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<document id="9E62C46D47A7D34FC0D311A398AEA213" ID-CLB-Dataset="110554" ID-DOI="10.1649/0010-065X-69.mo4.167" ID-GBIF-Dataset="459316b8-ecf9-4bcb-a14b-100658d73012" ID-ISSN="1938-4394" ID-Zenodo-Dep="4911739" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623095475750" checkinUser="carolina" docAuthor="Flores, Gustavo E. &amp; Aballay, Fernando H." docDate="2015" docId="182B9F50FFEFFFDFFD71FE61FCE7FB71" docLanguage="en" docName="TheColeopBull.69.mo14.167-179.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Achanius (Ambigatus) Fairmaire 1892" docType="treatment" docVersion="7" lastPageNumber="174" masterDocId="E412E728FFE8FFD8FFDCFFAFFFCCFFEA" masterDocTitle="Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson" masterLastPageNumber="179" masterPageNumber="167" pageNumber="174" updateTime="1699002214655" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="74E729DDB4C746FE831F3D314390B28A">Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson</mods:title>
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<mods:namePart id="2DB37688F89EBFECBF32251E716B9F3C">Flores, Gustavo E.</mods:namePart>
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<mods:namePart id="21741AA3FD3B538C9B77A51AAE34EFFF">Aballay, Fernando H.</mods:namePart>
<mods:affiliation id="35547D5B2BEDAC4D8DF449EDBD79B19C">CONICET, Laboratorio de Entomología Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT-CONICET Mendoza) Casilla de correo 507, 5500 Mendoza, ARGENTINA</mods:affiliation>
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<mods:title id="BD37D60741DA214B1245B9F3544D8834">The Coleopterists Bulletin</mods:title>
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<mods:date id="42963EC52D8C0FDEA5AD5E237B35D4AA">2015</mods:date>
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<mods:title id="05504FCB92935F7E8D062A58155084B3">mo 14</mods:title>
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Subgenus
<taxonomicName id="578255C5FFEFFFDFFCCFFE61FB58FE09" authority="Fairmaire, 1892" authorityName="Fairmaire" authorityYear="1892" box="[787,1172,462,483]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFCCFFE61FCA2FE09" bold="true" box="[787,878,462,483]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFCA2FE61FC28FE09" bold="true" box="[894,996,462,483]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<bibRefCitation id="F41353B7FFEFFFDFFC28FE61FB58FE09" author="Fairmaire" box="[1012,1172,462,483]" firstAuthor="Fairmaire" pageId="7" pageNumber="174" pagination="242 - 253" refId="ref8723" refString="Fairmaire, L. 1892. Description de quelques Coleopteres argentins. Bulletin de la Societe Entomologique de Belgique 36 (3): 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
</taxonomicName>
</emphasis>
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<subSubSection id="D8987DCDFFEFFFDFFC93FE46FC3FFE14" box="[847,1011,488,510]" pageId="7" pageNumber="174" type="description">
<paragraph id="903D2E46FFEFFFDFFC93FE46FC3FFE14" blockId="7.[685,1172,462,510]" box="[847,1011,488,510]" pageId="7" pageNumber="174">
(
<figureCitation id="08B932C3FFEFFFDFFC8AFE46FC04FE14" box="[854,968,488,510]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="8.[197,238,1662,1681]" captionStartId-1="9.[175,216,981,1000]" captionTargetBox-0="[208,1149,201,1640]" captionTargetBox-1="[151,1171,194,957]" captionTargetId-0="figure-1@8.[190,1176,183,1645]" captionTargetId-1="figure-1@9.[151,1173,193,957]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figs. 1316. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." captionText-1="Figs. 1720. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi-0="http://doi.org/10.5281/zenodo.4911747" figureDoi-1="http://doi.org/10.5281/zenodo.4911749" httpUri-0="https://zenodo.org/record/4911747/files/figure.png" httpUri-1="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Figs. 1420</figureCitation>
,
<figureCitation id="08B932C3FFEFFFDFFC08FE46FC20FE14" box="[980,1004,489,510]" captionStart="Figs" captionStartId="9.[175,215,1662,1681]" captionTargetBox="[151,1171,1074,1639]" captionTargetId="figure-36@9.[151,1173,1074,1640]" captionTargetPageId="9" captionText="Figs. 2122. Ovipositor in ventral view, spiculum and internal female reproductive tract. 21) Vaniosus profana; 22) Achanius (Ambigatus) rufonitens. Abbreviations: c = coxite; o = oviduct; p = paraproct; s = spermatheca; sag = spermathecal accessory gland; sp = spiculum; v = vagina. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911753" httpUri="https://zenodo.org/record/4911753/files/figure.png" pageId="7" pageNumber="174">22</figureCitation>
)
</paragraph>
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<subSubSection id="D8987DCDFFEFFFDFFD71FDBBFC4AFD92" pageId="7" pageNumber="174" type="reference_group">
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<treatmentCitationGroup id="B0920968FFEFFFDFFD71FDBBFC4AFD92" pageId="7" pageNumber="174">
<treatmentCitation id="11230857FFEFFFDFFD71FDBBFC29FDC2" author="Fairmaire" box="[685,997,531,553]" page="246" pageId="7" pageNumber="174" year="1892">
<taxonomicName id="578255C5FFEFFFDFFD71FDBBFC29FDC2" authority="Fairmaire 1892: 246" authorityName="Fairmaire" authorityPageNumber="246" authorityYear="1892" box="[685,997,531,553]" class="Insecta" family="Tenebrionidae" genus="Ambigatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis id="A2F6F254FFEFFFDFFD71FDBBFCDFFDC3" box="[685,787,532,553]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
<bibRefCitation id="F41353B7FFEFFFDFFCC7FDBCFC29FDC2" author="Fairmaire" box="[795,997,531,552]" firstAuthor="Fairmaire" page="246" pageId="7" pageNumber="174" pagination="242 - 253" refId="ref8723" refString="Fairmaire, L. 1892. Description de quelques Coleopteres argentins. Bulletin de la Societe Entomologique de Belgique 36 (3): 242 - 253." type="journal article" year="1892">Fairmaire 1892: 246</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="11230857FFEFFFDFFC2CFDBCFB58FDC2" author="Gebien &amp; Tenebrionidae II &amp; In &amp; Coleopterorum Catalogus XVII" box="[1008,1172,531,552]" page="81" pageId="7" pageNumber="174" year="1910">
<bibRefCitation id="F41353B7FFEFFFDFFC2CFDBCFB58FDC2" author="Gebien" box="[1008,1172,531,552]" firstAuthor="Gebien" page="81" pageId="7" pageNumber="174" pagination="167 - 354" refId="ref9004" refString="Gebien, H. 1910. Tenebrionidae II [pp. 167 - 354]. In: Coleopterorum Catalogus XVII. Pars 22. (W. Junk and S. Schenkling, editors). W. Junk, Berlin, Germany." type="journal article" year="1910">Gebien 1910: 81</bibRefCitation>
</treatmentCitation>
(catalog), 1937: 579 (catalog);
<treatmentCitation id="11230857FFEFFFDFFC2DFD81FB58FDA9" author="Bruch" box="[1009,1172,558,579]" page="264" pageId="7" pageNumber="174" year="1915">
<bibRefCitation id="F41353B7FFEFFFDFFC2DFD81FB58FDA9" author="Bruch" box="[1009,1172,558,579]" firstAuthor="Bruch" page="264" pageId="7" pageNumber="174" pagination="235 - 302" refId="ref8543" refString="Bruch, C. 1915. Catalogo sistematico de los coleopteros de la Republica Argentina. Pars VI. Revista del Museo de La Plata Serie 2 (19): 235 - 302." type="journal article" year="1915">Bruch 1915: 264</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFEFFFDFFCEFFDE6FBEBFDB4" author="Blackwelder" box="[819,1063,585,606]" page="511" pageId="7" pageNumber="174" year="1945">
<bibRefCitation id="F41353B7FFEFFFDFFCEFFDE6FBEBFDB4" author="Blackwelder" box="[819,1063,585,606]" firstAuthor="Blackwelder" page="511" pageId="7" pageNumber="174" pagination="343 - 550" refId="ref8449" refString="Blackwelder, R. E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 511</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFEFFFDFFD1BFDCCFC4EFD92" author="Doyen" box="[711,898,611,632]" page="500" pageId="7" pageNumber="174" year="1994">
<bibRefCitation id="F41353B7FFEFFFDFFD1BFDCCFC4EFD92" author="Doyen" box="[711,898,611,632]" firstAuthor="Doyen" page="500" pageId="7" pageNumber="174" pagination="443 - 514" refId="ref8578" refString="Doyen, J. T. 1994. Cladistic relationships among Pimeliinae Tenebrionidae (Coleoptera). Journal of the New York Entomological Society 101: 443 - 514." type="journal article" year="1994">Doyen 1994: 500</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFC4CFDCCFC58FD22" pageId="7" pageNumber="174" type="type_taxon">
<paragraph id="903D2E46FFEFFFDFFC4CFDCCFC15FD79" blockId="7.[685,1172,531,712]" pageId="7" pageNumber="174">
<typeStatus id="4F3990E4FFEFFFDFFC4CFDCCFC0EFD92" box="[912,962,611,632]" pageId="7" pageNumber="174">Type</typeStatus>
species:
<taxonomicName id="578255C5FFEFFFDFFBFBFDCCFC19FD79" authority="Fairmaire, 1892" authorityName="Fairmaire" authorityYear="1892" class="Insecta" family="Tenebrionidae" genus="Ambigatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="stricticollis">
<emphasis id="A2F6F254FFEFFFDFFBFBFDCCFCFDFD79" italics="true" pageId="7" pageNumber="174">Ambigatus stricticollis</emphasis>
<bibRefCitation id="F41353B7FFEFFFDFFCE5FDD1FC19FD79" author="Fairmaire" box="[825,981,638,659]" firstAuthor="Fairmaire" pageId="7" pageNumber="174" pagination="242 - 253" refId="ref8723" refString="Fairmaire, L. 1892. Description de quelques Coleopteres argentins. Bulletin de la Societe Entomologique de Belgique 36 (3): 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="903D2E46FFEFFFDFFD71FD36FC58FD22" blockId="7.[685,1172,531,712]" pageId="7" pageNumber="174">
<treatmentCitationGroup id="B0920968FFEFFFDFFD71FD36FC58FD22" pageId="7" pageNumber="174">
<taxonomicName id="578255C5FFEFFFDFFD71FD36FC48FD47" authority="" authorityName="Fairmaire" authorityYear="1892" box="[685,900,664,686]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFD71FD36FCC9FD44" box="[685,773,665,686]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFCC8FD36FCB6FD44" box="[788,890,665,686]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="11230857FFEFFFDFFC53FD37FBE7FD47" author="Kulzer" box="[911,1067,664,685]" page="31" pageId="7" pageNumber="174" year="1950">
<bibRefCitation id="F41353B7FFEFFFDFFC53FD37FBE7FD47" author="Kulzer" box="[911,1067,664,685]" firstAuthor="Kulzer" page="31" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">Kulzer 1950: 31</bibRefCitation>
</treatmentCitation>
(revision), 1956: 901 (revision).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFD19FD72FC83FC9D" pageId="7" pageNumber="174" type="diagnosis">
<paragraph id="903D2E46FFEFFFDFFD19FD72FC83FC9D" blockId="7.[685,1173,733,1179]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFD19FD72FCF8FD18" bold="true" box="[709,820,733,754]" pageId="7" pageNumber="174">Diagnosis.</emphasis>
<bibRefCitation id="F41353B7FFEFFFDFFC9FFD72FC02FD18" author="Kulzer" box="[835,974,733,754]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">
Kulzer (
<quantity id="577A83A3FFEFFFDFFC46FD72FC02FD18" box="[922,974,733,754]" metricMagnitude="1" metricUnit="m" metricValue="4.953" pageId="7" pageNumber="174" unit="in" value="1950.0">1950</quantity>
</bibRefCitation>
in key) stated the humeri are not elevated and hind wings are present. This study: metasternum 3X length of mesosternum; metacoxal width 3X metacoxal length; distance between meso and metacoxae 2.5X metacoxal length (
<figureCitation id="08B932C3FFEFFFDFFD25FCCDFC88FC9D" box="[761,836,866,887]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 1720. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFD19FCD2FB5EFBFD" pageId="7" pageNumber="174" type="biology_ecology">
<paragraph id="903D2E46FFEFFFDFFD19FCD2FB5EFBFD" blockId="7.[685,1173,733,1179]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFD19FCD2FCDBFC78" bold="true" box="[709,791,893,914]" pageId="7" pageNumber="174">Biology.</emphasis>
Species of
<taxonomicName id="578255C5FFEFFFDFFC48FCD2FBA2FC78" authority="" authorityName="Fairmaire" authorityYear="1892" box="[916,1134,893,914]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFC48FCD2FC22FC78" box="[916,1006,893,914]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFC22FCD2FBA9FC78" box="[1022,1125,893,914]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
</taxonomicName>
are winged, capable of flying, and of crepuscular and nocturnal habits. They were collected at white and UV lights, also using pitfall traps partly filled with water and propylene glycol (
<bibRefCitation id="F41353B7FFEFFFDFFC09FC48FB5CFC16" author="Flores" box="[981,1168,999,1020]" etAl="et al." firstAuthor="Flores" pageId="7" pageNumber="174" pagination="71 - 90" refId="ref8833" refString="Flores, G. E., S. J. Lagos, and S. Roig-Junent. 2004. Artropodos epigeos que viven bajo la copa del algarrobo (Prosopis flexuosa) en la Reserva Telteca (Mendoza, Argentina). Multequina 13: 71 - 90." type="journal article" year="2004">
<collectingCountry id="E8956ED6FFEFFFDFFC09FC48FBD8FC16" box="[981,1044,999,1020]" name="Indonesia" pageId="7" pageNumber="174">Flores</collectingCountry>
<emphasis id="A2F6F254FFEFFFDFFBC1FC48FB87FC16" box="[1053,1099,999,1020]" italics="true" pageId="7" pageNumber="174">et al</emphasis>
. 2004
</bibRefCitation>
;
<bibRefCitation id="F41353B7FFEFFFDFFD71FBADFCE9FBFD" author="Lagos" box="[685,805,1026,1047]" firstAuthor="Lagos" pageId="7" pageNumber="174" refId="ref9312" refString="Lagos, S. J. 2004. Diversidad biologica de las comunidades epigeas de artropodos en areas pastoreadas y no pastoreadas del Monte (Argentina). PhD dissertation, Programa de Postgrado en Ciencias Biologicas (PROBIOL), Universidad Nacional de Cuyo, Mendoza, Argentina." type="book" year="2004">Lagos 2004</bibRefCitation>
), and cattle dung-baited pitfall traps.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFD19FBB3FCE7FB71" pageId="7" pageNumber="174" type="discussion">
<paragraph id="903D2E46FFEFFFDFFD19FBB3FCE7FB71" blockId="7.[685,1173,733,1179]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFD19FBB3FC4AFBDB" bold="true" box="[709,902,1052,1073]" pageId="7" pageNumber="174">Species Included.</emphasis>
<taxonomicName id="578255C5FFEFFFDFFC48FBB3FBA7FBDB" box="[916,1131,1052,1073]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="rufonitens">
<emphasis id="A2F6F254FFEFFFDFFC48FBB3FBA7FBDB" box="[916,1131,1052,1073]" italics="true" pageId="7" pageNumber="174">Achanius rufonitens</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFBA2FBB3FCD8FBA6" authorityName="Fairmaire" authorityYear="1892" class="Insecta" family="Tenebrionidae" genus="Ambigatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="stricticollis">
<emphasis id="A2F6F254FFEFFFDFFBA2FBB3FCD8FBA6" italics="true" pageId="7" pageNumber="174">A. stricticollis</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFCFDFB98FCB1FBA6" box="[801,893,1079,1100]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A2F6F254FFEFFFDFFCFDFB98FCB1FBA6" box="[801,893,1079,1100]" italics="true" pageId="7" pageNumber="174">A. bicolor</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFC55FB98FBE0FBA6" box="[905,1068,1079,1100]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="rhinosomoides">
<emphasis id="A2F6F254FFEFFFDFFC55FB98FBE0FBA6" box="[905,1068,1079,1100]" italics="true" pageId="7" pageNumber="174">A. rhinosomoides</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFBE4FB98FB41FBA6" box="[1080,1165,1079,1100]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="A2F6F254FFEFFFDFFBE4FB98FB41FBA6" box="[1080,1165,1079,1100]" italics="true" pageId="7" pageNumber="174">A. piceus</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFD71FBFDFCD3FB8D" box="[685,799,1106,1127]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="A2F6F254FFEFFFDFFD71FBFDFCD3FB8D" box="[685,799,1106,1127]" italics="true" pageId="7" pageNumber="174">A. wittmeri</emphasis>
</taxonomicName>
(
<bibRefCitation id="F41353B7FFEFFFDFFCF3FBFEFC7EFB8C" author="Kulzer" box="[815,946,1105,1126]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">Kulzer 1950</bibRefCitation>
,
<bibRefCitation id="F41353B7FFEFFFDFFC1CFBFEFC39FB8C" author="Kulzer" box="[960,1013,1105,1126]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">1956</bibRefCitation>
), and
<taxonomicName id="578255C5FFEFFFDFFBEBFBFDFC0BFB6B" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="antofagastensis" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFBEBFBFDFB58FB8D" box="[1079,1172,1106,1127]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFD69FBC3FCD1FB6B" box="[693,797,1132,1153]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFCF3FBC3FC0BFB6B" box="[815,967,1132,1153]" italics="true" pageId="7" pageNumber="174">antofagastensis</emphasis>
</taxonomicName>
<collectingCountry id="E8956ED6FFEFFFDFFC0CFBC3FBC1FB6B" box="[976,1037,1132,1153]" name="Indonesia" pageId="7" pageNumber="174">Flores</collectingCountry>
and Aballay,
<taxonomicNameLabel id="B9C54F2FFFEFFFDFFD71FB29FCEBFB71" box="[685,807,1158,1179]" pageId="7" pageNumber="174" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>