treatments-xml/data/8C/0B/43/8C0B432BB6C9510386E5CA951D570889.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="8C0B432BB6C9510386E5CA951D570889" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus distinctus" docType="treatment" docVersion="3" lastPageNumber="182" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="181" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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<mods:date>2013</mods:date>
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<mods:number>271</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<treatment ID-GBIF-Taxon="152040875" LSID="urn:lsid:plazi:treatment:8C0B432BB6C9510386E5CA951D570889" httpUri="http://treatment.plazi.org/id/8C0B432BB6C9510386E5CA951D570889" lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181">
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<paragraph pageId="180" pageNumber="181">
<taxonomicName LSID="http://species-id.net/wiki/Operclipygus_distinctus" authority="Hinton 1935" authorityYear="1935" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus distinctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="distinctus">Operclipygus distinctus (Hinton 1935)</taxonomicName>
Figs 48L49 
<normalizedToken originalValue="A–BMap">A-BMap</normalizedToken>
17
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="reference_group">
<paragraph pageId="180" pageNumber="181">
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister distinctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="distinctus">Phelister distinctus</taxonomicName>
Hinton, 1935a: 589; 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus distinctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="distinctus">Operclipygus distinctus</taxonomicName>
: 
<bibRefCitation author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel 1976</bibRefCitation>
: 257.
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="type locality">
<paragraph pageId="180" pageNumber="181">Type locality.</paragraph>
<paragraph pageId="180" pageNumber="181">
BRAZIL: 
<normalizedToken originalValue="Pará">Para</normalizedToken>
: 
<normalizedToken originalValue="Santarém">Santarem</normalizedToken>
[
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-2.4333334">2°26'S</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-54.7">54°42'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="type material">
<paragraph pageId="180" pageNumber="181">Type material.</paragraph>
<paragraph pageId="180" pageNumber="181">
Lectotype, here designated: &quot;H.H. Smith&quot; / &quot;Santarem&quot; / &quot; 
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister distinctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="distinctus">Phelister distinctus</taxonomicName>
Hntn.&quot; / &quot;LECTOTYPE 
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister distinctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="distinctus">Phelister distinctus</taxonomicName>
Hinton M.S.Caterino &amp; A.K.Tishechkin des. 2010&quot; (BMNH). This species was described from an unspecified number of specimens, and the lectotype designation fixes primary type status on the only known original specimen.
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="other material">
<paragraph pageId="180" pageNumber="181">Other material.</paragraph>
<paragraph pageId="180" pageNumber="181">
BRAZIL: 
<normalizedToken originalValue="Pará">Para</normalizedToken>
: 1:IPEAN, Utinga, 
<normalizedToken originalValue="Belém">Belem</normalizedToken>
, 
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.45">1°27'S</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.433334">48°26'W</geoCoordinate>
, v.1985, FIT (CHND), 1: viii.1985, FIT (CHND), 1: xi.1984, FIT (FMNH); 1: Barcarena, 
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.5">1°30'S</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.616665">48°37'W</geoCoordinate>
, 13-25.vi.1991, FIT (MSCC); 1: 
<normalizedToken originalValue="Tucuruí">Tucurui</normalizedToken>
, 
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.75">3°45'S</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-49.666668">49°40'W</geoCoordinate>
, 23. 
<normalizedToken originalValue="vi–">vi-</normalizedToken>
7.vii.1986, FIT (AKTC). ECUADOR: Orellana: 1: 
<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
Res. Stn., 
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.67444444">00°40'28&quot;S</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-76.64722">76°38'50&quot;W</geoCoordinate>
, 215m, 5-10.ix.1999, FIT, primary forest, E.G. Riley (TAMU). FRENCH GUIANA: 1: 
<normalizedToken originalValue="Rés">Res</normalizedToken>
. 
<normalizedToken originalValue="Trésor">Tresor</normalizedToken>
, route de Kaw Pk18, 
<geoCoordinate direction="north" orientation="latitude" precision="9" value="4.6105">4°36.63'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-52.279">52°16.74'W</geoCoordinate>
, 225m, 6.xi.2009, FIT, SEAG (MNHN); 1: Mont. Tabulaire, 
<normalizedToken originalValue="Itoupé">Itoupe</normalizedToken>
, 
<geoCoordinate direction="north" orientation="latitude" precision="9" value="3.022">3°1.32'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-53.084167">53°5.05'W</geoCoordinate>
, 800m, 17.iii.2010, FIT, SEAG (CHND); 1: 
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
des Nouragues, 
<normalizedToken originalValue="Régina">Regina</normalizedToken>
, 
<geoCoordinate direction="north" orientation="latitude" precision="9" value="4.037833">4°2.27'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-52.6725">52°40.35'W</geoCoordinate>
, 3.xi.2009, FIT, SEAG (CHND). PERU: Madre de Dios: 1: Tambopata, 24.x.1982, rotten palm flowers, L. Watrous &amp; G. Mazurek (FMNH).
</paragraph>
</subSubSection>
<subSubSection lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181" type="description">
<paragraph pageId="180" pageNumber="181">Diagnostic description.</paragraph>
<paragraph lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181">
Length: 1.87-2.22 mm, width: 1.56-1.90 mm; body rufobrunneus to rufopiceous, elongate oval, moderately strongly convex; frons depressed, finely punctate, frontal portion of frontal stria detached from lateral; supraorbital stria complete, but narrowly detached from sides of frontal; epistoma slightly emarginate; labrum rectangular, about twice as wide as long; pronotum curved downward in anterior corners, nearly vertical on either side; pronotal disk lacking prescutellar impression, but strongly depressed across base; ground punctation of pronotal disk very fine and sparse, with ~15 shallowly impressed coarser punctures near sides; marginal pronotal stria obsolete for width of head; lateral submarginal stria complete, continuous with anterior submarginal stria; pronotal gland openings close together behind eye, between end of marginal stria and submarginal stria; elytra strongly swollen in subhumeral interval, with single complete epipleural stria, outer subhumeral stria complete, inner subhumeral stria present in apical half to three-fourths, frequently fragmented basally, rarely absent, striae 1-3 complete, 4th stria present in apical half, 5th stria present in apical third, sutural stria present in apical two-thirds, the sutural stria widening anteriorly; prosternal keel very weakly emarginate at base, carinal striae enclosing triangular area which is finely microsculptured, meeting anteriorly; mesoventrite faintly projecting, with complete marginal stria; mesometaventral arch nearly reaches mesoventral stria, sinuate near mesocoxae, continued by lateral metaventral stria posterolaterally toward middle of metacoxa, slightly curved outward at apex; central portion of metaventral disk weakly depressed on either side; 1st abdominal ventrite with single lateral stria; propygidium with large, elongate punctures separated by about their diameters; pygidial disk strongly convex, with dense, fine ground punctation and numerous conspicuous coarser punctures interspersed; marginal pygidial sulcus distinct, complete, crenulate, with distinct series of punctures within. Male genitalia largely indistinguishable from trhose of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
(see Figs 48 
<normalizedToken originalValue="A–D">A-D</normalizedToken>
), except that S8 has its sides more strongly convergent, halves 
<pageBreakToken pageId="181" pageNumber="182" start="start">ventrally</pageBreakToken>
more strongly divergent along ventral midline, and apical guides narrower; tegmen curved ventrad in apical third(Fig. 48L), but not strongly bent as in 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="181" pageNumber="182" type="remarks">
<paragraph pageId="181" pageNumber="182">Remarks.</paragraph>
<paragraph pageId="181" pageNumber="182">
This species is the only previously described member of a small subgroup of four very similar species (including the following three), all of which have their pronotal gland openings very close together along the anterior margin. All are more or less piceous, and relatively strongly convex, with conspicuously dual punctation on the pygidium (Fig. 49B). 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus distinctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="distinctus">Operclipygus distinctus</taxonomicName>
has the most convex pygidium of the four, with the largest secondary punctures. It also lacks an outer lateral stria on the 1st abdominal ventrite, which the last two species in this subgroup have.
</paragraph>
<caption pageId="181" pageNumber="182">
<paragraph pageId="181" pageNumber="182">
Figure 48. Male genitalia of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
group. A T8 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
B S8 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
C T9 &amp; T10 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
D S9 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
E S9 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus atlanticus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="atlanticus">Operclipygus atlanticus</taxonomicName>
F S8 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus atlanticus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="atlanticus">Operclipygus atlanticus</taxonomicName>
G T9 &amp; T10 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus atlanticus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="atlanticus">Operclipygus atlanticus</taxonomicName>
H S8 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus gilli" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="gilli">Operclipygus gilli</taxonomicName>
I S9 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus gilli" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="gilli">Operclipygus gilli</taxonomicName>
J S9 of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus proximus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="proximus">Operclipygus proximus</taxonomicName>
K Aedeagus, dorsal and lateral views, of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
L Aedeagus, lateral view, of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus distinctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="distinctus">Operclipygus distinctus</taxonomicName>
M Aedeagus, dorsal and lateral views, of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus atlanticus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="atlanticus">Operclipygus atlanticus</taxonomicName>
N Aedeagus, dorsal and lateral views, of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus gilli" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="gilli">Operclipygus gilli</taxonomicName>
O Aedeagus, dorsal and lateral views, of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus proximus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="proximus">Operclipygus proximus</taxonomicName>
P Aedeagus, dorsal and lateral views, of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus prolixus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="prolixus">Operclipygus prolixus</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="181" pageNumber="182">
<paragraph pageId="181" pageNumber="182">
Map 17. Records of the 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
group.
</paragraph>
</caption>
<caption pageId="181" pageNumber="182">
<paragraph pageId="181" pageNumber="182">
Figure 49. 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
group. A Dorsal habitus of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus distinctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="distinctus">Operclipygus distinctus</taxonomicName>
B Pygidia of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus distinctus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="distinctus">Operclipygus distinctus</taxonomicName>
C Dorsal habitus of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus atlanticus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="atlanticus">Operclipygus atlanticus</taxonomicName>
D Dorsal habitus of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus gilli" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="gilli">Operclipygus gilli</taxonomicName>
E Ventral habitus of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus gilli" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="gilli">Operclipygus gilli</taxonomicName>
F Dorsal habitus of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus proximus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="proximus">Operclipygus proximus</taxonomicName>
G Dorsal habitus of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus prolixus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="prolixus">Operclipygus prolixus</taxonomicName>
H Frons of 
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus prolixus" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="prolixus">Operclipygus prolixus</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
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