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<document id="AA0A4772541478524BA21C5F15A7157D" ID="10.11646/zootaxa.4282.1.4SLASH11465" ID-CLB-Dataset="28726" ID-DOI="10.11646/zootaxa.4282.1.4" ID-GBIF-Dataset="e901e52f-3189-4226-bb52-3ffed859d5a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="818445" ID-ZooBank="473BB1D9-438F-47FF-9717-D95EE950412E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1498474875696" checkinUser="plazi" docAuthor="Haber, William A." docDate="2017" docId="0F7B8794FFB8FFE8AFC1E2AC041D9308" docLanguage="en" docName="zootaxa.4282.1.4SLASH11465.pdf" docOrigin="Zootaxa 4282 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epigomphus morrisoni Haber 2017, new species" docType="treatment" docVersion="7" lastPageNumber="88" masterDocId="F342FFECFFB6FFE7AF56E44C055A9574" masterDocTitle="Three new species of Epigomphus (Odonata: Gomphidae) from Costa Rica" masterLastPageNumber="94" masterPageNumber="73" pageNumber="87" updateTime="1698449431867" updateUser="ExternalLinkService">
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<mods:title id="319D81658489538E8388D85EEB2DDF74">Three new species of Epigomphus (Odonata: Gomphidae) from Costa Rica</mods:title>
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<mods:title id="2017FAC720863E118712CEF878ACEB2E">Zootaxa</mods:title>
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<mods:date id="63E84FFC7A982ADBF6750217CBB77D08">2017</mods:date>
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<treatment id="0F7B8794FFB8FFE8AFC1E2AC041D9308" ID-DOI="http://doi.org/10.5281/zenodo.5999729" ID-GBIF-Taxon="150795646" ID-Zenodo-Dep="5999729" LSID="urn:lsid:plazi:treatment:0F7B8794FFB8FFE8AFC1E2AC041D9308" httpUri="http://treatment.plazi.org/id/0F7B8794FFB8FFE8AFC1E2AC041D9308" lastPageId="15" lastPageNumber="88" pageId="14" pageNumber="87">
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<paragraph id="876D3682FFB8FFE9AFC1E2AC071F938F" blockId="14.[151,581,1760,1821]" box="[151,581,1760,1787]" pageId="14" pageNumber="87">
<heading id="DC2581EEFFB8FFE9AFC1E2AC071F938F" bold="true" box="[151,581,1760,1787]" fontSize="11" level="1" pageId="14" pageNumber="87" reason="1">
<emphasis id="B5A6EA90FFB8FFE9AFC1E2AC071F938F" bold="true" box="[151,581,1760,1787]" pageId="14" pageNumber="87">
<taxonomicName id="40D24D01FFB8FFE9AFC1E2AC04F4938E" ID-CoL="9X2MM" authority="Haber, 2017" authorityName="Haber" authorityYear="2017" box="[151,430,1760,1786]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="87" phylum="Arthropoda" rank="species" species="morrisoni" status="sp. nov.">
<emphasis id="B5A6EA90FFB8FFE9AFC1E2AC04F4938E" bold="true" box="[151,430,1760,1786]" italics="true" pageId="14" pageNumber="87">Epigomphus morrisoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AE9557EBFFB8FFE9AEE3E2AD071F938F" box="[437,581,1761,1787]" pageId="14" pageNumber="87" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="876D3682FFB8FFE9AFC1E34807669268" blockId="14.[151,581,1760,1821]" box="[151,572,1796,1821]" pageId="14" pageNumber="87">
<figureCitation id="1FE92A07FFB8FFE9AFC1E34805A49268" box="[151,254,1796,1821]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1536]" captionTargetBox="[159,1423,223,1470]" captionTargetId="figure@2.[151,1436,195,1490]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus bosquenuboso: lateral scans of a) male; b) female. E. morrisoni: c) lateral scan of male. E. wagneri: d) lateral scan of male." httpUri="https://zenodo.org/record/818447/files/figure.png" pageId="14" pageNumber="87">Figures 1</figureCitation>
c; 3b; 4b; 5c, d; 9b; 10b; 12.
</paragraph>
</subSubSection>
<subSubSection id="CFC86509FFB8FFE9AFC1E30006D592FC" pageId="14" pageNumber="87" type="description">
<paragraph id="876D3682FFB8FFE9AFC1E30006D592FC" blockId="14.[151,1436,1868,2037]" pageId="14" pageNumber="87">
<materialsCitation id="37BA3CDFFFB8FFE9AFC1E30006D592FC" ID-GBIF-Occurrence="1950907567" collectingDate="2010-06-22" collectionCode="INBIO, INB" country="Costa Rica" county="Specimens" elevation="725" location="Rio Guacimal" municipality="Monteverde" pageId="14" pageNumber="87" specimenCode="INB0004319187" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<emphasis id="B5A6EA90FFB8FFE9AFC1E30007139211" bold="true" box="[151,585,1868,1893]" pageId="14" pageNumber="87">
<collectingCounty id="6E0C4E0EFFB8FFE9AFC1E300044F9211" box="[151,277,1868,1893]" pageId="14" pageNumber="87">Specimens</collectingCounty>
examined.
<typeStatus id="58698820FFB8FFE9AECBE30007509211" box="[413,522,1868,1893]" pageId="14" pageNumber="87" type="holotype">Holotype</typeStatus>
male
</emphasis>
:
<collectingCountry id="FFC57612FFB8FFE9AD0EE30007A49210" box="[600,766,1868,1892]" name="Costa Rica" pageId="14" pageNumber="87">COSTA RICA</collectingCountry>
, Provincia
<collectingRegion id="4516F860FFB8FFE9AC2DE30106A39210" box="[891,1017,1869,1892]" country="Costa Rica" name="Puntarenas" pageId="14" pageNumber="87">Puntarenas</collectingRegion>
,
<collectingMunicipality id="6709ACF8FFB8FFE9AB52E30001D49211" box="[1028,1166,1868,1893]" pageId="14" pageNumber="87">Monteverde</collectingMunicipality>
,
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(10.28, - 84.82),
<elevation id="0CFFD1B1FFB8FFE9AFBBE323046C92F3" box="[237,310,1903,1928]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="14" pageNumber="87" unit="m" value="725.0">
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,
<collectingDate id="E328E9AAFFB8FFE9AE14E33C048092FC" box="[322,474,1904,1928]" pageId="14" pageNumber="87" value="2010-06-22">
<date id="F36C1042FFB8FFE9AE14E33C048092FC" box="[322,474,1904,1928]" pageId="14" pageNumber="87" value="2010-06-22">22 June 2010</date>
</collectingDate>
, leg. WAH (
<collectionCode id="E1C3AE47FFB8FFE9AD32E33C07EC92FC" box="[612,694,1904,1928]" pageId="14" pageNumber="87">INBIO</collectionCode>
,
<specimenCode id="D7749EF9FFB8FFE9AD97E33C06DE92FC" box="[705,900,1904,1928]" collectionCode="INB" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14937" name="Instituto Nacional de Biodiversidad" pageId="14" pageNumber="87">
<collectionCode id="E1C3AE47FFB8FFE9AD97E33C07AB92FC" box="[705,753,1904,1928]" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14937" name="Instituto Nacional de Biodiversidad" pageId="14" pageNumber="87">INB</collectionCode>
0004319187
</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CFC86509FFB8FFE9AF91E3D805A09281" pageId="14" pageNumber="87" type="etymology">
<paragraph id="876D3682FFB8FFE9AF91E3D805A09281" blockId="14.[151,1436,1868,2037]" pageId="14" pageNumber="87">
<emphasis id="B5A6EA90FFB8FFE9AF91E3D8041492D9" bold="true" box="[199,334,1940,1965]" pageId="14" pageNumber="87">Etymology.</emphasis>
This species is named for my friend and colleague Fredric Morrison, a great naturalist, devotee of the
<taxonomicName id="40D24D01FFB8FFE9AF96E3F4041C92A5" authorityName="Rambur" authorityYear="1842" box="[192,326,1976,2001]" class="Insecta" family="Gomphidae" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="87" phylum="Arthropoda" rank="family">Gomphidae</taxonomicName>
, and trusted companion who has accompanied me on many
<taxonomicName id="40D24D01FFB8FFE9ACBEE3F4011D92A5" authorityName="Fabricius" authorityYear="1793" box="[1000,1095,1976,2001]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="87" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
expeditions in
<collectingCountry id="FFC57612FFB8FFE9ABA5E3F4003792A5" box="[1267,1389,1976,2001]" name="Costa Rica" pageId="14" pageNumber="87">Costa Rica</collectingCountry>
and
<collectingCountry id="FFC57612FFB8FFE9AFC1E39005AD9281" box="[151,247,2012,2037]" name="Ecuador" pageId="14" pageNumber="87">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CFC86509FFB9FFE8AF91E4DB04F89047" pageId="15" pageNumber="88" type="description">
<paragraph id="876D3682FFB9FFE8AF91E4DB007A94FC" blockId="15.[151,1437,151,1661]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E4DB073895C4" bold="true" box="[199,610,151,176]" pageId="15" pageNumber="88">
Description. Male
<typeStatus id="58698820FFB9FFE8AEF3E4DB075495C4" box="[421,526,151,176]" pageId="15" pageNumber="88" type="holotype">holotype</typeStatus>
. Head.
</emphasis>
Maximum width
<quantity id="402A9B67FFB9FFE8AC79E4D406D395DB" box="[815,905,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="15" pageNumber="88" unit="mm" value="7.2">7.2 mm</quantity>
; shiny dark brown above (
<figureCitation id="1FE92A07FFB9FFE8ABEEE4DB01A095C4" box="[1208,1274,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1536]" captionTargetBox="[159,1423,223,1470]" captionTargetId="figure@2.[151,1436,195,1490]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus bosquenuboso: lateral scans of a) male; b) female. E. morrisoni: c) lateral scan of male. E. wagneri: d) lateral scan of male." httpUri="https://zenodo.org/record/818447/files/figure.png" pageId="15" pageNumber="88">Fig. 1</figureCitation>
c, 9b); labrum dark brown with a blue-gray spot near lateral margin; base of mandible and gena pale; top of frons with a pale stripe covering about 2/3 of surface, narrowed in middle and divided by a dark line; antennae dark brown; eyes blue in life; lateral ocelli equidistant from eye margin and mid line; postocellar surface nearly flat, barely impressed; posterior margin of occiput with a rounded ridge, a pair of lateral conical tubercles on dorsum that are collapsed in
<typeStatus id="58698820FFB9FFE8AE70E50004D19411" box="[294,395,332,357]" pageId="15" pageNumber="88" type="holotype">holotype</typeStatus>
, rear margin with central third indented, a pair of low rounded tubercles on posterior side of ridge close to eye margins; underside of head dark brown on upper half shading to cream on lower half.
</paragraph>
<paragraph id="876D3682FFB9FFE8AF91E5D8073396F4" blockId="15.[151,1437,151,1661]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E5D8047C94D9" bold="true" box="[199,294,404,429]" pageId="15" pageNumber="88">Thorax.</emphasis>
dark brown with pattern of stripes similar to
<taxonomicName id="40D24D01FFB9FFE8AC6FE5D9068294D8" authorityName="Selys, 1878" authorityYear="1878" box="[825,984,404,428]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8AC6FE5D9068294D8" box="[825,984,404,428]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
(
<figureCitation id="1FE92A07FFB9FFE8ACBFE5D8017694D8" box="[1001,1068,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1536]" captionTargetBox="[159,1423,223,1470]" captionTargetId="figure@2.[151,1436,195,1490]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus bosquenuboso: lateral scans of a) male; b) female. E. morrisoni: c) lateral scan of male. E. wagneri: d) lateral scan of male." httpUri="https://zenodo.org/record/818447/files/figure.png" pageId="15" pageNumber="88">Fig. 1</figureCitation>
c; 9b). Prothorax fore lobe with anterior margin pale; mid lobe with a pair of contiguous pale spots in center and a pair of larger pale spots on lateral ends. Pterothorax: mesepisternum with two pale antehumeral stripes, upper stripe separate from pale collar stripe and not reaching posterior margin, lower stripe narrowed in middle; mesepimeron with a complete central stripe, narrowed posteriorly, then widened at end; metepisternal pale stripe broad at base, narrowed to a point and almost separated into a posterior spot; metepimeron pale except for a narrow brown upper border and a dark central bar on anterior half; mesinfraepisternum with a pale triangular patch at lower posterior corner; metinfraepisternum mostly pale, shading to brown in upper third; metasternum pale, unmarked.
<emphasis id="B5A6EA90FFB9FFE8ACEAE6C306AE97DC" bold="true" box="[956,1012,655,680]" pageId="15" pageNumber="88">Legs</emphasis>
: fore and mid legs with femur dark brown on anterior side, shading to light brown on posterior side; no pale stripe on inner surface of fore femur; hind femur pale brown, dark only at tip.
<emphasis id="B5A6EA90FFB9FFE8AD64E69B07249784" bold="true" box="[562,638,727,752]" pageId="15" pageNumber="88">Wings</emphasis>
hyaline with dark brown venation (
<figureCitation id="1FE92A07FFB9FFE8AB4FE69B010D9784" box="[1049,1111,727,752]" captionStart="FIGURE 3" captionStartId="4.[151,250,1989,2011]" captionTargetBox="[254,1323,206,1953]" captionTargetId="figure@4.[241,1340,193,1963]" captionTargetPageId="4" captionText="FIGURE 3. Wings of male Epigomphus, dorsal view: a) E. bosquenuboso, b) E. morrisoni, c) E. wagneri." httpUri="https://zenodo.org/record/818451/files/figure.png" pageId="15" pageNumber="88">Fig 3</figureCitation>
b); basal subcostal crossvein present; cubito-anal crossveins
<quantity id="402A9B67FFB9FFE8AEAEE6B0077E9661" box="[504,548,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="15" pageNumber="88" unit="in" value="3.0">3 in</quantity>
FW,
<quantity id="402A9B67FFB9FFE8AD36E6B007D69661" box="[608,652,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="88" unit="in" value="2.0">2 in</quantity>
HW; FW Ax 19, 17 with 7 thickened; Px 13, 14; HW Ax 14, 13 with 7, 6 thickened; Px 13, 12; pterostigma dark brown, opaque,
<quantity id="402A9B67FFB9FFE8AC0FE75306E99643" box="[857,947,799,824]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="15" pageNumber="88" unit="mm" value="3.6">3.6 mm</quantity>
in FW,
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in HW; crossveins above supratriangle between arculus and point of branching of RP 5, 6 in FW, 3, 4 in HW; post-triangular cell space in FW with two rows of cells for 10, 9 cells.
</paragraph>
<paragraph id="876D3682FFB9FFE8AF91E7C005B69164" blockId="15.[151,1437,151,1661]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E7C0046696D1" bold="true" box="[199,316,908,933]" pageId="15" pageNumber="88">Abdomen</emphasis>
dark brown with pale markings cream-yellow; dorsum of S14 with a pale dorsal line (widened on S12); sides of S3 pale on basal 3/4; sides of S46 with pale basal triangles; S7 pale in basal 3/4; side of S8 with a tiny pale spot on lower basal corner; S9 unmarked; S10 with a low hump barely divided by a central longitudinal groove.
</paragraph>
<paragraph id="876D3682FFB9FFE8AF91E050075D91B3" blockId="15.[151,1437,151,1661]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E05004B39141" bold="true" box="[199,489,1052,1077]" pageId="15" pageNumber="88">Abdominal appendages.</emphasis>
Cercus as in
<taxonomicName id="40D24D01FFB9FFE8ADDFE051067D9140" authorityName="Selys, 1878" authorityYear="1878" box="[649,807,1052,1076]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8ADDFE051067D9140" box="[649,807,1052,1076]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
;
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long along dorsal line, apex with line of 78 small teeth; epiproct
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along dorsal line, slightly shorter than cercus, similar in shape to
<taxonomicName id="40D24D01FFB9FFE8ABA3E00D00CE912C" authorityName="Selys, 1878" authorityYear="1878" box="[1269,1428,1088,1112]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8ABA3E00D00CE912C" box="[1269,1428,1088,1112]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
, except apex squared off and bearing a strong dorsally directed tooth, which is slightly recurved at tip and bent slightly inward; sinus between branches 95o; on dorsal side, margin of sinus with a rounded bump extending about 3/4 of sinus length (
<figureCitation id="1FE92A07FFB9FFE8AE23E0E204E991B3" box="[373,435,1198,1223]" captionStart="FIGURE 4" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[159,1408,205,1820]" captionTargetId="figure@5.[151,1436,193,1840]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal appendages of Epigomphus species: a) E. bosquenuboso lateral view, ventral view; b) E. morrisoni, lateral view, ventral view; c) E. wagneri lateral view, ventral view." httpUri="https://zenodo.org/record/818453/files/figure.png" pageId="15" pageNumber="88">Fig 4</figureCitation>
c; 10b).
</paragraph>
<paragraph id="876D3682FFB9FFE8AF91E09E0197919F" blockId="15.[151,1437,151,1661]" box="[199,1229,1234,1259]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E09E04E0919F" bold="true" box="[199,442,1234,1259]" pageId="15" pageNumber="88">Secondary genitalia.</emphasis>
Vesica spermalis similar in shape to that of
<taxonomicName id="40D24D01FFB9FFE8ACF1E09F0119919F" authorityName="Selys, 1878" authorityYear="1878" box="[935,1091,1235,1259]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8ACF1E09F0119919F" box="[935,1091,1235,1259]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
(
<figureCitation id="1FE92A07FFB9FFE8AB04E09E01CF919F" box="[1106,1173,1234,1259]" captionStart="FIGURE 5" captionStartId="6.[151,250,1613,1635]" captionTargetBox="[182,1401,230,1561]" captionTargetId="figure@6.[151,1436,193,1591]" captionTargetPageId="6" captionText="FIGURE 5. Distal segment of vesica spermalis of Epigomphus species with lateral view on left, dorsal view on right. E. bosquenuboso: a, b; E. morrisoni: c, d; E. wagneri: e, f. E. bosquenuboso female: g) dorsal view of head; h) vulvar lamina." httpUri="https://zenodo.org/record/818455/files/figure.png" pageId="15" pageNumber="88">Fig. 5</figureCitation>
c, d).
</paragraph>
<paragraph id="876D3682FFB9FFE8AF91E0BA0198907B" blockId="15.[151,1437,151,1661]" box="[199,1218,1270,1295]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E0BA049F907B" bold="true" box="[199,453,1270,1295]" pageId="15" pageNumber="88">Measurements (mm).</emphasis>
Total length 50; abdomen 38; FW 35; HW 33, maximum width 8.2.
</paragraph>
<paragraph id="876D3682FFB9FFE8AF91E15604F89047" blockId="15.[151,1437,151,1661]" box="[199,418,1306,1331]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E156047F9047" bold="true" box="[199,293,1306,1331]" pageId="15" pageNumber="88">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="CFC86509FFB9FFE8AF91E10D041D9308" pageId="15" pageNumber="88" type="diagnosis">
<paragraph id="876D3682FFB9FFE8AF91E10D01DC90D1" blockId="15.[151,1437,151,1661]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E10D041B902E" bold="true" box="[199,321,1345,1370]" pageId="15" pageNumber="88">Diagnosis.</emphasis>
Thoracic pattern and cercus shape as in
<taxonomicName id="40D24D01FFB9FFE8AC40E10E06E9902D" authorityName="Selys, 1878" authorityYear="1878" box="[790,947,1345,1369]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8AC40E10E06E9902D" box="[790,947,1345,1369]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
, branches of epiproct forming a 95o inside angle compared with 6478o in
<taxonomicName id="40D24D01FFB9FFE8AD5CE12507F090F4" authorityName="Selys, 1878" authorityYear="1878" box="[522,682,1384,1408]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8AD5CE12507F090F4" box="[522,682,1384,1408]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
(mean 72.4 o, s.d. 4.97, n=10), apex of epiproct with a strongly developed dorsal hook that is inclined slightly inward (slightly outward in
<taxonomicName id="40D24D01FFB9FFE8AC8DE1C1012C90D0" authorityName="Selys, 1878" authorityYear="1878" box="[987,1142,1420,1444]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8AC8DE1C1012C90D0" box="[987,1142,1420,1444]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="876D3682FFB9FFE8AF91E1FC041D9308" blockId="15.[151,1437,151,1661]" pageId="15" pageNumber="88">
<emphasis id="B5A6EA90FFB9FFE8AF91E1FC04DB90BD" bold="true" box="[199,385,1456,1481]" pageId="15" pageNumber="88">Natural history</emphasis>
. To date
<taxonomicName id="40D24D01FFB9FFE8AEA5E1FD07D990BC" authorityName="Haber" authorityYear="2017" box="[499,643,1457,1480]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="morrisoni">
<emphasis id="B5A6EA90FFB9FFE8AEA5E1FD07D990BC" box="[499,643,1457,1480]" italics="true" pageId="15" pageNumber="88">E. morrisoni</emphasis>
</taxonomicName>
is known only from one male from the
<typeStatus id="58698820FFB9FFE8AB37E1FD01CB90BD" box="[1121,1169,1457,1481]" pageId="15" pageNumber="88">type</typeStatus>
locality. No additional individuals were found in many subsequent visits to the area. The
<typeStatus id="58698820FFB9FFE8ACF0E199068C9099" box="[934,982,1493,1517]" pageId="15" pageNumber="88">type</typeStatus>
specimen was a mature male that was perched (apparently defending a territory) on an upright dead stick projecting from a shallow, partly shaded pool in a spring outflow at the interface of a forest and pasture about 11:00 hr. Larvae and adults of
<taxonomicName id="40D24D01FFB9FFE8ABF9E25100169340" authorityName="Selys, 1878" authorityYear="1878" box="[1199,1356,1564,1588]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B5A6EA90FFB9FFE8ABF9E25100169340" box="[1199,1356,1564,1588]" italics="true" pageId="15" pageNumber="88">E. subobtusus</emphasis>
</taxonomicName>
and
<taxonomicName id="40D24D01FFB9FFE8AAD2E251044B932C" authorityName="Calvert" authorityYear="1903" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="tumefactus">
<emphasis id="B5A6EA90FFB9FFE8AAD2E251044B932C" italics="true" pageId="15" pageNumber="88">E. tumefactus</emphasis>
</taxonomicName>
were collected at the same pool, but no larvae that might have been
<taxonomicName id="40D24D01FFB9FFE8AB59E20D01C6932C" authorityName="Haber" authorityYear="2017" box="[1039,1180,1601,1624]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="morrisoni">
<emphasis id="B5A6EA90FFB9FFE8AB59E20D01C6932C" box="[1039,1180,1601,1624]" italics="true" pageId="15" pageNumber="88">E. morrisoni</emphasis>
</taxonomicName>
were found there or in nearby streams.
</paragraph>
</subSubSection>
</treatment>
</document>