of fore wing</td>
<td colspan="2" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.83</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.68-1.93</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zookeys.1059.70500.figure2" httpUri="https://binary.pensoft.net/fig/584167" pageId="0" pageNumber="23" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Figure 2.</emphasis>
Dorsal (top) and lateral views of the 
<taxonomicName authorityName="Scheffrahn" authorityYear="2021" class="Insecta" family="Kalotermitidae" genus="Glyptotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyptotermes hickmani" order="Blattodea" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">Glyptotermes hickmani</emphasis>
</taxonomicName>
sp. nov. female dealate.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">Soldier</emphasis>
(Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal (top) (B), lateral / oblique, and ventral views of the Glyptotermes hickmani sp. nov. soldier head capsule and pronotum. Arrow = frontal horn." figureDoi="10.3897/zookeys.1059.70500.figure3" httpUri="https://binary.pensoft.net/fig/584168" pageId="0" pageNumber="23">3</figureCitation>
, Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements (mm) of Glyptotermes hickmani sp. nov. and G. canellae (Mueller) soldiers." httpUri="http://table.plazi.org/id/0AB0B6FCE002E515112F302A2405A06F" pageId="0" pageNumber="23" tableUuid="0AB0B6FCE002E515112F302A2405A06F">2</tableCitation>
). Head, in lateral view, grading from light orange at the cervical margin to dark brown at frontal horns (darker in live habitus, Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live habitus of Glyptotermes hickmani sp. nov. imagos A and soldier B and G. canellae (Mueller) soldier C and imagos D." figureDoi="10.3897/zookeys.1059.70500.figure6" httpUri="https://binary.pensoft.net/fig/584171" pageId="0" pageNumber="23">6B</figureCitation>
). Mandibles concolorous with frons. Head capsule nearly cuboid; about as wide as high; with distinct narrowing near anterior fourth. Pronotum more than twice as wide as long, widest near front third; anterior margin weakly concave, posterior margin barely concave in middle. In dorsal and ventral views, setae on head and pronotum medium to long; about 4-5 short setae on basal hump of mandibles. In lateral view, setae less numerous except for setae on postmentum. Mandibles robust, form about two-thirds of total head length; curve apically near anterior half, humps at basal fourth evenly rounded beyond outer blades. Dentition as in Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal (top) (B), lateral / oblique, and ventral views of the Glyptotermes hickmani sp. nov. soldier head capsule and pronotum. Arrow = frontal horn." figureDoi="10.3897/zookeys.1059.70500.figure3" httpUri="https://binary.pensoft.net/fig/584168" pageId="0" pageNumber="23">3</figureCitation>
. Frons bilobed; lobes rounded in lateral view; frontal furrow between lobes extends to postclypeus. Angle formed by vertex and frons about 75°; lateral view of frons punctuated by large, knob-like, frontal horns. Eye spots very faint. In ventral view, posterior margin of postmentum slightly wider than middle. Genal horns rounded from above, project slightly beyond narrowest point of head capsule. Antenna with 11-12 articles; formula 2&gt; 3 &lt;4 &lt;5. Femora swollen.
</paragraph>
<caption ID-Table-UUID="0AB0B6FCE002E515112F302A2405A06F" httpUri="http://table.plazi.org/id/0AB0B6FCE002E515112F302A2405A06F" pageId="0" pageNumber="23" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Table 2.</emphasis>
Measurements (mm) of 
<taxonomicName authorityName="Scheffrahn" authorityYear="2021" class="Insecta" family="Kalotermitidae" genus="Glyptotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyptotermes hickmani" order="Blattodea" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">Glyptotermes hickmani</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="G. canellae" pageId="0" pageNumber="23" rank="species" species="canellae">
<emphasis italics="true" pageId="0" pageNumber="23">G. canellae</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
) soldiers.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="23">
<table pageId="0" pageNumber="23">
<tr pageId="0" pageNumber="23">
<th colspan="1" pageId="0" pageNumber="23" rowspan="3">Characters</th>
<th colspan="2" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName lsidName="G. hickmani" pageId="0" pageNumber="23" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">G. hickmani</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName lsidName="G. canellae" pageId="0" pageNumber="23" rank="species" species="canellae">
<emphasis italics="true" pageId="0" pageNumber="23">G. canellae</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="23">
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">(n = 9)</th>
<th colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">(n = 2)</th>
<th colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</th>
</tr>
<tr pageId="0" pageNumber="23">
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">mean</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">range</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">mean</th>
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">range</th>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of head to side base of mandibles</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.44</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.09-1.70</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.84</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.81-1.88</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Width of head</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.22</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.00-1.32</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.37</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.32-1.42</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Height of head</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.11</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.95-1.18</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.19-1.19</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of left mandible</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.04</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.88-1.14</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.18-1.23</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Maximum width of postmentum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.38</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.35-0.40</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.51</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.49-0.53</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Minimum width of postmentum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.29</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.26-0.33</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.37</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.35-0.39</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of postmentum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.12</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.89-1.25</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.12</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.12-1.12</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Median length of pronotum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.54</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.40-0.61</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.59</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.58-0.60</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Maximum length of pronotum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.65</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.51-0.74</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.73</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.72-0.74</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Width of pronotum</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.91-1.32</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.26</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1.23-1.30</td>
</tr>
<tr pageId="0" pageNumber="23">
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Length of hind tibia</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.72</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.61-0.79</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">0.88-0.88</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zookeys.1059.70500.figure3" httpUri="https://binary.pensoft.net/fig/584168" pageId="0" pageNumber="23" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Figure 3.</emphasis>
Dorsal (top) (
<emphasis bold="true" pageId="0" pageNumber="23">B</emphasis>
), lateral/oblique, and ventral views of the 
<taxonomicName authorityName="Scheffrahn" authorityYear="2021" class="Insecta" family="Kalotermitidae" genus="Glyptotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyptotermes hickmani" order="Blattodea" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="hickmani">
<emphasis italics="true" pageId="0" pageNumber="23">Glyptotermes hickmani</emphasis>
</taxonomicName>
sp. nov. soldier head capsule and pronotum. Arrow = frontal horn.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="distribution">
<paragraph pageId="0" pageNumber="23">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="23">
See Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Glyptotermes hickmani sp. nov. and G. canellae (Mueller) in Paraguay and literature localities of G. canellae in Argentina and Brazil." figureDoi="10.3897/zookeys.1059.70500.figure1" httpUri="https://binary.pensoft.net/fig/584166" pageId="0" pageNumber="23">1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="etymology">
<paragraph pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="23">Named after Robert B. (Bob) Hickman, a friend and able termite collector, who joined us on the Paraguay expedition.</paragraph>
</subSubSection>
</treatment>
</document>