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<document id="FC692B214C00DAD780CF717F1B1DE8BD" ID-CLB-Dataset="56755" ID-DOI="10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FF366A8AFA799C2F1687BA9D" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Myotis horsfieldii" docType="treatment" docVersion="16" lastPageNumber="968" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="967" updateTime="1718993610009" updateUser="carolina">
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<mods:title id="2277F2CD142B7F813FBA80ECF6131F6E">Vespertilionidae</mods:title>
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<mods:namePart id="7A915020F9B462586841001948E2D778">Russell A. Mittermeier</mods:namePart>
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<mods:title id="B3A92AC6AF623CABD9AA9AEA26629E17">Handbook of the Mammals of the World – Volume 9 Bats</mods:title>
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<treatment id="4C3D87E8FF366A8AFA799C2F1687BA9D" ID-DOI="http://doi.org/10.5281/zenodo.6577946" ID-GBIF-Taxon="195628114" ID-Zenodo-Dep="6577946" LSID="urn:lsid:plazi:treatment:4C3D87E8FF366A8AFA799C2F1687BA9D" httpUri="http://treatment.plazi.org/id/4C3D87E8FF366A8AFA799C2F1687BA9D" lastPageId="206" lastPageNumber="968" pageId="205" pageNumber="967">
<subSubSection id="8C8E6575FF366A89FA799C2F1BD8B196" box="[1413,1496,2678,2724]" pageId="205" pageNumber="967" type="multiple">
<paragraph id="C42B36FEFF366A89FA799C2F1BD8B196" blockId="205.[1411,2436,2678,2807]" box="[1413,1496,2678,2724]" pageId="205" pageNumber="967">
<heading id="9F638192FF366A89FA799C2F1BD8B196" box="[1413,1496,2678,2724]" pageId="205" pageNumber="967">
<figureCitation id="5CAF2A7BFF366A89FA799C2F1BD8B196" box="[1413,1496,2678,2724]" captionStart="Plate 73: Vespertilionidae" captionStartId="199.[131,161,3286,3311]" captionTargetBox="[12,2760,18,3659]" captionTargetPageId="198" captionText="449. Hairy-faced Myotis (Myotis annectans), 450. Frosted Myotis (Myotis pruinosus), 451. Yanbaru Whiskered Myotis (Myotis yanbarensis), 452. Long-toed Myotis (Myotis secundus), 453. Bornean Whiskered Myotis (Myotis borneoensis), 454. Malaysian Whiskered Myotis (Myotis federatus), 455. Indochinese Myotis (Myotis indochinensis), 456. Peyton’s Myotis (Myotis peytoni), 457. Burmese Whiskered Myotis (Myotis montivagus), 458. Long-fingered Myotis (Myotis capaccinii), 459. Chinese Water Myotis (Myotis laniger), 460. Csorba’s Myotis (Myotis csorbai), 461. Kashmir Cave Myotis (Myotis longipes), 462. Annamit Myotis (Myotis annamiticus), 463. Phan Luong’s Myotis (Myotis phanluongi), 464. Himalayan Whiskered Myotis (Myotis siligorensis), 465. Chestnut Myotis (Myotis badius), 466. Ridley’s Myotis (Myotis ridleyi), 467. Thick-thumbed Myotis (Myotis rosseti), 468. Horsfield’s Myotis (Myotis horsfieldii), 469. Pallid Large-footed Myotis (Myotis macrotarsus), 470. Stalker’s Myotis (Myotis stalkeri), 471. Gray Large-footed Myotis (Myotis adversus), 472. Maluku Myotis (Myotis moluccarum), 473. Southern Myotis (Myotis macropus), 474. Lesser Large-footed Myotis (Myotis hasseltii)" figureDoi="http://doi.org/10.5281/zenodo.6398948" httpUri="https://zenodo.org/record/6398948/files/figure.png" pageId="205" pageNumber="967">468.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF366A89FA159C2F195EB196" box="[1513,1886,2678,2724]" pageId="205" pageNumber="967" type="vernacular_names">
<paragraph id="C42B36FEFF366A89FA159C2F195EB196" blockId="205.[1411,2436,2678,2807]" box="[1513,1886,2678,2724]" pageId="205" pageNumber="967">
<heading id="9F638192FF366A89FA159C2F195EB196" box="[1513,1886,2678,2724]" pageId="205" pageNumber="967">
<vernacularName id="4A9746D0FF366A89FA159C2F195EB196" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1513,1886,2678,2724]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" language="eng" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Horsfield’s Myotis</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF366A89F86E9C2F16D0B196" box="[1938,2256,2678,2724]" pageId="205" pageNumber="967" type="nomenclature">
<paragraph id="C42B36FEFF366A89F86E9C2F16D0B196" blockId="205.[1411,2436,2678,2807]" box="[1938,2256,2678,2724]" pageId="205" pageNumber="967">
<heading id="9F638192FF366A89F86E9C2F16D0B196" box="[1938,2256,2678,2724]" pageId="205" pageNumber="967">
<taxonomicName id="03944D7DFF366A89F86E9C2F16D0B196" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1938,2256,2678,2724]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">
<emphasis id="F6E0EAECFF366A89F86E9C2F16D0B196" box="[1938,2256,2678,2724]" italics="true" pageId="205" pageNumber="967">Myotis horsfieldii</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF366A89FA799CEE1708B1C1" pageId="205" pageNumber="967" type="vernacular_names">
<paragraph id="C42B36FEFF366A89FA799CEE1783B1FE" blockId="205.[1411,2436,2678,2807]" box="[1413,2435,2743,2764]" pageId="205" pageNumber="967">
<heading id="9F638192FF366A89FA799CEE1783B1FE" box="[1413,2435,2743,2764]" pageId="205" pageNumber="967">
<emphasis id="F6E0EAECFF366A89FA799CEE1BD0B1FE" bold="true" box="[1413,1488,2743,2764]" pageId="205" pageNumber="967">French:</emphasis>
<vernacularName id="4A9746D0FF366A89FA269CEE1893B1FE" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1498,1683,2743,2764]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" language="fra" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Murin de Horsfield</vernacularName>
/
<emphasis id="F6E0EAECFF366A89F95B9CEE1902B1FE" bold="true" box="[1703,1794,2743,2764]" pageId="205" pageNumber="967">German:</emphasis>
<vernacularName id="4A9746D0FF366A89F8F09CEE162EB1FE" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1804,2094,2743,2764]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" language="deu" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Horsfield-LangfuRfledermaus</vernacularName>
/
<emphasis id="F6E0EAECFF366A89F7B89CEE169FB1FE" bold="true" box="[2116,2207,2743,2764]" pageId="205" pageNumber="967">Spanish:</emphasis>
<vernacularName id="4A9746D0FF366A89F7569CEE1783B1FE" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[2218,2435,2743,2764]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" language="esp" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Ratonero de Horsfield</vernacularName>
</heading>
</paragraph>
<paragraph id="C42B36FEFF366A89FA789C871708B1C1" blockId="205.[1411,2436,2678,2807]" box="[1412,2312,2782,2803]" pageId="205" pageNumber="967">
<heading id="9F638192FF366A89FA789C871708B1C1" box="[1412,2312,2782,2803]" pageId="205" pageNumber="967">
<emphasis id="F6E0EAECFF366A89FA789C87187BB1C1" bold="true" box="[1412,1659,2782,2803]" pageId="205" pageNumber="967">Other common names:</emphasis>
<vernacularName id="4A9746D0FF366A89F9799C871976B1C1" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1669,1910,2782,2803]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" language="eng" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Common Asiatic Myotis</vernacularName>
,
<vernacularName id="4A9746D0FF366A89F8789C871615B1C1" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[1924,2069,2782,2803]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" language="eng" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Horsfield's Bat</vernacularName>
,
<vernacularName id="4A9746D0FF366A89F7DF9C871708B1C1" ID-CoL="8LZBB" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[2083,2312,2782,2803]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" language="eng" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Lesser Large-tooth Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF366A89F8149D7E1632B059" pageId="205" pageNumber="967" type="reference_group">
<paragraph id="C42B36FEFF366A89F8149D7E1632B059" blockId="205.[2023,2616,2855,3277]" pageId="205" pageNumber="967">
<emphasis id="F6E0EAECFF366A89F8149D7E1683B076" bold="true" box="[2024,2179,2855,2884]" pageId="205" pageNumber="967">Taxonomy.</emphasis>
<taxonomicName id="03944D7DFF366A89F76F9D7E162DB059" ID-CoL="85317cef-ba8c-4dd5-be2d-3528232b04ed" authority="Temminck, 1840" authorityName="Temminck" authorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Vespertilio horsfieldii Temminck, 1840</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF366A89F7C19D1317DBB059" box="[2109,2523,2890,2923]" pageId="205" pageNumber="967" type="materials_examined">
<paragraph id="C42B36FEFF366A89F7C19D1317DBB059" blockId="205.[2023,2616,2855,3277]" box="[2109,2523,2890,2923]" pageId="205" pageNumber="967">
<materialsCitation id="74FC3CA3FF366A89F7C19D1317D8B059" ID-GBIF-Occurrence="3783319457" box="[2109,2520,2890,2923]" pageId="205" pageNumber="967">
Mount Gede,
<collectingRegion id="0650F81CFF366A89F7009D13173AB059" box="[2300,2362,2890,2923]" country="Indonesia" name="Jawa Timur" pageId="205" pageNumber="967">Java</collectingRegion>
,
<collectingCountry id="BC83766EFF366A89F6B79D1317D8B059" box="[2379,2520,2890,2923]" name="Indonesia" pageId="205" pageNumber="967">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF366A89F8159D281765B797" pageId="205" pageNumber="967" type="discussion">
<paragraph id="C42B36FEFF366A89F8159D281765B797" blockId="205.[2023,2616,2855,3277]" pageId="205" pageNumber="967">
Subgenus
<taxonomicName id="03944D7DFF366A89F7839D2816D3B0A0" authority="Kaup, 1829" box="[2175,2259,2929,2962]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="genus">Myotis</taxonomicName>
;
<taxonomicName id="03944D7DFF366A89F7149D281762B0A0" authorityName="Temminck" authorityYear="1840" box="[2280,2402,2929,2962]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">horsfieldii</taxonomicName>
species group.
<collectingRegion id="0650F81CFF366A89F8159DC01618B088" box="[2025,2072,2969,3002]" country="South Sudan" name="Lakes" pageId="205" pageNumber="967">See</collectingRegion>
<taxonomicName id="03944D7DFF366A89F7D59DC016ACB088" authorityName="Thomas" authorityYear="1898" box="[2089,2220,2969,3002]" class="Mammalia" family="Vespertilionidae" genus="Pipistrellus" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="970" phylum="Chordata" rank="species" species="ridleyi">M. ridleyi</taxonomicName>
.
<taxonomicName id="03944D7DFF366A89F73D9DC01715B088" authority="Kaup, 1829" box="[2241,2325,2969,3002]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="genus">Myotis</taxonomicName>
horsfieldit was previously included under
<taxonomicName id="03944D7DFF366A89F6E69D9917B5B0D3" baseAuthorityName="Horsfield" baseAuthorityYear="1824" box="[2330,2485,3008,3041]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="adversus">M. adversus</taxonomicName>
and later found closely related to
<taxonomicName id="03944D7DFF366A89F6C49DBE17FCB73A" baseAuthorityName="Waterhouse" baseAuthorityYear="1845" box="[2360,2556,3047,3080]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="macrotarsus">M. macrotarsus</taxonomicName>
and M.
<taxonomicName id="03944D7DFF366A89F7E09A561680B702" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[2076,2176,3087,3120]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="hasseltii">hasseltii</taxonomicName>
, with
<taxonomicName id="03944D7DFF366A89F7219A561765B702" authorityName="E. H. Taylor" authorityYear="1934" box="[2269,2405,3087,3120]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="browni">M. browni</taxonomicName>
in a more basal position.
<taxonomicName id="03944D7DFF366A89F75E9A6F177DB765" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[2210,2429,3126,3159]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="species" species="horsfieldii">Myotis horsfieldii</taxonomicName>
represents a species complex in need of deep revision. Five subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF366A8AF81B9AF51DB2BABF" lastPageId="206" lastPageNumber="968" pageId="205" pageNumber="967" type="distribution">
<caption id="90EB6676FF366A8AF81B9AF51DB2BABF" ID-DOI="http://doi.org/10.5281/zenodo.6398930" ID-Zenodo-Dep="6398930" httpUri="https://zenodo.org/record/6398930/files/figure.png" inLine="true" lastPageId="206" lastPageNumber="968" pageId="205" pageNumber="967" targetBox="[1408,1999,2860,3273]" targetPageId="205">
<paragraph id="C42B36FEFF366A89F81B9AF51777B7FF" blockId="205.[2023,2616,2855,3277]" box="[2023,2423,3244,3277]" pageId="205" pageNumber="967">
<emphasis id="F6E0EAECFF366A89F81B9AF51777B7FF" bold="true" box="[2023,2423,3244,3277]" pageId="205" pageNumber="967">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="C42B36FEFF366A89FA7F9A8A1432B629" blockId="205.[1410,2616,3283,3473]" pageId="205" pageNumber="967">
<taxonomicName id="03944D7DFF366A89FA7F9A8A1432B629" authority="Temminck, 1840" authorityName="Temminck" authorityYear="1840" baseAuthorityName="Temminck" baseAuthorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="subSpecies" species="horsfieldii" subSpecies="horsfieldii">M.h.horsfieldiiTemminck,1840—PeninsularMalaysia(includingLangkawiandPen-angIs),Singapore,Borneo,SSumatra(Lampung),Java,Bali,Lombok,andSulawesi.</taxonomicName>
</paragraph>
<paragraph id="C42B36FEFF366A89FA7F9B7B187EB65A" blockId="205.[1410,2616,3283,3473]" pageId="205" pageNumber="967">
<taxonomicName id="03944D7DFF366A89FA7F9B7B187EB65A" authority="Shamel, 1942" authorityName="Shamel" authorityYear="1942" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="subSpecies" species="horsfieldii" subSpecies="deignani">M.h.deignaniShamel,1942—SEChina(Guangdong,Hongkong,andHainan)andmainlandSEAsia.</taxonomicName>
</paragraph>
<paragraph id="C42B36FEFF366A89FA7E9B2916AAB6A3" blockId="205.[1410,2616,3283,3473]" box="[1410,2218,3440,3473]" pageId="205" pageNumber="967">
<taxonomicName id="03944D7DFF366A89FA7E9B2916AAB6A3" authority="K. Andersen, 1907" authorityName="K. Andersen" authorityYear="1907" box="[1410,2218,3440,3473]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="205" pageNumber="967" phylum="Chordata" rank="subSpecies" species="horsfieldii" subSpecies="dryas">M.h.dryasK.Andersen,1907—AndamanandNicobarIs.</taxonomicName>
</paragraph>
<paragraph id="C42B36FEFF356A8AFF5D97441D14BA0C" blockId="206.[160,1379,285,3479]" box="[161,788,285,318]" pageId="206" pageNumber="968">
<taxonomicName id="03944D7DFF356A8AFF5D97441D14BA0C" authority="E. H. Taylor, 1934" authorityName="Taylor" authorityYear="1934" box="[161,788,285,318]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="subSpecies" species="horsfieldii" subSpecies="jeannei">M.h.jeanne:E.H.Taylor,1934—Philippines.</taxonomicName>
</paragraph>
<paragraph id="C42B36FEFF356A8AFF5D971D1DB2BABF" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<taxonomicName id="03944D7DFF356A8AFF5D971D1C1EBA57" authority="Thomas, 1915" authorityName="Thomas" authorityYear="1915" box="[161,542,324,357]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="subSpecies" species="horsfieldii" subSpecies="peshwa">M. h. peshwa Thomas, 1915</taxonomicName>
— C, W &amp; NE
<collectingCountry id="BC83766EFF356A8AFD10971D1D35BA57" box="[748,821,324,357]" name="India" pageId="206" pageNumber="968">India</collectingCountry>
(
<collectingRegion id="0650F81CFF356A8AFCB1971D1A3BBA57" box="[845,1083,324,357]" country="India" name="Madhya Pradesh" pageId="206" pageNumber="968">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="0650F81CFF356A8AFBB6971D1AA5BA57" box="[1098,1189,324,357]" country="India" name="Assam" pageId="206" pageNumber="968">Assam</collectingRegion>
,
<collectingRegion id="0650F81CFF356A8AFB4B971D1B4FBA57" box="[1207,1359,324,357]" country="India" name="Meghalaya" pageId="206" pageNumber="968">Meghalaya</collectingRegion>
,
<collectingRegion id="0650F81CFF356A8AFF5C972D1F53BABF" box="[160,339,372,397]" country="India" name="Maharashtra" pageId="206" pageNumber="968">Maharashtra</collectingRegion>
,
<collectingCountry id="BC83766EFF356A8AFE98972D1F9EBABF" box="[356,414,372,397]" name="India" pageId="206" pageNumber="968">Goa</collectingCountry>
,
<collectingRegion id="0650F81CFF356A8AFE53972D1C3FBABF" box="[431,575,372,397]" country="India" name="Karnataka" pageId="206" pageNumber="968">Karnataka</collectingRegion>
,
<collectingRegion id="0650F81CFF356A8AFDB3972D1CABBABF" box="[591,683,372,397]" country="India" name="Kerala" pageId="206" pageNumber="968">Kerala</collectingRegion>
, and
<collectingRegion id="0650F81CFF356A8AFD04972D1DA4BABF" box="[760,932,372,397]" country="India" name="Tamil Nadu" pageId="206" pageNumber="968">Tamil Nadu</collectingRegion>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF5C97C31FB4BD97" pageId="206" pageNumber="968" type="description">
<paragraph id="C42B36FEFF356A8AFF5C97C31FB4BD97" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF5C97C31F9CBA81" bold="true" box="[160,412,410,435]" pageId="206" pageNumber="968">Descriptive notes.</emphasis>
Head-body
<quantity id="036C9B1BFF356A8AFDAB97C31CE7BA81" box="[599,743,410,435]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.1" metricValueMin="4.4" pageId="206" pageNumber="968" unit="mm" value="47.5" valueMax="51.0" valueMin="44.0">44-51 mm</quantity>
,tail
<quantity id="036C9B1BFF356A8AFCCA97C31DCDBA81" box="[822,973,410,435]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.3" metricValueMin="3.3" pageId="206" pageNumber="968" unit="mm" value="38.0" valueMax="43.0" valueMin="33.0">33-43 mm</quantity>
, ear 13-5—-
<quantity id="036C9B1BFF356A8AFB9597C31AC8BA81" box="[1129,1224,410,435]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="206" pageNumber="968" unit="mm" value="17.0">17 mm</quantity>
, hindfoot
<quantity id="036C9B1BFF356A8AFF5E97E31F25BAE9" box="[162,293,442,475]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="206" pageNumber="968" unit="mm" value="9.0" valueMax="11.0" valueMin="7.0">7-11 mm</quantity>
, forearm 33-8-41-
<quantity id="036C9B1BFF356A8AFDD497E31C7CBAE9" box="[552,636,442,475]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="206" pageNumber="968" unit="mm" value="5.0">5 mm</quantity>
; weight 5-7-
<quantity id="036C9B1BFF356A8AFCD797E31D59BAE9" box="[811,857,442,475]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="206" pageNumber="968" unit="g" value="5.0">5 g</quantity>
. Specimens from
<collectingCountry id="BC83766EFF356A8AFBAB97E31AA0BAE9" box="[1111,1184,442,475]" name="India" pageId="206" pageNumber="968">India</collectingCountry>
(peshwa) are slightly larger (forearm 36-5—41-
<quantity id="036C9B1BFF356A8AFD8497B81CCDB930" box="[632,717,481,514]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="206" pageNumber="968" unit="mm" value="5.0">5 mm</quantity>
) and have browner pelage. Subspecies dryas from the Andaman Islands is on average smaller (forearm 34-8-37-
<quantity id="036C9B1BFF356A8AFBA694501AB1B918" box="[1114,1201,521,554]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="206" pageNumber="968" unit="mm" value="1.0">1 mm</quantity>
). Fur is soft and dense,feet large, and ears long. Upperparts are dark brown to almost black, with dark brown hair tips that can be grizzled with gray in some individuals. Underparts are gray to dark gray, with pale gray or buffy hair tips and very dark roots. Fur surrounding anal region is nearly white. Ears are dark brown to black and relatively tall compared with size of head and have smoothly convex anterior borders,slightly concave posterior borders, and bluntly pointed tips. Tragus is narrowly pointed and is less than one-half the height of pinna. Feet are moderately large and exceed one-half the length of tibia. Claws are small. Interfemoral and wing membranes are chocolate-brown. Wing membrane attaches to metatarsus below ankle. Subspecies deignani has tricolored hairs: broad band of dark basal hairs, followed by pale buff band, and then narrow dark brown band, tipped with flaxen. Mingling of these colors gives it a grizzled appearance compared with individuals from Java, which does not have a tricolored appearance and is larger in external and cranial measurements. Baculum is small and saddle-shaped, with rounded tip (specimen from
<collectingCountry id="BC83766EFF356A8AFBDA92681A75BF60" box="[1062,1141,1073,1106]" name="India" pageId="206" pageNumber="968">India</collectingCountry>
). Of two penial bones from
<collectingCountry id="BC83766EFF356A8AFEB592001FC4BF48" box="[329,452,1113,1146]" name="Vietnam" pageId="206" pageNumber="968">Vietnam</collectingCountry>
, one (probably immature specimen) was similar to that described from
<collectingCountry id="BC83766EFF356A8AFF0B92DD1F42BF93" box="[247,322,1156,1185]" name="India" pageId="206" pageNumber="968">India</collectingCountry>
. The second was larger, c.
<quantity id="036C9B1BFF356A8AFD3892DD1D4CBF93" box="[708,844,1156,1185]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="7.2" metricValueMin="0.0" pageId="206" pageNumber="968" unit="mm" value="36.0" valueMax="72.0" valueMin="0.0">0-72 mm</quantity>
, with almost parallel sides, abruptly narrowing to blunt tip; its upper side had abrupt tooth-like elevation on its basal onequarter. Skull is medium-sized, with moderately broad (7-2-7-
<quantity id="036C9B1BFF356A8AFBEA92961A6DBFC2" box="[1046,1133,1231,1264]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="206" pageNumber="968" unit="mm" value="7.0">7 mm</quantity>
) braincase. Rostrum is robust, with shallow depression in its midline. Sagittal crest is absent or very weak. Supraocciptal is bulbous. Zygomata are well developed and outwardly flared. C! is nearly twice the height of P*. P? is ¢.50% of crown area of P?, usually lies in tooth row oris slightly displaced internally, and is in close contact with P*. Upper molars are typically myotodont. P, is ¢.50% the height of the canine, which is higher than P. P,, which occasionally can be displaced internally, is sometimes in close contact with P,. Condylo-basal lengths are 14-3-14-
<quantity id="036C9B1BFF356A8AFD6893BD1CE9BD37" box="[660,745,1508,1541]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="206" pageNumber="968" unit="mm" value="8.0">8 mm</quantity>
; maxillary tooth row lengths are 5-8-
<quantity id="036C9B1BFF356A8AFB0093BD1B54BD37" box="[1276,1364,1508,1541]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="206" pageNumber="968" unit="mm" value="6.0">6 mm</quantity>
. Chromosomal complement has 2n = 44 and FN = 52 (
<collectingCountry id="BC83766EFF356A8AFC5B90551A29BD1F" box="[935,1065,1548,1581]" name="Thailand" pageId="206" pageNumber="968">Thailand</collectingCountry>
, Peninsular
<collectingCountry id="BC83766EFF356A8AFB2090551B54BD1F" box="[1244,1364,1548,1581]" name="Malaysia" pageId="206" pageNumber="968">Malaysia</collectingCountry>
, and
<collectingRegion id="0650F81CFF356A8AFF19906E1F39BD66" box="[229,313,1591,1620]" country="Malaysia" name="Sabah" pageId="206" pageNumber="968">Sabah</collectingRegion>
in Borneo) and 2n = 44 and FN = 50 (
<collectingRegion id="0650F81CFF356A8AFC95906E1DD4BD66" box="[873,980,1591,1620]" country="China" name="Hainan" pageId="206" pageNumber="968">Hainan</collectingRegion>
,
<collectingCountry id="BC83766EFF356A8AFC1B906E1A42BD66" box="[999,1090,1591,1620]" name="China" pageId="206" pageNumber="968">China</collectingCountry>
). This difference in numbers of chromosomal arms was due to use of different criteria for classification of chromosome types.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF5990F01A00BCED" pageId="206" pageNumber="968" type="biology_ecology">
<paragraph id="C42B36FEFF356A8AFF5990F01A00BCED" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF5990F01F14BDF8" bold="true" box="[165,276,1705,1738]" pageId="206" pageNumber="968">Habitat.</emphasis>
Wide variety of habitats including wooded areas with local supply of freshwater, mixed and hill forests, and tea estates (
<collectingCountry id="BC83766EFF356A8AFCD3908C1D7FBDC0" box="[815,895,1749,1778]" name="India" pageId="206" pageNumber="968">India</collectingCountry>
); lowland forests and agricultural areas, adjacent to limestone karst (
<collectingCountry id="BC83766EFF356A8AFD6D90A11D1EBC2B" box="[657,798,1784,1817]" name="Myanmar" pageId="206" pageNumber="968">Myanmar</collectingCountry>
); dry dipterocarp and dry evergreen forests (
<collectingCountry id="BC83766EFF356A8AFF10917D1F72BC73" box="[236,370,1828,1857]" name="Thailand" pageId="206" pageNumber="968">Thailand</collectingCountry>
); lowland forests,riverine forests, disturbed forests, and open land near villages (
<collectingCountry id="BC83766EFF356A8AFEDF91121FA1BC5A" box="[291,417,1867,1896]" name="Vietnam" pageId="206" pageNumber="968">Vietnam</collectingCountry>
); lowland and hill forest (
<collectingCountry id="BC83766EFF356A8AFCE691121D98BC5A" box="[794,920,1867,1896]" name="Malaysia" pageId="206" pageNumber="968">Malaysia</collectingCountry>
); lowland forests, close to large rivers and streams (Borneo and south-western Sumatra); secondary and primary lowland, montane, and mossy forests and agricultural areas (
<collectingCountry id="BC83766EFF356A8AFC3391CF1A75BC85" box="[975,1141,1942,1975]" name="Philippines" pageId="206" pageNumber="968">Philippines</collectingCountry>
); and secondary forests (Sulawesi) from sea level up to elevations of ¢.
<quantity id="036C9B1BFF356A8AFC7291E71DFCBCED" box="[910,1020,1982,2015]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="206" pageNumber="968" unit="m" value="1450.0">1450 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF5991BC1F67B394" pageId="206" pageNumber="968" type="food_feeding">
<paragraph id="C42B36FEFF356A8AFF5991BC1F67B394" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF5991BC1FB6B334" bold="true" box="[165,438,2021,2054]" pageId="206" pageNumber="968">Food and Feeding.</emphasis>
Horsfield’s
<taxonomicName id="03944D7DFF356A8AFD9291BC1CC8B334" authority="Kaup, 1829" box="[622,712,2021,2054]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
typically flies in circles c.
<quantity id="036C9B1BFF356A8AFBBD91BC1A9DB334" box="[1089,1181,2021,2054]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="206" pageNumber="968" unit="cm" value="10.0">10 cm</quantity>
above water, similar to European Daubenton’s
<taxonomicName id="03944D7DFF356A8AFD6A9E541CF0B31C" authority="Kaup, 1829" box="[662,752,2061,2094]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
(
<taxonomicName id="03944D7DFF356A8AFCF79E541DDFB31C" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[779,991,2061,2094]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="975" phylum="Chordata" rank="species" species="daubentonii">M. daubentonii</taxonomicName>
). It was observed foraging near forest streams, skimming insects from water surface in some instances. On Da Lat plateau,
<collectingCountry id="BC83766EFF356A8AFEA49E051FD3B34F" box="[344,467,2140,2173]" name="Vietnam" pageId="206" pageNumber="968">Vietnam</collectingCountry>
, it foraged in the air, similar to the Nepalese Whiskered
<taxonomicName id="03944D7DFF356A8AFAFE9E051B5CB34F" authority="Kaup, 1829" box="[1282,1372,2140,2173]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
(
<taxonomicName id="03944D7DFF356A8AFF489ED01F5BB394" baseAuthorityName="Gray" baseAuthorityYear="1846" box="[180,347,2185,2214]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="970" phylum="Chordata" rank="species" species="muricola">M. muricola</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF5A9EF21CD9B22E" pageId="206" pageNumber="968" type="breeding">
<paragraph id="C42B36FEFF356A8AFF5A9EF21CD9B22E" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF5A9EF21F2DB3FE" bold="true" box="[166,301,2219,2252]" pageId="206" pageNumber="968">Breeding.</emphasis>
In
<collectingCountry id="BC83766EFF356A8AFE979EF21FB5B3FE" box="[363,437,2219,2252]" name="India" pageId="206" pageNumber="968">India</collectingCountry>
(
<collectingRegion id="0650F81CFF356A8AFE339EF21CC6B3FE" box="[463,710,2219,2252]" country="India" name="Madhya Pradesh" pageId="206" pageNumber="968">Madhya Pradesh</collectingRegion>
), pregnant Horsfield’s
<taxonomicName id="03944D7DFF356A8AFBE79EF21A75B3FE" authority="Kaup, 1829" box="[1051,1141,2219,2252]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
were caught in February-March. In
<collectingCountry id="BC83766EFF356A8AFE2C9E8B1C47B3C1" box="[464,583,2258,2291]" name="Malaysia" pageId="206" pageNumber="968">Malaysia</collectingCountry>
, pregnant females were recorded in January-June and October; a few females had twin fetuses.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF599F781CA4B16A" pageId="206" pageNumber="968" type="activity">
<paragraph id="C42B36FEFF356A8AFF599F781CA4B16A" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF599F781F92B270" bold="true" box="[165,402,2337,2370]" pageId="206" pageNumber="968">Activity patterns.</emphasis>
Horsfield’s
<taxonomicName id="03944D7DFF356A8AFDBD9F781C9BB270" authority="Kaup, 1829" box="[577,667,2337,2370]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
is nocturnal. On Pulau Langkawi,
<collectingCountry id="BC83766EFF356A8AFB779F781B03B270" box="[1163,1283,2337,2370]" name="Malaysia" pageId="206" pageNumber="968">Malaysia</collectingCountry>
, it has been found roosting in limestone caves. In
<collectingCountry id="BC83766EFF356A8AFCE69F141D65B258" box="[794,869,2381,2410]" name="India" pageId="206" pageNumber="968">India</collectingCountry>
, it often roosts in culverts, tunnels, and houses and under bridges. In
<collectingRegion id="0650F81CFF356A8AFD5E9F291D90B2A3" box="[674,912,2416,2449]" country="India" name="Madhya Pradesh" pageId="206" pageNumber="968">Madhya Pradesh</collectingRegion>
, it was found in leaves of a palmyra palm. On the
<collectingCountry id="BC83766EFF356A8AFE479FCE1C5BB28A" box="[443,603,2455,2488]" name="Philippines" pageId="206" pageNumber="968">Philippines</collectingCountry>
, it roosted beneath a large rock over a stream. It might depend on caves near water sources; it is usually absent where streams are lacking. It disappears when water dries up. Echolocation calls are steeply FM without any CF component; frequencies are 45-100 kHz. Call characteristics imply that it can detect prey close to substrate or over water.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF549C041B27B059" pageId="206" pageNumber="968" type="biology_ecology">
<paragraph id="C42B36FEFF356A8AFF549C041B27B059" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF549C041D65B14C" bold="true" box="[168,869,2653,2686]" pageId="206" pageNumber="968">Movements, Home range and Social organization.</emphasis>
Horsfield’s
<taxonomicName id="03944D7DFF356A8AFBEE9C041A6CB14C" authority="Kaup, 1829" box="[1042,1132,2653,2686]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
is often the most common insectivorous bat species in some areas. It was the most frequently captured species in tunneltraps, representing 45% and 66% of tunnel trap captures on the west and south slopes of Mount Isarog, Luzon. It normally roosts in small (2-5) to mediumsized groups. One roost had more than 100 individuals. In
<collectingRegion id="0650F81CFF356A8AFC139CA21A4AB02E" box="[1007,1098,2811,2844]" country="India" name="Assam" pageId="206" pageNumber="968">Assam</collectingRegion>
, it was netted while coming out of a culvert where a group of 4-5 individuals was roosting. Horsfield’s
<taxonomicName id="03944D7DFF356A8AFF569D131F04B059" authority="Kaup, 1829" box="[170,260,2890,2923]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
often share roosts with species of Rhinolophus, Hipposideros, and
<taxonomicName id="03944D7DFF356A8AFB749D131B24B059" box="[1160,1316,2890,2923]" class="Mammalia" family="Miniopteridae" genus="Miniopterus" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Miniopterus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF569D281C14B76B" pageId="206" pageNumber="968" type="conservation">
<paragraph id="C42B36FEFF356A8AFF569D281C14B76B" blockId="206.[160,1379,285,3479]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF569D281C06B0A0" bold="true" box="[170,518,2929,2962]" pageId="206" pageNumber="968">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. Horsfield’s
<taxonomicName id="03944D7DFF356A8AFF569DC01F04B088" authority="Kaup, 1829" box="[170,260,2969,3002]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="206" pageNumber="968" phylum="Chordata" rank="genus">Myotis</taxonomicName>
is widespread, presumably has a large population, occurs in a number of protected areas, tolerates some habitat modification, and is unlikely to be declining fast enough to qualify for listing in a more threatened category. In South Asia,its habitat is being deforested for timber, firewood, and agriculture. It is also threatened by disturbance of roosting sites.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FF356A8AFF509A3E1687BA9D" pageId="206" pageNumber="968" type="bibRefCitation_list">
<paragraph id="C42B36FEFF356A8AFF509A3E1687BA9D" blockId="206.[160,1379,285,3479]" lastBlockId="206.[1435,2641,287,434]" pageId="206" pageNumber="968">
<emphasis id="F6E0EAECFF356A8AFF509A3E1F45B7B2" bold="true" box="[172,325,3175,3200]" pageId="206" pageNumber="968">Bibliography.</emphasis>
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</paragraph>
</subSubSection>
</treatment>
</document>
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