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<document id="BFF182CD8D3237C51E2408947E99B8E6" ID-CLB-Dataset="56755" ID-DOI="10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FF706AC8FA7F90EA1CBDB800" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Kerivoula papillosa" docType="treatment" docVersion="13" lastPageNumber="900" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="899" updateTime="1718993610009" updateUser="carolina">
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<mods:title id="2C71AFB2DE211FAB2A502A4F22A29488">Vespertilionidae</mods:title>
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<mods:namePart id="40D73F8E43C663B673EE9D2A08539641">Russell A. Mittermeier</mods:namePart>
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<treatment id="4C3D87E8FF706AC8FA7F90EA1CBDB800" ID-DOI="http://doi.org/10.5281/zenodo.6581364" ID-GBIF-Taxon="195628326" ID-Zenodo-Dep="6581364" LSID="urn:lsid:plazi:treatment:4C3D87E8FF706AC8FA7F90EA1CBDB800" httpUri="http://treatment.plazi.org/id/4C3D87E8FF706AC8FA7F90EA1CBDB800" lastPageId="140" lastPageNumber="900" pageId="139" pageNumber="899">
<subSubSection id="8C8E6575FF706ACFFA7F90EA1BD5BDD3" box="[1411,1493,1715,1761]" pageId="139" pageNumber="899" type="multiple">
<paragraph id="C42B36FEFF706ACFFA7F90EA1BD5BDD3" blockId="139.[1408,2459,1715,1840]" box="[1411,1493,1715,1761]" pageId="139" pageNumber="899">
<heading id="9F638192FF706ACFFA7F90EA1BD5BDD3" box="[1411,1493,1715,1761]" pageId="139" pageNumber="899">
<figureCitation id="5CAF2A7BFF706ACFFA7F90EA1BD5BDD3" box="[1411,1493,1715,1761]" captionStart="Plate 67: Vespertilionidae" captionStartId="135.[114,144,3260,3285]" captionTargetBox="[11,2751,19,3659]" captionTargetPageId="134" captionText="293. Groove-toothed Trumpet-eared Bat (Phoniscus atrox), 294. Peterss Trumpet-eared Bat (Phoniscus jagorii), 295. Dubious Trumpet-eared Bat (Phoniscus aerosus), 296. Golden-tipped Bat (Phonuscus papuensis), 297. Hardwickes Woolly Bat (Kerivoula hardwickii), 298. Flatskulled Woolly Bat (Kerivoula depressa), 299. Sri Lankan Woolly Bat (Kerivoula malpasi), 300. Cryptic Woolly Bat (Kerivoula crypta), 301. Dark Woolly Bat (Kerivoula furva), 302. Kachin Woolly Bat (Kerivoula kachinensis), 303. Indochinese Woolly Bat (Kerivoula dongduongana), 304. Krau Woolly Bat (Kerwoula krauensis), 305. Titanias Woolly Bat (Kerwoula titania), 306. Least Woolly Bat (Kerivoula minuta), 307. Small Woolly Bat (Kerivoula intermedia), 308. Papillose Woolly Bat (Kerivoula papillosa), 309. Lenis Woolly Bat (Kerivoula lenis), 310. Clear-winged Woolly Bat (Kerivoula pellucida), 311. Painted Woolly Bat (Kerwoula picta), 312. Whiteheads Woolly Bat (Kerivoula whiteheadi), 313. Flores Woolly Bat (Kerivoula flora), 314. Fly River Woolly Bat (Kerivoula muscina), 315. Saint Aignans Woolly Bat (Kerivoula agnella), 316. Bismarck Woolly Bat (Kerwoula myrella), 317. Lesser Woolly Bat (Kerivoula lanosa), 318. Ethiopian Woolly Bat (Kerwoula eriophora), 319. Tanzanian Woolly Bat (Kerwwoula africana), 320. Smiths Woolly Bat (Kerivoula smithii), 321. Coppery Woolly Bat (Kerivoula cuprosa), 322. Spurrells Woolly Bat (Kerivoula phalaena), 323. Damara Woolly Bat (Kerwoula argentata)" figureDoi="http://doi.org/10.5281/zenodo.6398545" httpUri="https://zenodo.org/record/6398545/files/figure.png" pageId="139" pageNumber="899">308.</figureCitation>
</heading>
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<subSubSection id="8C8E6575FF706ACFFA1A90EA198DBDD3" box="[1510,1933,1715,1761]" pageId="139" pageNumber="899" type="vernacular_names">
<paragraph id="C42B36FEFF706ACFFA1A90EA198DBDD3" blockId="139.[1408,2459,1715,1840]" box="[1510,1933,1715,1761]" pageId="139" pageNumber="899">
<heading id="9F638192FF706ACFFA1A90EA198DBDD3" box="[1510,1933,1715,1761]" pageId="139" pageNumber="899">
<vernacularName id="4A9746D0FF706ACFFA1A90EA198DBDD3" ID-CoL="3R624" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1510,1933,1715,1761]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" language="eng" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">Papillose Woolly Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFF84290EA172CBDD3" box="[1982,2348,1715,1761]" pageId="139" pageNumber="899" type="nomenclature">
<paragraph id="C42B36FEFF706ACFF84290EA172CBDD3" blockId="139.[1408,2459,1715,1840]" box="[1982,2348,1715,1761]" pageId="139" pageNumber="899">
<heading id="9F638192FF706ACFF84290EA172CBDD3" box="[1982,2348,1715,1761]" pageId="139" pageNumber="899">
<taxonomicName id="03944D7DFF706ACFF84290EA172CBDD3" ID-CoL="3R624" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1982,2348,1715,1761]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">
<emphasis id="F6E0EAECFF706ACFF84290EA172CBDD3" box="[1982,2348,1715,1761]" italics="true" pageId="139" pageNumber="899">Kerivoula papillosa</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFFA7E90AD1901BC1C" pageId="139" pageNumber="899" type="vernacular_names">
<paragraph id="C42B36FEFF706ACFFA7E90AD1799BC3B" blockId="139.[1408,2459,1715,1840]" box="[1410,2457,1780,1801]" pageId="139" pageNumber="899">
<heading id="9F638192FF706ACFFA7E90AD1799BC3B" box="[1410,2457,1780,1801]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA7E90AD1BCDBC3B" bold="true" box="[1410,1485,1780,1801]" pageId="139" pageNumber="899">French:</emphasis>
<vernacularName id="4A9746D0FF706ACFFA2B90AD18BCBC3B" ID-CoL="3R624" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1495,1724,1780,1801]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" language="fra" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">Kérivoule mamelonnée</vernacularName>
/
<emphasis id="F6E0EAECFF706ACFF92D90AD192CBC3B" bold="true" box="[1745,1836,1780,1801]" pageId="139" pageNumber="899">German:</emphasis>
<vernacularName id="4A9746D0FF706ACFF8CE90AD1621BC3B" ID-CoL="3R624" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1842,2081,1780,1801]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" language="deu" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">Warzen-Wollfledermaus</vernacularName>
/
<emphasis id="F6E0EAECFF706ACFF7CA90AD1692BC3B" bold="true" box="[2102,2194,1780,1801]" pageId="139" pageNumber="899">Spanish:</emphasis>
<vernacularName id="4A9746D0FF706ACFF76790AD1799BC3B" ID-CoL="3R624" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[2203,2457,1780,1801]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" language="esp" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">
<vernacularName id="4A9746D0FF706ACFF76790AD1705BC3B" ID-CoL="3R624" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[2203,2309,1780,1801]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" language="esp" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">Querivoula</vernacularName>
piloso de
<collectingRegion id="0650F81CFF706ACFF69390AD1799BC3B" box="[2415,2457,1780,1801]" country="Indonesia" name="Jawa Timur" pageId="139" pageNumber="899">Java</collectingRegion>
</vernacularName>
</heading>
</paragraph>
<paragraph id="C42B36FEFF706ACFFA7D91401901BC1C" blockId="139.[1408,2459,1715,1840]" box="[1409,1793,1817,1838]" pageId="139" pageNumber="899">
<heading id="9F638192FF706ACFFA7D91401901BC1C" box="[1409,1793,1817,1838]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA7D91401878BC1C" bold="true" box="[1409,1656,1817,1838]" pageId="139" pageNumber="899">Other common names:</emphasis>
<vernacularName id="4A9746D0FF706ACFF97F91401901BC1C" ID-CoL="3R624" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1667,1793,1817,1838]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" language="eng" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">Papillose Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFF8199139162CBC9A" pageId="139" pageNumber="899" type="reference_group">
<paragraph id="C42B36FEFF706ACFF8199139162CBC9A" blockId="139.[2019,2614,1888,2315]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFF8199139167CBCB3" bold="true" box="[2021,2172,1888,1921]" pageId="139" pageNumber="899">Taxonomy.</emphasis>
<taxonomicName id="03944D7DFF706ACFF76D91391628BC9A" authority="Temminck, 1840" authorityName="Temminck" authorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosus">Vespertilio papillosus Temminck, 1840</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFF7C591D6141AB35F" pageId="139" pageNumber="899" type="materials_examined">
<paragraph id="C42B36FEFF706ACFF7C591D6141AB35F" blockId="139.[2019,2614,1888,2315]" pageId="139" pageNumber="899">
<materialsCitation id="74FC3CA3FF706ACFF7C591D6141AB35F" ID-GBIF-Occurrence="3783319603" pageId="139" pageNumber="899">
“Lile de
<collectingRegion id="0650F81CFF706ACFF75391D616ECBC9A" box="[2223,2284,1935,1960]" country="Indonesia" name="Jawa Timur" pageId="139" pageNumber="899">Java</collectingRegion>
, dans le district sauvage de
<collectingRegion id="0650F81CFF706ACFF7ED91F7167DBCFD" box="[2065,2173,1966,1999]" country="Indonesia" name="Banten" pageId="139" pageNumber="899">Bantam</collectingRegion>
. Se trouve aussi a
<collectingRegion id="0650F81CFF706ACFF68591F717EBBCFD" box="[2425,2539,1966,1999]" country="Indonesia" name="Sumatera Utara" pageId="139" pageNumber="899">Sumatra</collectingRegion>
dans le district de Padang [= Theisland ofJava, in the wild district of
<collectingRegion id="0650F81CFF706ACFF7169E581756B32C" box="[2282,2390,2049,2078]" country="Indonesia" name="Banten" pageId="139" pageNumber="899">Bantam</collectingRegion>
. It is also found in
<collectingRegion id="0650F81CFF706ACFF7F79E7D167DB377" box="[2059,2173,2084,2117]" country="Indonesia" name="Sumatera Utara" pageId="139" pageNumber="899">Sumatra</collectingRegion>
in the district of Padang].” Restricted by G. H. H. Tate in 1940 to “
<collectingRegion id="0650F81CFF706ACFF62E9E151409B35F" box="[2514,2569,2124,2157]" country="Indonesia" name="Jawa Timur" pageId="139" pageNumber="899">Java</collectingRegion>
.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFF81F9E2A1419B2C4" pageId="139" pageNumber="899" type="discussion">
<paragraph id="C42B36FEFF706ACFF81F9E2A1419B2C4" blockId="139.[2019,2614,1888,2315]" lastBlockId="139.[1400,2612,2320,3497]" pageId="139" pageNumber="899">
As currently defined,
<taxonomicName id="03944D7DFF706ACFF6F09E2A17A0B3A6" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[2316,2464,2163,2196]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">K. papillosa</taxonomicName>
represents a species complex that is paraphyletic with K. lenas, which togetherare sisterto the hardwickspecies group. There appearto be several sympatrically distributed forms (at least three), and manyspecimens currently attributed to
<taxonomicName id="03944D7DFF706ACFFA179F6E187FB26A" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1515,1663,2359,2392]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">K. papillosa</taxonomicName>
likely represent
<taxonomicName id="03944D7DFF706ACFF89E9F6E19C0B26A" box="[1890,1984,2359,2392]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="lenis">K. lenis</taxonomicName>
. Genetic and morphological data from across the distribution are needed to determine taxonomy of this complex.
<taxonomicName id="03944D7DFF706ACFF6BF9F0617FDB2B2" box="[2371,2557,2399,2432]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="lenis">Kerivoula lenis</taxonomicName>
was previously considered a subspecies of
<taxonomicName id="03944D7DFF706ACFF88A9FDE1607B29A" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1910,2055,2439,2472]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="papillosa">K. papillosa</taxonomicName>
, but genetic and morphological data support its status as a separate species, especially because the two are broadly sympatric. Identity of
<collectingCountry id="BC83766EFF706ACFFA2E9F8C186AB2C4" box="[1490,1642,2517,2550]" name="Philippines" pageId="139" pageNumber="899">Philippines</collectingCountry>
specimens needs to be addressed further. Two subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFFA829FA61899B188" pageId="139" pageNumber="899" type="distribution">
<caption id="90EB6676FF706ACFFA829FA61899B188" ID-DOI="http://doi.org/10.5281/zenodo.6398464" ID-Zenodo-Dep="6398464" httpUri="https://zenodo.org/record/6398464/files/figure.png" inLine="true" pageId="139" pageNumber="899" targetBox="[1405,1996,1892,2306]" targetPageId="139">
<paragraph id="C42B36FEFF706ACFFA829FA6190EB12E" blockId="139.[1400,2612,2320,3497]" box="[1406,1806,2559,2588]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA829FA6190EB12E" bold="true" box="[1406,1806,2559,2588]" pageId="139" pageNumber="899">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="C42B36FEFF706ACFFA829C701785B15F" blockId="139.[1400,2612,2320,3497]" pageId="139" pageNumber="899">
<taxonomicName id="03944D7DFF706ACFFA829C701785B15F" authority="Temminck, 1840" authorityName="Temminck" authorityYear="1840" baseAuthorityName="Temminck" baseAuthorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="subSpecies" species="papillosa" subSpecies="papillosa">K.p.papillosaTemminck,1840—Sumatra,N&amp;WBorneo,Java,N&amp;CSulawesi,includ-ingnearbyKabaenaandButonIs,andPhilippines(LuzonandBoholIs).</taxonomicName>
</paragraph>
<paragraph id="C42B36FEFF706ACFFA829C251899B188" blockId="139.[1400,2612,2320,3497]" pageId="139" pageNumber="899">
<taxonomicName id="03944D7DFF706ACFFA829C251906B1A7" authority="Chasen, 1940" authorityName="Chasen" authorityYear="1940" box="[1406,1798,2684,2709]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="subSpecies" species="papillosa" subSpecies="malayana">K. p. malayana Chasen, 1940</taxonomicName>
— WC &amp; S
<collectingCountry id="BC83766EFF706ACFF8609C25161EB1A7" box="[1948,2078,2684,2709]" name="Thailand" pageId="139" pageNumber="899">Thailand</collectingCountry>
, NE
<collectingCountry id="BC83766EFF706ACFF79F9C2516A5B1A7" box="[2147,2213,2684,2709]" name="Laos" pageId="139" pageNumber="899">Laos</collectingCountry>
,
<collectingCountry id="BC83766EFF706ACFF74F9C25172DB1A7" box="[2227,2349,2684,2709]" name="Vietnam" pageId="139" pageNumber="899">Vietnam</collectingCountry>
, C
<collectingCountry id="BC83766EFF706ACFF6A09C2517EEB1A7" box="[2396,2542,2684,2709]" name="Cambodia" pageId="139" pageNumber="899">Cambodia</collectingCountry>
, and Peninsular
<collectingCountry id="BC83766EFF706ACFF9E19CC41895B188" box="[1565,1685,2717,2746]" name="Malaysia" pageId="139" pageNumber="899">Malaysia</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFFA819C9F1707B7A7" pageId="139" pageNumber="899" type="description">
<paragraph id="C42B36FEFF706ACFFA819C9F1707B7A7" blockId="139.[1400,2612,2320,3497]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA819C9F1877B1D1" bold="true" box="[1405,1655,2758,2787]" pageId="139" pageNumber="899">Descriptive notes.</emphasis>
Head-body 40-3-52-
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, tail 47-56-
<quantity id="036C9B1BFF706ACFF75E9C9F16F7B1D1" box="[2210,2295,2758,2787]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="139" pageNumber="899" unit="mm" value="4.0">4 mm</quantity>
, ear
<quantity id="036C9B1BFF706ACFF6BF9C9F17D6B1D1" box="[2371,2518,2758,2787]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="139" pageNumber="899" unit="mm" value="15.5" valueMax="17.0" valueMin="14.0">14-17 mm</quantity>
, hindfoot 7-9-
<quantity id="036C9B1BFF706ACFFA059CB71862B039" box="[1529,1634,2798,2827]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="139" pageNumber="899" unit="mm" value="10.0">10 mm</quantity>
, forearm
<quantity id="036C9B1BFF706ACFF90E9CB71988B039" box="[1778,1928,2798,2827]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="4.9" metricValueMin="3.9" pageId="139" pageNumber="899" unit="mm" value="44.0" valueMax="49.0" valueMin="39.0">39-49 mm</quantity>
; weight
<quantity id="036C9B1BFF706ACFF7F89CB71663B039" box="[2052,2147,2798,2827]" metricMagnitude="-3" metricUnit="kg" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="139" pageNumber="899" unit="g" value="9.5" valueMax="13.0" valueMin="6.0">6-13 g</quantity>
. Furis dense and woolly. Dorsal pelage is brown to buffy brown; venter is paler. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragus is narrow and tall, with virtuallystraight anterior margin except for very slight convexity near tip and concave posterior margin with small hooked basal lobe. Membranes are light brown, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes, and calcar is long. Skull has high braincase and domed forehead, with more robust rostrum than in the
<taxonomicName id="03944D7DFF706ACFFA809A701831B774" authority="Woolly" authorityName="Woolly" box="[1404,1585,3113,3142]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="lenis">Lenis Woolly</taxonomicName>
Bat (
<taxonomicName id="03944D7DFF706ACFF97D9A7018ECB774" box="[1665,1772,3113,3142]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="lenis">K. lenis</taxonomicName>
). Compared to the
<taxonomicName id="03944D7DFF706ACFF7FB9A7016BCB774" authority="Woolly" authorityName="Woolly" box="[2055,2236,3113,3142]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="139" pageNumber="899" phylum="Chordata" rank="species" species="lenis">Lenis Woolly</taxonomicName>
Bat, dentition is more robust overall, upper premolars are wider, and lower premolars are larger, especially P. Chromosomal complement has 2n = 38 and FN = 54 (
<collectingCountry id="BC83766EFF706ACFF78F9A2D16F9B7A7" box="[2163,2297,3188,3221]" name="Thailand" pageId="139" pageNumber="899">Thailand</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFFA879AC51654B7D1" pageId="139" pageNumber="899" type="biology_ecology">
<paragraph id="C42B36FEFF706ACFFA879AC51654B7D1" blockId="139.[1400,2612,2320,3497]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA879AC51BEAB78F" bold="true" box="[1403,1514,3228,3261]" pageId="139" pageNumber="899">Habitat.</emphasis>
Lowland mixed deciduous dipterocarp forests and secondary lowland forests (Luzon) from sea level up to elevations of ¢.
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.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFFA869AB0196FB638" box="[1402,1903,3305,3338]" pageId="139" pageNumber="899" type="food_feeding">
<paragraph id="C42B36FEFF706ACFFA869AB0196FB638" blockId="139.[1400,2612,2320,3497]" box="[1402,1903,3305,3338]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA869AB01880B638" bold="true" box="[1402,1664,3305,3338]" pageId="139" pageNumber="899">Food and Feeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706ACFFA869B4D18F0B603" box="[1402,1776,3348,3377]" pageId="139" pageNumber="899" type="breeding">
<paragraph id="C42B36FEFF706ACFFA869B4D18F0B603" blockId="139.[1400,2612,2320,3497]" box="[1402,1776,3348,3377]" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA869B4D1800B603" bold="true" box="[1402,1536,3348,3377]" pageId="139" pageNumber="899">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF706AC8FA849B631AC1BA94" lastPageId="140" lastPageNumber="900" pageId="139" pageNumber="899" type="activity">
<paragraph id="C42B36FEFF706AC8FA849B631AC1BA94" blockId="139.[1400,2612,2320,3497]" lastBlockId="140.[162,1369,313,818]" lastPageId="140" lastPageNumber="900" pageId="139" pageNumber="899">
<emphasis id="F6E0EAECFF706ACFFA849B63185CB669" bold="true" box="[1400,1628,3386,3419]" pageId="139" pageNumber="899">Activity patterns.</emphasis>
The Papillose Woolly Bat roosts in small tree hollows. Call shape is steep FM sweep. Calls of two (B and C group) ofthe three genetic lineages in
<collectingCountry id="BC83766EFF706ACFF69B9B3817E4B6B0" box="[2407,2532,3425,3458]" name="Thailand" pageId="139" pageNumber="899">Thailand</collectingCountry>
were recorded: B group, start frequencies of 180-196 kHz (mean 190-3 kHz), end frequencies of 60-69 kHz (65-5 kHz), and peak frequencies of 109-9-157-6 kHz (131-6 kHz) and C group,start frequencies of 169-207-7 kHz (189-8 kHz), end frequencies of 63-5- 89-8 kHz (75-7 kHz), and peak frequencies of 112-1-161-4 kHz (140-8 kHz).
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF776AC8FF5897F51C9CBAC4" pageId="140" pageNumber="900" type="biology_ecology">
<paragraph id="C42B36FEFF776AC8FF5897F51C9CBAC4" blockId="140.[162,1369,313,818]" pageId="140" pageNumber="900">
<emphasis id="F6E0EAECFF776AC8FF5897F51D59BAFF" bold="true" box="[164,857,428,461]" pageId="140" pageNumber="900">Movements, Home range and Social organization.</emphasis>
The Papillose Woolly Bat reportedly roosts in groups of 1-14 individuals.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF776AC8FF5897A21A67B988" pageId="140" pageNumber="900" type="conservation">
<paragraph id="C42B36FEFF776AC8FF5897A21A67B988" blockId="140.[162,1369,313,818]" pageId="140" pageNumber="900">
<emphasis id="F6E0EAECFF776AC8FF5897A21C01B92E" bold="true" box="[164,513,507,540]" pageId="140" pageNumber="900">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Papillose Woolly Bat appears to be common in much of its wide distribution. Nevertheless, its taxonomy is unresolved, and it likely represents several species that would have smaller distributions. Further taxonomic studies are needed before a proper assessment can be made. Deforestation might be a threat in some regions.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF776AC8FF5A94911CBDB800" pageId="140" pageNumber="900" type="bibRefCitation_list">
<paragraph id="C42B36FEFF776AC8FF5A94911CBDB800" blockId="140.[162,1369,313,818]" pageId="140" pageNumber="900">
<emphasis id="F6E0EAECFF776AC8FF5A94911F3EB9D3" bold="true" box="[166,318,712,737]" pageId="140" pageNumber="900">Bibliography.</emphasis>
Douangboubpha et al. (2016), Duya et al. (2007), Francis (2008a), Hasan &amp; Abdullah (2011), Hutson &amp;
<collectingRegion id="0650F81CFF776AC8FEEA94B61F73B83A" box="[278,371,751,776]" country="Jamaica" name="Kingston" pageId="140" pageNumber="900">Kingston</collectingRegion>
(2008e), Khan et al. (2010), Kruskop (2013a), Kuo Haochih et al. (2017), Struebig, Horsburgh et al. (2008), Tate (1940), Vanitharani et al. (2003).
</paragraph>
</subSubSection>
</treatment>
</document>