155 lines
21 KiB
XML
155 lines
21 KiB
XML
<document id="A25D98176186826566CF565A5C3785A0" ID-CLB-Dataset="4969" ID-DOI="10.11646/zootaxa.4984.1.24" ID-GBIF-Dataset="bfd25ea4-72d4-4089-b954-6ba5755638e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="4927456" ID-ZooBank="0A3B8E9A-BA67-433A-A9C3-970DBD21F175" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623431039949" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2021" docId="03FECE14FF877F3CFF49ACF3FB50F91F" docLanguage="en" docName="zootaxa.4984.1.24.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deelemanikara Jäger 2021, gen. nov." docType="treatment" docVersion="6" lastPageNumber="339" masterDocId="FFC7B66CFF837F38FFDEAF05FFE0FFFE" masterDocTitle="Two new enigmatic genera of huntsman spiders from Madagascar (Araneae Sparassidae)" masterLastPageNumber="346" masterPageNumber="335" pageNumber="339" updateTime="1699010494404" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD5D96552E38A8204F7CC9C613503AB6">Two new enigmatic genera of huntsman spiders from Madagascar (Araneae Sparassidae)</mods:title>
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<mods:name id="B8186DB21BB4B58A15C8EC46E7282F12" type="personal">
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<mods:namePart id="E0DC7C87099B2C326DE86F7B2F57DC43">Jäger, Peter</mods:namePart>
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<mods:title id="7A0CF5590F3A9F6D9B319AD017DEBC24">Zootaxa</mods:title>
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<mods:part id="1004EA5980A60DBAC9A44B412E721490">
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<mods:date id="628C48D055FE9D201597A1E0D095395D">2021</mods:date>
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<mods:number id="C358AF88B6985B5E48878903C773FD75">2021-06-10</mods:number>
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<mods:number id="1CF9934A25275EB6D402DBBEECB0CED5">4984</mods:number>
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<mods:identifier id="AD509FD8605C7F817272934AE0AB1B12" type="DOI">10.11646/zootaxa.4984.1.24</mods:identifier>
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<treatment id="03FECE14FF877F3CFF49ACF3FB50F91F" ID-DOI="http://doi.org/10.5281/zenodo.4927464" ID-GBIF-Taxon="184162280" ID-Zenodo-Dep="4927464" LSID="urn:lsid:plazi:treatment:03FECE14FF877F3CFF49ACF3FB50F91F" httpUri="http://treatment.plazi.org/id/03FECE14FF877F3CFF49ACF3FB50F91F" lastPageNumber="339" pageId="4" pageNumber="339">
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<subSubSection id="C34D2C89FF877F3CFF49ACF3FE26FBEF" box="[151,454,1014,1041]" pageId="4" pageNumber="339" type="nomenclature">
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<paragraph id="8BE87F02FF877F3CFF49ACF3FE26FBEF" blockId="4.[151,454,1014,1041]" box="[151,454,1014,1041]" pageId="4" pageNumber="339">
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<heading id="D0A0C86EFF877F3CFF49ACF3FE26FBEF" bold="true" box="[151,454,1014,1041]" fontSize="11" level="1" pageId="4" pageNumber="339" reason="1">
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<emphasis id="B923A310FF877F3CFF49ACF3FE26FBEF" bold="true" box="[151,454,1014,1041]" pageId="4" pageNumber="339">
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<taxonomicName id="4C570481FF877F3CFF49ACF3FEB1FBEF" ID-CoL="95F68" authority="Jäger, 2021" authorityName="Jäger" authorityYear="2021" box="[151,337,1014,1041]" class="Arachnida" family="Sparassidae" genus="Deelemanikara" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B923A310FF877F3CFF49ACF3FEB1FBEF" bold="true" box="[151,337,1014,1041]" italics="true" pageId="4" pageNumber="339">Deelemanikara</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2101E6BFF877F3CFE86ACF2FE26FBEF" box="[344,454,1015,1041]" pageId="4" pageNumber="339" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2C89FF877F3CFF49AB3AFD5EFBA7" box="[151,702,1086,1113]" pageId="4" pageNumber="339" type="type_taxon">
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<paragraph id="8BE87F02FF877F3CFF49AB3AFD5EFBA7" blockId="4.[151,1437,1086,1761]" box="[151,702,1086,1113]" pageId="4" pageNumber="339">
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<emphasis id="B923A310FF877F3CFF49AB3AFED2FBA6" bold="true" box="[151,306,1086,1113]" pageId="4" pageNumber="339">
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<typeStatus id="54ECC1A0FF877F3CFF49AB3AFF32FBA7" box="[151,210,1087,1113]" pageId="4" pageNumber="339">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C570481FF877F3CFEE7AB3AFDA7FBA7" ID-CoL="95FKQ" authority="Jäger, 2021" authorityName="Jäger" authorityYear="2021" box="[313,583,1087,1113]" class="Arachnida" family="Sparassidae" genus="Deelemanikara" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="christae" status="sp. nov.">
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<emphasis id="B923A310FF877F3CFEE7AB3AFDA7FBA7" box="[313,583,1087,1113]" italics="true" pageId="4" pageNumber="339">Deelemanikara christae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2101E6BFF877F3CFD90AB3AFD5EFBA7" box="[590,702,1087,1113]" pageId="4" pageNumber="339" rank="species">spec. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2C89FF877F3CFF19AB66FDB9FB3B" pageId="4" pageNumber="339" type="etymology">
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<paragraph id="8BE87F02FF877F3CFF19AB66FDB9FB3B" blockId="4.[151,1437,1086,1761]" pageId="4" pageNumber="339">
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<emphasis id="B923A310FF877F3CFF19AB66FEAEFB83" bold="true" box="[199,334,1123,1149]" pageId="4" pageNumber="339">Etymology.</emphasis>
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Named in honour of Christa Deeleman-Reinhold on the occasion of her 90
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<superScript id="7C22D24AFF877F3CFB49AB67FB44FB8E" attach="left" box="[1175,1188,1122,1136]" fontSize="6" pageId="4" pageNumber="339">th</superScript>
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birthday in 2020. The generic name is a combination of the family name “Deeleman” and the Malagasy word for
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<collectingCountry id="F3403F92FF877F3CFBACAB82FB19FB5F" box="[1138,1273,1159,1185]" name="Madagascar" pageId="4" pageNumber="339">Madagascar</collectingCountry>
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“
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<collectingCountry id="F3403F92FF877F3CFAD9AB82FEDBFB3B" name="Madagascar" pageId="4" pageNumber="339">Repoblikan’i Madagasikara</collectingCountry>
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”. The gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2C89FF877F3CFF19ABCBFBCCFAAB" pageId="4" pageNumber="339" type="diagnosis">
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<paragraph id="8BE87F02FF877F3CFF19ABCBFBCCFAAB" blockId="4.[151,1437,1086,1761]" pageId="4" pageNumber="339">
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<emphasis id="B923A310FF877F3CFF19ABCBFEA2FB16" bold="true" box="[199,322,1230,1256]" pageId="4" pageNumber="339">Diagnosis.</emphasis>
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<taxonomicName id="4C570481FF877F3CFE96ABCAFE15FB17" authority="Jäger, 2021" authorityName="Jäger" authorityYear="2021" box="[328,501,1231,1257]" class="Arachnida" family="Sparassidae" genus="Deelemanikara" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B923A310FF877F3CFE96ABCAFE15FB17" box="[328,501,1231,1257]" italics="true" pageId="4" pageNumber="339">Deelemanikara</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2101E6BFF877F3CFE25ABCAFD81FB17" box="[507,609,1231,1257]" pageId="4" pageNumber="339" rank="genus">gen. nov.</taxonomicNameLabel>
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can be recognised by the following combination of characters: 1. Epigyne with anterior pocket (
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<figureCitation id="136C6387FF877F3CFE59ABF6FE3DFAF3" box="[391,477,1267,1293]" captionStart="FIGURES 14–22" captionStartId="5.[151,264,1874,1898]" captionTargetBox="[205,1372,181,1851]" captionTargetId="figure-16@5.[205,1382,181,1851]" captionTargetPageId="5" captionText="FIGURES 14–22. Deelemanikara christae gen. & spec. nov., holotype female (Madagascar, Beparasy, MRCA 142.748). 14–17 Copulatory organ (14 epigyne, ventral; 15 epigyne, lateral; 16 vulva, dorsal; 17 schematic course of internal duct system, dorsal). 18 Eyes, dorsal. 19 Left chelicera, ventral. 20 Trilobate membrane of metatarsus, right leg II, dorsal. 21 Right palpal claw, prolateral. 9 Prolateral claw of left leg II, prolateral. AP—anterior pocket, CO—copulatory opening (with mating plugs), EF—epigynal field, FD—fertilization duct." figureDoi="http://doi.org/10.5281/zenodo.4927468" httpUri="https://zenodo.org/record/4927468/files/figure.png" pageId="4" pageNumber="339">Figs 14</figureCitation>
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,
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<figureCitation id="136C6387FF877F3CFE37ABF6FDBAFAF3" box="[489,602,1267,1293]" captionStart="FIGURES 23–29" captionStartId="6.[151,264,1598,1622]" captionTargetBox="[151,1436,189,1574]" captionTargetId="figure-22@6.[151,1436,181,1574]" captionTargetPageId="6" captionText="FIGURES 23–29. Deelemanikara christae gen. & spec. nov., Madagascar (23–26 2 females,Antsirabe, MRCA 142.587; 27–29 holotype female, Beparasy, MRCA 142.748). 23–24 epigyne, ventral. 25–26 Distal tip of metatarsi showing insertion points of bristles, ventral (25 leg III, 26 leg IV). 27 Anterior pocket, frontal. 28 Left part of vulva, ventral. 29 Schematic course of left internal duct system, ventral.AP—anterior pocket, EP—epigynal plug, TB—transversal bulge of median septum." figureDoi="http://doi.org/10.5281/zenodo.4927472" httpUri="https://zenodo.org/record/4927472/files/figure.png" pageId="4" pageNumber="339">23–24, 27</figureCitation>
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) situated on a distinct transversal ledge (best seen in lateral view:
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<figureCitation id="136C6387FF877F3CFAE3ABF6FA6FFAF3" box="[1341,1423,1266,1293]" captionStart="FIGURES 14–22" captionStartId="5.[151,264,1874,1898]" captionTargetBox="[205,1372,181,1851]" captionTargetId="figure-16@5.[205,1382,181,1851]" captionTargetPageId="5" captionText="FIGURES 14–22. Deelemanikara christae gen. & spec. nov., holotype female (Madagascar, Beparasy, MRCA 142.748). 14–17 Copulatory organ (14 epigyne, ventral; 15 epigyne, lateral; 16 vulva, dorsal; 17 schematic course of internal duct system, dorsal). 18 Eyes, dorsal. 19 Left chelicera, ventral. 20 Trilobate membrane of metatarsus, right leg II, dorsal. 21 Right palpal claw, prolateral. 9 Prolateral claw of left leg II, prolateral. AP—anterior pocket, CO—copulatory opening (with mating plugs), EF—epigynal field, FD—fertilization duct." figureDoi="http://doi.org/10.5281/zenodo.4927468" httpUri="https://zenodo.org/record/4927468/files/figure.png" pageId="4" pageNumber="339">Fig. 15</figureCitation>
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). 2. Both eye rows slightly procurved (
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<figureCitation id="136C6387FF877F3CFDEAAA12FD67FACF" box="[564,647,1303,1329]" captionStart="FIGURES 14–22" captionStartId="5.[151,264,1874,1898]" captionTargetBox="[205,1372,181,1851]" captionTargetId="figure-16@5.[205,1382,181,1851]" captionTargetPageId="5" captionText="FIGURES 14–22. Deelemanikara christae gen. & spec. nov., holotype female (Madagascar, Beparasy, MRCA 142.748). 14–17 Copulatory organ (14 epigyne, ventral; 15 epigyne, lateral; 16 vulva, dorsal; 17 schematic course of internal duct system, dorsal). 18 Eyes, dorsal. 19 Left chelicera, ventral. 20 Trilobate membrane of metatarsus, right leg II, dorsal. 21 Right palpal claw, prolateral. 9 Prolateral claw of left leg II, prolateral. AP—anterior pocket, CO—copulatory opening (with mating plugs), EF—epigynal field, FD—fertilization duct." figureDoi="http://doi.org/10.5281/zenodo.4927468" httpUri="https://zenodo.org/record/4927468/files/figure.png" pageId="4" pageNumber="339">Fig. 18</figureCitation>
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). 3. Cheliceral furrow with 3 promarginal teeth and most intermarginal denticles arranged in a single line (
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<figureCitation id="136C6387FF877F3CFDC0AA3EFD91FAAB" box="[542,625,1339,1365]" captionStart="FIGURES 14–22" captionStartId="5.[151,264,1874,1898]" captionTargetBox="[205,1372,181,1851]" captionTargetId="figure-16@5.[205,1382,181,1851]" captionTargetPageId="5" captionText="FIGURES 14–22. Deelemanikara christae gen. & spec. nov., holotype female (Madagascar, Beparasy, MRCA 142.748). 14–17 Copulatory organ (14 epigyne, ventral; 15 epigyne, lateral; 16 vulva, dorsal; 17 schematic course of internal duct system, dorsal). 18 Eyes, dorsal. 19 Left chelicera, ventral. 20 Trilobate membrane of metatarsus, right leg II, dorsal. 21 Right palpal claw, prolateral. 9 Prolateral claw of left leg II, prolateral. AP—anterior pocket, CO—copulatory opening (with mating plugs), EF—epigynal field, FD—fertilization duct." figureDoi="http://doi.org/10.5281/zenodo.4927468" httpUri="https://zenodo.org/record/4927468/files/figure.png" pageId="4" pageNumber="339">Fig. 19</figureCitation>
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). 4. Leg claws with convex dorsal side.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2C89FF877F3CFF19AA5BFD72FA63" pageId="4" pageNumber="339" type="description">
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<paragraph id="8BE87F02FF877F3CFF19AA5BFCD3FA87" blockId="4.[151,1437,1086,1761]" box="[199,819,1374,1401]" pageId="4" pageNumber="339">
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<emphasis id="B923A310FF877F3CFF19AA5BFEB9FA86" bold="true" box="[199,345,1374,1400]" pageId="4" pageNumber="339">Description.</emphasis>
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See description of the
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<typeStatus id="54ECC1A0FF877F3CFD84AA5AFD6AFA87" box="[602,650,1375,1401]" pageId="4" pageNumber="339">type</typeStatus>
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species below.
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</paragraph>
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<paragraph id="8BE87F02FF877F3CFF19AA87FD72FA63" blockId="4.[151,1437,1086,1761]" box="[199,658,1410,1437]" pageId="4" pageNumber="339">
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<emphasis id="B923A310FF877F3CFF19AA87FE73FA62" bold="true" box="[199,403,1410,1436]" pageId="4" pageNumber="339">Species included.</emphasis>
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Only the
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<typeStatus id="54ECC1A0FF877F3CFDDCAA86FDD2FA63" box="[514,562,1411,1437]" pageId="4" pageNumber="339">type</typeStatus>
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species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2C89FF877F3CFF19AAA3FB50F91F" pageId="4" pageNumber="339" type="distribution">
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<paragraph id="8BE87F02FF877F3CFF19AAA3FEA7FA18" blockId="4.[151,1437,1086,1761]" pageId="4" pageNumber="339">
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<emphasis id="B923A310FF877F3CFF19AAA3FE81FA3E" bold="true" box="[199,353,1446,1472]" pageId="4" pageNumber="339">Distribution.</emphasis>
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Known from two localities in central
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<collectingCountry id="F3403F92FF877F3CFCD8AAA2FC6DFA3F" box="[774,909,1447,1473]" name="Madagascar" pageId="4" pageNumber="339">Madagascar</collectingCountry>
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(
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<collectingRegion id="4993B1E0FF877F3CFC45AAA2FBF6FA3F" box="[923,1046,1447,1473]" country="Madagascar" name="Toamasina" pageId="4" pageNumber="339">Toamasina</collectingRegion>
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and
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<collectingRegion id="4993B1E0FF877F3CFB94AAA2FB02FA3F" box="[1098,1250,1447,1473]" country="Madagascar" name="Antananarivo" pageId="4" pageNumber="339">Antananarivo</collectingRegion>
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Provinces) (
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<figureCitation id="136C6387FF877F3CFAB1AAA2FF57FA1B" captionStart="FIGURES 30–36" captionStartId="7.[151,264,1699,1723]" captionTargetBox="[151,1435,363,1667]" captionTargetId="figure-90@7.[151,1436,344,1675]" captionTargetPageId="7" captionText="FIGURES 30–36. 30–35 Deelemanikara christae gen. & spec. nov., females from Madagascar (30–33 holotype from Beparasy, MRCA 142.748; 34–35 female from Antsirabe, MRCA 142.587), habitus (30 dorsal, 31, 35 ventral, 32, 34 frontal, 33 lateral). 36 Localities of Martensikara jocheni gen. & spec. nov. (green triangle) and of Deelemanikara christae gen. & spec. nov. (red circles) on Madagascar." figureDoi="http://doi.org/10.5281/zenodo.4927479" httpUri="https://zenodo.org/record/4927479/files/figure.png" pageId="4" pageNumber="339">Fig. 36</figureCitation>
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: red circles).
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</paragraph>
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<paragraph id="8BE87F02FF877F3CFF19AAEAFB50F91F" blockId="4.[151,1437,1086,1761]" pageId="4" pageNumber="339">
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<emphasis id="B923A310FF877F3CFF19AAEAFEF1F9F7" bold="true" box="[199,273,1519,1545]" pageId="4" pageNumber="339">Notes.</emphasis>
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<taxonomicName id="4C570481FF877F3CFEC6AAEAFE25F9F7" authority="Jäger, 2021" authorityName="Jäger" authorityYear="2021" box="[280,453,1519,1545]" class="Arachnida" family="Sparassidae" genus="Deelemanikara" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B923A310FF877F3CFEC6AAEAFE25F9F7" box="[280,453,1519,1545]" italics="true" pageId="4" pageNumber="339">Deelemanikara</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2101E6BFF877F3CFE12AAEAFDD2F9F7" box="[460,562,1519,1545]" pageId="4" pageNumber="339" rank="genus">gen. nov.</taxonomicNameLabel>
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currently cannot be assigned to any subfamily. It combines characters of
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<taxonomicName id="4C570481FF877F3CFAB4AAEAFEF3F9D3" authorityName="Thorell" authorityYear="1873" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Heteropodinae">Heteropodinae</taxonomicName>
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(cheliceral dentition, trilobate membrane, one cheliceral escort seta), Sparassinae (leg tibiae lacking distal pair of ventral spines),
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<taxonomicName id="4C570481FF877F3CFE00A932FD02F9AF" authority="Simon, 1880" authorityName="Simon" authorityYear="1880" box="[478,738,1591,1617]" class="Arachnida" family="Sparassidae" genus="Damastes" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923A310FF877F3CFE00A932FDACF9AE" box="[478,588,1591,1616]" italics="true" pageId="4" pageNumber="339">Damastes</emphasis>
|
||
<bibRefCitation id="EFC602F3FF877F3CFD8DA932FD02F9AF" author="Simon, E." box="[595,738,1591,1617]" pageId="4" pageNumber="339" pagination="223 - 351" refId="ref7374" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe Linneenne de Bordeaux, 34, 223 - 351." type="journal article" year="1880">Simon, 1880</bibRefCitation>
|
||
</taxonomicName>
|
||
(eye arrangement, ventro-distal metatarsal bristles) and
|
||
<taxonomicName id="4C570481FF877F3CFA87A932FF37F98B" authorityName="Jager" authorityYear="2020" class="Arachnida" family="Sparassidae" genus="Thunberga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923A310FF877F3CFA87A932FF37F98B" italics="true" pageId="4" pageNumber="339">Thunberga</emphasis>
|
||
</taxonomicName>
|
||
(anterior epigynal pocket, ventro-distal metatarsal bristles). Moreover, the very variable leg spination (especially on femora) and the convex dorsal margin of leg claws (the latter known in a more extreme form from
|
||
<taxonomicName id="4C570481FF877F3CFA9AA97AFEA1F943" authority="L. Koch, 1875" authorityName="L. Koch" authorityYear="1875" class="Arachnida" family="Sparassidae" genus="Prychia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923A310FF877F3CFA9AA97AFA7DF967" box="[1348,1437,1663,1689]" italics="true" pageId="4" pageNumber="339">Prychia</emphasis>
|
||
L.
|
||
<bibRefCitation id="EFC602F3FF877F3CFF67A9A6FEA1F943" author="Koch, L." box="[185,321,1698,1725]" pageId="4" pageNumber="339" pagination="577 - 740" refId="ref6928" refString="Koch, L. (1875) s. n. In: Die Arachniden Australiens 1. Bauer & Raspe, Nurnberg, pp. 577 - 740." type="book chapter" year="1875">Koch, 1875</bibRefCitation>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="EFC602F3FF877F3CFEA3A9A6FE1AF943" author="Jager, P." box="[381,506,1699,1725]" pageId="4" pageNumber="339" pagination="53 - 59" refId="ref5814" refString="Jager, P. (1998) First results of a taxonomic revision of the SE Asian Sparassidae (Araneae). In: Selden, P. A. (Ed.), Proceedings of the 17 th European Colloquium of Arachnology, Edinburgh, 1997. British Arachnological Society, Burnham Beeches, Bucks, pp. 53 - 59." type="book chapter" year="1998">Jäger 1998</bibRefCitation>
|
||
) are unique characters that do not fit any of the known subfamilies. Therefore a diagnosis differentiating the new genus from its most closely related taxa cannot be provided.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |