151 lines
17 KiB
XML
151 lines
17 KiB
XML
<document id="9CB8576ED3037FBFEC90A64CE35CA4F3" ID-DOI="10.11646/zootaxa.3919.3.5" ID-GBIF-Dataset="12e779cb-148a-4853-ac3f-affef036a340" ID-ISSN="1175-5326" ID-Zenodo-Dep="234051" ID-ZooBank="743AF37E-54B4-4BCB-A3E8-93092F779A20" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460993193249" checkinUser="plazi" docAuthor="Blake, James A." docDate="2015" docId="9A3E87FEEB7EFFFFD2F7FA85FC58FF48" docLanguage="en" docName="zt03919p552.pdf" docOrigin="Zootaxa 3919 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tharyx Webster and Benedict 1887" docType="treatment" docVersion="9" lastPageNumber="533" masterDocId="6607FF86EB61FFDFD260FFB6FFC0FFB0" masterDocTitle="New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic" masterLastPageNumber="552" masterPageNumber="501" pageNumber="532" updateTime="1698562195018" updateUser="plazi">
|
|
<mods:mods id="EF4D4E8A88C67BBD6CF98F7E8CFEF395" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="BA319EEFC9062354044ABC7A25F8FDEC">
|
|
<mods:title id="A7376D9F55C14EECB9613841F70152F4">New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="53876F3042B5257F6EFCBDA80F5D0631" type="personal">
|
|
<mods:role id="36B9E570E92AE155FE834E2B529BD0F0">
|
|
<mods:roleTerm id="9ABD6D4C62277A68DA0BF35FE18C484B">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="6B29EF5DA6CBD4BB7CD38197FFB1373E">Blake, James A.</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="0DE02FADF3DBE4F9D0C2C073A7BC0467">text</mods:typeOfResource>
|
|
<mods:relatedItem id="0A3FABAEF2995A2436B7745DFB11AB3E" type="host">
|
|
<mods:titleInfo id="FF35AF737687BF3E0EB26235994E38D6">
|
|
<mods:title id="4BEBCD9154AEB416741B8CB5B636EC81">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="D9F12F4DF67AD0C161BF994275A714D9">
|
|
<mods:date id="AB1584FBCF58035798398039AE628216">2015</mods:date>
|
|
<mods:detail id="284DB8A7C997BBF470E66AF7B5D0EEAC" type="volume">
|
|
<mods:number id="B25BE773143D07353E9BE810400F0F96">3919</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="A020904778DBCB90FCB66C91B0F015D6" type="issue">
|
|
<mods:number id="A9BA58B98589BA4ED44B6F3773B04AF9">3</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="BAE44348648629076E7A48E810981ABF" unit="page">
|
|
<mods:start id="4A34B4573043F4E022396BF2C0CA4C28">501</mods:start>
|
|
<mods:end id="9E312D7DEFC5C4F2CB1A71CE31E76487">552</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="DC6039172CA07D967DD7F563045D2F5F">journal article</mods:classification>
|
|
<mods:identifier id="59AF9A3641007A815EB22E1A507A2E27" type="DOI">10.11646/zootaxa.3919.3.5</mods:identifier>
|
|
<mods:identifier id="66F87753117D31E294E8C63C27A14344" type="GBIF-Dataset">12e779cb-148a-4853-ac3f-affef036a340</mods:identifier>
|
|
<mods:identifier id="BF74E065E3B436F9B5D67551983743C9" type="ISSN">1175-5326</mods:identifier>
|
|
<mods:identifier id="D253C33767E02ED4097F7DA0341F3BCE" type="Zenodo-Dep">234051</mods:identifier>
|
|
<mods:identifier id="5BE11B31D8E32E2AB99349F07AEB1975" type="ZooBank">743AF37E-54B4-4BCB-A3E8-93092F779A20</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="9A3E87FEEB7EFFFFD2F7FA85FC58FF48" ID-DOI="http://doi.org/10.5281/zenodo.5664948" ID-GBIF-Taxon="119596878" ID-Zenodo-Dep="5664948" LSID="urn:lsid:plazi:treatment:9A3E87FEEB7EFFFFD2F7FA85FC58FF48" httpUri="http://treatment.plazi.org/id/9A3E87FEEB7EFFFFD2F7FA85FC58FF48" lastPageId="32" lastPageNumber="533" pageId="31" pageNumber="532">
|
|
<subSubSection id="5A8D6563EB7EFFC0D2F7FA85FD6AFAFD" box="[151,682,1331,1357]" pageId="31" pageNumber="532" type="nomenclature">
|
|
<paragraph id="122836E8EB7EFFC0D2F7FA85FD6AFAFD" blockId="31.[151,682,1331,1357]" box="[151,682,1331,1357]" pageId="31" pageNumber="532">
|
|
<heading id="49608184EB7EFFC0D2F7FA85FD6AFAFD" bold="true" box="[151,682,1331,1357]" fontSize="11" level="1" pageId="31" pageNumber="532" reason="1">
|
|
<emphasis id="20E3EAFAEB7EFFC0D2F7FA85FD6AFAFD" bold="true" box="[151,682,1331,1357]" pageId="31" pageNumber="532">
|
|
Genus
|
|
<taxonomicName id="D5974D6BEB7EFFC0D28EFA85FD6AFAFD" ID-CoL="84V7M" authority="Webster and Benedict, 1887" authorityName="Webster and Benedict" authorityYear="1887" box="[238,682,1331,1357]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D28EFA85FE86FAFD" bold="true" box="[238,326,1331,1357]" italics="true" pageId="31" pageNumber="532">Tharyx</emphasis>
|
|
Webster and Benedict, 1887
|
|
</taxonomicName>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5A8D6563EB7EFFC0D2F7FACCFCA5FA20" box="[151,869,1402,1424]" pageId="31" pageNumber="532" type="materials_examined">
|
|
<paragraph id="122836E8EB7EFFC0D2F7FACCFCA5FA20" blockId="31.[151,869,1402,1424]" box="[151,869,1402,1424]" pageId="31" pageNumber="532">
|
|
<emphasis id="20E3EAFAEB7EFFC0D2F7FACCFEDCFA20" bold="true" box="[151,284,1402,1424]" pageId="31" pageNumber="532">
|
|
<typeStatus id="CD2C884AEB7EFFC0D2F7FACCFF0CFA20" box="[151,204,1402,1424]" pageId="31" pageNumber="532">Type</typeStatus>
|
|
species
|
|
</emphasis>
|
|
:
|
|
<taxonomicName id="D5974D6BEB7EFFC0D34AFACCFD0EFA20" authority="Webster & Benedict, 1887" authorityName="Webster & Benedict" authorityYear="1887" box="[298,718,1402,1424]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="species" species="acutus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D34AFACCFE7AFA20" box="[298,442,1402,1424]" italics="true" pageId="31" pageNumber="532">Tharyx acutus</emphasis>
|
|
Webster & Benedict, 1887
|
|
</taxonomicName>
|
|
, by monotypy.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5A8D6563EB7EFFC0D2F7FA0AFA57F91E" pageId="31" pageNumber="532" type="diagnosis">
|
|
<paragraph id="122836E8EB7EFFC0D2F7FA0AFA57F91E" blockId="31.[151,1436,1468,2033]" pageId="31" pageNumber="532">
|
|
<emphasis id="20E3EAFAEB7EFFC0D2F7FA0AFECAFA65" bold="true" box="[151,266,1468,1493]" pageId="31" pageNumber="532">Diagnosis</emphasis>
|
|
. Prostomium conical; peristomium elongate, with pair of grooved dorsal tentacles arising on posterior margin anterior to setiger 1; first pair of branchiae typically arising immediately posterior to dorsal tentacles either on posterior margin of peristomium or on setiger 1; abdominal segments sometimes beadlike. Noto- and neurosetae arising close to one another, not widely separated. Setae include simple capillaries in anterior and middle setigers, acicular spines in posterior setigers with irregular notched tips, sometimes appearing more or less bidentate, with pair of stunted or rounded knobs but never with distinct, sharply pointed teeth; spines present either in both posterior noto- and neuropodia or only in neuropodia. Pygidium with terminal anus and small ventral lobe or disk.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5A8D6563EB7EFFFFD2A7F90EFC58FF48" lastPageId="32" lastPageNumber="533" pageId="31" pageNumber="532" type="discussion">
|
|
<paragraph id="122836E8EB7EFFC0D2A7F90EFC1BF841" blockId="31.[151,1436,1468,2033]" pageId="31" pageNumber="532">
|
|
<emphasis id="20E3EAFAEB7EFFC0D2A7F90EFEFBF961" bold="true" box="[199,315,1720,1745]" pageId="31" pageNumber="532">Remarks.</emphasis>
|
|
<bibRefCitation id="76064B19EB7EFFC0D323F90EFE17F961" author="Blake" box="[323,471,1720,1745]" pageId="31" pageNumber="532" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. & J. B. Kirkegaard (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement No. 5, 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
|
|
determined that the
|
|
<typeStatus id="CD2C884AEB7EFFC0D0A2F90FFD32F961" box="[706,754,1721,1745]" pageId="31" pageNumber="532">type</typeStatus>
|
|
species of
|
|
<taxonomicName id="D5974D6BEB7EFFC0D113F90EFC02F961" box="[883,962,1720,1745]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D113F90EFC02F961" box="[883,962,1720,1745]" italics="true" pageId="31" pageNumber="532">Tharyx</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="D5974D6BEB7EFFC0D1B1F90CFAABF961" authority="Webster & Benedict, 1887" authorityName="Webster & Benedict" authorityYear="1887" box="[977,1387,1720,1745]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="species" species="acutus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D1B1F90CFBF7F961" box="[977,1079,1722,1745]" italics="true" pageId="31" pageNumber="532">T. acutus</emphasis>
|
|
Webster & Benedict, 1887
|
|
</taxonomicName>
|
|
had knob-tipped acicular spines in addition to capillaries. At the time,
|
|
<taxonomicName id="D5974D6BEB7EFFC0D1FDF96BFC2CF946" box="[925,1004,1757,1782]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D1FDF96BFC2CF946" box="[925,1004,1757,1782]" italics="true" pageId="31" pageNumber="532">Tharyx</emphasis>
|
|
</taxonomicName>
|
|
species were defined as having only capillaries (
|
|
<bibRefCitation id="76064B19EB7EFFC0D37BF8B7FE04F8A9" author="Hartman" box="[283,452,1792,1817]" pageId="31" pageNumber="532" refString="Hartman, O. (1961) Polychaetous annelids from California. Allan Hancock Pacific Expeditions, 25, 1 - 226, 34 plates." type="book chapter" year="1961">Hartman 1961</bibRefCitation>
|
|
).
|
|
<bibRefCitation id="76064B19EB7EFFC0D3BAF8B6FDAFF8A9" author="Blake" box="[474,623,1792,1817]" pageId="31" pageNumber="532" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. & J. B. Kirkegaard (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement No. 5, 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
|
|
limited
|
|
<taxonomicName id="D5974D6BEB7EFFC0D0B1F8B6FCE1F8A9" box="[721,801,1792,1817]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D0B1F8B6FCE1F8A9" box="[721,801,1792,1817]" italics="true" pageId="31" pageNumber="532">Tharyx</emphasis>
|
|
</taxonomicName>
|
|
to those species having knob-tipped spines and moved species having only simple or serrated capillaries to the genera
|
|
<taxonomicName id="D5974D6BEB7EFFC0D1F7F893FB0BF88D" authority="Blake, 1991" authorityName="Blake" authorityYear="1991" box="[919,1227,1829,1854]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D1F7F893FBF1F88E" box="[919,1073,1829,1854]" italics="true" pageId="31" pageNumber="532">Aphelochaeta</emphasis>
|
|
<bibRefCitation id="76064B19EB7EFFC0D65DF893FB0BF88D" author="Blake" box="[1085,1227,1829,1854]" pageId="31" pageNumber="532" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. & J. B. Kirkegaard (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement No. 5, 17 - 30." type="book chapter" year="1991">Blake, 1991</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="D5974D6BEB7EFFC0D76DF893FEF3F8D1" authority="Laubier, 1961" authorityName="Laubier" authorityYear="1961" class="Polychaeta" family="Cirratulidae" genus="Monticellina" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D76DF893FA5CF88E" box="[1293,1436,1829,1854]" italics="true" pageId="31" pageNumber="532">Monticellina</emphasis>
|
|
Laubier, 1961
|
|
</taxonomicName>
|
|
respectively. The genus
|
|
<taxonomicName id="D5974D6BEB7EFFC0D027F8FEFD56F8D1" box="[583,662,1864,1889]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D027F8FEFD56F8D1" box="[583,662,1864,1889]" italics="true" pageId="31" pageNumber="532">Tharyx</emphasis>
|
|
</taxonomicName>
|
|
superficially appears most closely related to
|
|
<taxonomicName id="D5974D6BEB7EFFC0D6EFF8FEFAD6F8D1" box="[1167,1302,1864,1889]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D6EFF8FEFAD6F8D1" box="[1167,1302,1864,1889]" italics="true" pageId="31" pageNumber="532">Caulleriella</emphasis>
|
|
</taxonomicName>
|
|
in that both genera have species with modified spines that are more or less bidentate.
|
|
<taxonomicName id="D5974D6BEB7EFFC0D669F8DBFB50F836" box="[1033,1168,1901,1926]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D669F8DBFB50F836" box="[1033,1168,1901,1926]" italics="true" pageId="31" pageNumber="532">Caulleriella</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="D5974D6BEB7EFFC0D6B1F8DBFAE1F836" box="[1233,1313,1901,1926]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D6B1F8DBFAE1F836" box="[1233,1313,1901,1926]" italics="true" pageId="31" pageNumber="532">Tharyx</emphasis>
|
|
</taxonomicName>
|
|
are easily distinguished, however, by the position of the noto- and neuropodia. In
|
|
<taxonomicName id="D5974D6BEB7EFFC0D1A2F826FB89F819" box="[962,1097,1936,1961]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB7EFFC0D1A2F826FB89F819" box="[962,1097,1936,1961]" italics="true" pageId="31" pageNumber="532">Caulleriella</emphasis>
|
|
</taxonomicName>
|
|
, the setal fascicles are widely separated from one another, so much so that in cross section of some species, they appear to be positioned at four corners. In
|
|
<emphasis id="20E3EAFAEB7EFFC0D375F86EFEACF841" box="[277,364,2008,2033]" italics="true" pageId="31" pageNumber="532">
|
|
<taxonomicName id="D5974D6BEB7EFFC0D375F86EFEA7F841" box="[277,359,2008,2033]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="532" phylum="Annelida" rank="genus">Tharyx</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
on the other hand, the setal fascicles are close together.
|
|
</paragraph>
|
|
<paragraph id="122836E8EB41FFFFD2A7FF2EFC58FF48" blockId="32.[151,1436,151,248]" pageId="32" pageNumber="533">
|
|
Two new species of T
|
|
<emphasis id="20E3EAFAEB41FFFFD3A1FF21FDC1FF00" box="[449,513,151,176]" italics="true" pageId="32" pageNumber="533">haryx</emphasis>
|
|
are described as part of this study. One species from Alaska had been previously referred to the genus
|
|
<taxonomicName id="D5974D6BEB41FFFFD3EEFF0AFDD2FF65" box="[398,530,188,213]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="32" pageNumber="533" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB41FFFFD3EEFF0AFDD2FF65" box="[398,530,188,213]" italics="true" pageId="32" pageNumber="533">Chaetozone</emphasis>
|
|
</taxonomicName>
|
|
. The second species, from the Puget Sound, Washington, was known in Puget Sound monitoring programs under a provisional
|
|
<taxonomicName id="D5974D6BEB41FFFFD0D9FF69FCC9FF48" box="[697,777,223,248]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="32" pageNumber="533" phylum="Annelida" rank="genus">
|
|
<emphasis id="20E3EAFAEB41FFFFD0D9FF69FCC9FF48" box="[697,777,223,248]" italics="true" pageId="32" pageNumber="533">Tharyx</emphasis>
|
|
</taxonomicName>
|
|
designation.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |