treatments-xml/data/E1/48/6C/E1486C4BB40B3D5696D056A00BE96A74.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.28.6378" ID-PMC="PMC3881412" ID-Pensoft-Pub="1314-2003-28-9" ID-Pensoft-UUID="177E3C2F8E03131A525AFFDBFFB1FFC0" ID-PubMed="24399890" ID-Zenodo-Dep="576180" ModsDocID="1314-2003-28-9" checkinTime="1451251822051" checkinUser="pensoft" docAuthor="Diazgranados, Mauricio &amp; Morillo, Gilberto" docDate="2013" docId="E1486C4BB40B3D5696D056A00BE96A74" docLanguage="en" docName="PhytoKeys 28: 9-18" docOrigin="PhytoKeys 28" docPubDate="2013-11-04" docSource="http://dx.doi.org/10.3897/phytokeys.28.6378" docTitle="Coespeletia palustris M. Diazgranados &amp; G. Morillo 2013, sp. nov." docType="treatment" docVersion="6" id="177E3C2F8E03131A525AFFDBFFB1FFC0" lastPageNumber="15" masterDocId="177E3C2F8E03131A525AFFDBFFB1FFC0" masterDocTitle="A new species of Coespeletia (Asteraceae, Millerieae) from Venezuela" masterLastPageNumber="18" masterPageNumber="9" pageNumber="11" updateTime="1668140934668" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new species of Coespeletia (Asteraceae, Millerieae) from Venezuela</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Diazgranados, Mauricio</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0448-5706</mods:nameIdentifier>
<mods:affiliation>Dept. of Botany, MRC 166, National Museum of Natural History, P. O. Box 37012, Smithsonian Institution, Washington D. C. 20013 - 7012, United States</mods:affiliation>
<mods:nameIdentifier type="email">m.diazgranados@kew.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Morillo, Gilberto</mods:namePart>
<mods:affiliation>Departamento de Botanica, Escuela de Ingenieria Forestal, Facultad de Ciencias Forestales y Ambientales, Universidad de Los Andes, Merida 5101 A, Venezuela</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-11-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>28</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>9</mods:start>
<mods:end>18</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.28.6378</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.28.6378</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-28-9</mods:identifier>
<mods:identifier type="Pensoft-UUID">177E3C2F8E03131A525AFFDBFFB1FFC0</mods:identifier>
<mods:identifier type="Zenodo-Dep">576180</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152024802" LSID="urn:lsid:plazi:treatment:E1486C4BB40B3D5696D056A00BE96A74" httpUri="http://treatment.plazi.org/id/E1486C4BB40B3D5696D056A00BE96A74" lastPageId="6" lastPageNumber="15" pageId="2" pageNumber="11">
<subSubSection pageId="2" pageNumber="11" type="nomenclature">
<paragraph pageId="2" pageNumber="11">
<taxonomicName LSID="E1486C4B-B40B-3D56-96D0-56A00BE96A74" authority="M. Diazgranados &amp; G. Morillo" authorityName="M. Diazgranados &amp; G. Morillo" authorityYear="2013" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Coespeletia palustris" order="Asterales" pageId="2" pageNumber="11" phylum="Magnoliophyta" rank="species" species="palustris" status="sp. nov.">Coespeletia palustris M. Diazgranados &amp; G. Morillo</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="11">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Coespeletia palustris: A abaxial view of the involucre B young leaves showing white basal portion C parts of the collection D capitulum of the holotype collection (Morillo et al. 14155) E acaulescent (or subsessile) habit and marshy habitat F whitish rosette from top. Coespeletia moritziana: G capitulum H caulescent habit and rocky habitat I yellowish rosette from top." httpUri="https://binary.pensoft.net/fig/10467" pageId="2" pageNumber="11">Figs 1</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map showing collections for Coespeletia palustris (red circles) and Coespeletia moritziana (green triangles). Photograph of marshes around Laguna de los Patos, with a population of Coespeletia palustris." httpUri="https://binary.pensoft.net/fig/10471" pageId="2" pageNumber="11">-4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="11" type="diagnosis">
<paragraph pageId="2" pageNumber="11">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="11">
Related to 
<taxonomicName authorityName="J. Cuatrec" authorityYear="1976" baseAuthorityName="Sch. Bip. ex Wedd." class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="2" pageNumber="11" phylum="Magnoliophyta" rank="species" species="moritziana">
<emphasis italics="true" pageId="2" pageNumber="11">Coespeletia moritziana</emphasis>
</taxonomicName>
but differs in indumentum primarily whitish, larger leaf sheaths, proximal portion of young leaves white, thicker scapes, which are ebracteate, larger phyllaries with pubescence unkempt, capitula with fewer flowers, which are much larger, and smaller pollen grains.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="11" type="type">
<paragraph pageId="2" pageNumber="11">Type.</paragraph>
<paragraph pageId="2" pageNumber="11">
Venezuela, 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
, 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Santo Domingo, alrededores de la Laguna de los Patos, en zona de turbera. Alt. 3729 m., 
<geoCoordinate degrees="8.77522" direction="north" orientation="latitude" precision="1" value="8.77522">8.77522 N</geoCoordinate>
, - 
<geoCoordinate degrees="70.80349" direction="east" orientation="longitude" precision="1" value="70.80349">70.80349 E</geoCoordinate>
, 29 September 2011, 
<emphasis italics="true" pageId="2" pageNumber="11">G. Morillo</emphasis>
, 
<emphasis italics="true" pageId="2" pageNumber="11">M. Diazgranados</emphasis>
, 
<emphasis italics="true" pageId="2" pageNumber="11">
L. 
<normalizedToken originalValue="Gámez">Gamez</normalizedToken>
</emphasis>
, 
<emphasis italics="true" pageId="2" pageNumber="11">
S. 
<normalizedToken originalValue="Rodríguez">Rodriguez</normalizedToken>
&amp; J. Parra 14155
</emphasis>
(holotype: MER, isotype: VEN). Additional collections from the type locality (paratypes): 
<emphasis italics="true" pageId="2" pageNumber="11">L. Gamez</emphasis>
, 
<emphasis italics="true" pageId="2" pageNumber="11">G. Morillo</emphasis>
, 
<emphasis italics="true" pageId="2" pageNumber="11">
J. Parra &amp; S. 
<normalizedToken originalValue="Rodríguez">Rodriguez</normalizedToken>
1097
</emphasis>
, 
<emphasis italics="true" pageId="2" pageNumber="11">1100</emphasis>
(MER).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="12" pageId="2" pageNumber="11" type="description">
<paragraph pageId="2" pageNumber="11">Description.</paragraph>
<paragraph pageId="2" pageNumber="11">Acaulescent (or subsessile) rosette of 60-80(-100) cm (including inflorescences) and of 40-60 cm in diameter; numerous inflorescences monocephalous, twice longer than leaves, with naked axes. Rosettes whitish, densely covered by whitish lanose indumentum, with little accumulation of marcescent leaves at the base. Leaves linear, with laminae (28-)32-36(-38) cm long, (0.7-)0.8-1.2(-1.3) cm wide, length to width ratio (28-)30(-35):1, thick, subsessile, with a pseudopetiole greenish and almost glabrous, 2 cm long and 0.5 cm wide. The leaves are covered by a dense lanose indumentum, grey-whitish. When young, leaves are golden yellow in the distal portion, and white in the proximal portion. The sheath is oblong, adaxially glabrescent and greenish or whitish, abaxially reddish-brown and slightly covered by hairs, (7-)7.5-9.5(-10) cm long, (2-)2.2-2.5(-2.7) cm wide.</paragraph>
<paragraph pageId="2" pageNumber="11">Inflorescences numerous (&gt;30), coetaneous, axillary, aphyllous, monocephalous, with a single nodding capitulum. Scape about twice longer than leaves, 60-80 cm long, 1.0-1.4 cm in diameter, erect, densely lanate, yellowish, relatively thick and stout, totally naked, without leafy bracts at the base.</paragraph>
<paragraph pageId="2" pageNumber="11">
Capitula radiate, patelliform, nodding, 4.5-6 cm in diameter including the ligules of the ray flowers; ligular circle well developed, as large as the involucre; discs (without counting ligular circle) 2.2-2.4 cm in diameter. Involucre sub-hemispherical, densely lanate, the pubescence unkempt, yellowish or whitish. Phyllaries pluriseriate, narrow and curved (almost curly), the outer ones 18-20 mm 
<normalizedToken originalValue="×">x</normalizedToken>
4.0-4.2 cm, linear to narrowly-lanceolate, becoming progressively shorter towards the inner-most phyllaries, 13-14 mm 
<normalizedToken originalValue="×">x</normalizedToken>
2.8-3.2 mm, only apically hairy.
</paragraph>
<paragraph lastPageId="3" lastPageNumber="12" pageId="2" pageNumber="11">
Ray flowers ligulate, 95-200 per capitulum, in 3-5 rows. Corolla abaxially dark pink, adaxially yellowish becoming brown-reddish towards the apex, carnosulous, 12-14.2 mm long; ligules elliptical or oblong, bi- or tridentate, with 6 reddish-purple conspicuous nerves. Tube glanduliferous, 0.6-1.0 mm in diameter and 3.8-4.8(-6.0) mm in length, dark pink, the glands pediculate capitate, 150-160 
<normalizedToken originalValue="μm">μm</normalizedToken>
long 
<normalizedToken originalValue="×">x</normalizedToken>
30-35 
<normalizedToken originalValue="μm">μm</normalizedToken>
<pageBreakToken pageId="3" pageNumber="12" start="start">in</pageBreakToken>
diameter; tubes with 2 linguiform appendages 1.5-2.2 mm long; ovary whitish when young, becoming red in the extremes. Paleas 12-16(-17.5) mm long, 2.7-3.0 mm wide, brownish, with 3 main nerves. Styles 8-9 mm long, 0.2-0.3 mm wide, with branches about (1.7-)2.0-3.2 mm long, papillose. Cypselae oblong, triangular, 3.6-3.9 mm 
<normalizedToken originalValue="×">x</normalizedToken>
(1.6-)1.7-2.3 mm.
</paragraph>
<paragraph pageId="3" pageNumber="12">
Disc flowers 215-280, (8-)9-10 mm long (not counting the anthers), of purple appearance; corolla 5-5.5 mm long, reddish, yellowish pink or translucid in the lower half, lobes 5, 1-1.1 mm long and 0.5-0.6 mm wide at the base; tube glanduliferous, 0.8 mm wide 
<normalizedToken originalValue="×">x</normalizedToken>
2.6-4.5 mm long; anthers purple, sometimes exceeding the corolla by 2 mm, slightly translucid; paleas 9.8-10.2 mm long, 1.5-1.7 mm wide, brownish, with 3 main nerves.
</paragraph>
<paragraph pageId="3" pageNumber="12">
Pollen yellow when fresh, tricolporate, 22.1-23.7 
<normalizedToken originalValue="μm">μm</normalizedToken>
in equatorial diameter (not including spines), 25-26 
<normalizedToken originalValue="μm">μm</normalizedToken>
in polar diameter; spines 110-116 total, equatorial spines 18, 2.3-3.2 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, slightly curved.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10467" pageId="3" pageNumber="12" start="Figure 1" startId="F1">
<paragraph pageId="3" pageNumber="12">
<emphasis bold="true" pageId="3" pageNumber="12">Figure 1.</emphasis>
<taxonomicName authorityName="M. Diazgranados &amp; G. Morillo" authorityYear="2013" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="12" phylum="Magnoliophyta" rank="species" species="palustris">
<emphasis italics="true" pageId="3" pageNumber="12">Coespeletia palustris</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="3" pageNumber="12">A</emphasis>
abaxial view of the involucre 
<emphasis bold="true" pageId="3" pageNumber="12">B</emphasis>
young leaves showing white basal portion 
<emphasis bold="true" pageId="3" pageNumber="12">C</emphasis>
parts of the collection 
<emphasis bold="true" pageId="3" pageNumber="12">D</emphasis>
capitulum of the holotype collection (Morillo et al. 14155) 
<emphasis bold="true" pageId="3" pageNumber="12">E</emphasis>
acaulescent (or subsessile) habit and marshy habitat 
<emphasis bold="true" pageId="3" pageNumber="12">F</emphasis>
whitish rosette from top. 
<taxonomicName authorityName="J. Cuatrec" authorityYear="1976" baseAuthorityName="Sch. Bip. ex Wedd." class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="12" phylum="Magnoliophyta" rank="species" species="moritziana">
<emphasis italics="true" pageId="3" pageNumber="12">Coespeletia moritziana</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="3" pageNumber="12">G</emphasis>
capitulum 
<emphasis bold="true" pageId="3" pageNumber="12">H</emphasis>
caulescent habit and rocky habitat 
<emphasis bold="true" pageId="3" pageNumber="12">I</emphasis>
yellowish rosette from top.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/10469" pageId="3" pageNumber="12" start="Figure 2" startId="F2">
<paragraph pageId="3" pageNumber="12">
<emphasis bold="true" pageId="3" pageNumber="12">Figure 2.</emphasis>
Illustrations of 
<taxonomicName authorityName="M. Diazgranados &amp; G. Morillo" authorityYear="2013" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="12" phylum="Magnoliophyta" rank="species" species="palustris">
<emphasis italics="true" pageId="3" pageNumber="12">Coespeletia palustris</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="3" pageNumber="12">A</emphasis>
Ray flower 
<emphasis bold="true" pageId="3" pageNumber="12">B</emphasis>
cypsela 
<emphasis bold="true" pageId="3" pageNumber="12">C</emphasis>
disc flower style 
<emphasis bold="true" pageId="3" pageNumber="12">D</emphasis>
disk flower 
<emphasis bold="true" pageId="3" pageNumber="12">E</emphasis>
disk flower palea 
<emphasis bold="true" pageId="3" pageNumber="12">F</emphasis>
outer phyllary 
<emphasis bold="true" pageId="3" pageNumber="12">G</emphasis>
inner (sterile) phyllary 
<emphasis bold="true" pageId="3" pageNumber="12">H</emphasis>
fertile ray flower palea.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/10470" pageId="3" pageNumber="12" start="Figure 3" startId="F3">
<paragraph pageId="3" pageNumber="12">
<emphasis bold="true" pageId="3" pageNumber="12">Figure 3.</emphasis>
Photomicrographs of 
<taxonomicName authorityName="M. Diazgranados &amp; G. Morillo" authorityYear="2013" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="12" phylum="Magnoliophyta" rank="species" species="palustris">
<emphasis italics="true" pageId="3" pageNumber="12">Coespeletia palustris</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="3" pageNumber="12">A</emphasis>
Tip of stigmatic branches in a disc flower 
<emphasis bold="true" pageId="3" pageNumber="12">B</emphasis>
pollen grain 
<emphasis bold="true" pageId="3" pageNumber="12">C</emphasis>
cypsela 
<emphasis bold="true" pageId="3" pageNumber="12">D</emphasis>
granduliferous tube of ray flower.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="12" type="distribution">
<paragraph pageId="3" pageNumber="12">Distribution</paragraph>
<paragraph pageId="3" pageNumber="12">
(
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map showing collections for Coespeletia palustris (red circles) and Coespeletia moritziana (green triangles). Photograph of marshes around Laguna de los Patos, with a population of Coespeletia palustris." httpUri="https://binary.pensoft.net/fig/10471" pageId="3" pageNumber="12">Fig. 4</figureCitation>
). Endemic to Venezuela. This species has been found in a few marshy areas of the 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Santo Domingo (e.g. around the Laguna de los Patos) and the Sierra de la Culata, always in small areas of less than 0.5 km2.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10471" pageId="3" pageNumber="12" start="Figure 4" startId="F4">
<paragraph pageId="3" pageNumber="12">
<emphasis bold="true" pageId="3" pageNumber="12">Figure 4.</emphasis>
Distribution map showing collections for 
<taxonomicName authorityName="M. Diazgranados &amp; G. Morillo" authorityYear="2013" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="12" phylum="Magnoliophyta" rank="species" species="palustris">
<emphasis italics="true" pageId="3" pageNumber="12">Coespeletia palustris</emphasis>
</taxonomicName>
(red circles) and 
<taxonomicName authorityName="J. Cuatrec" authorityYear="1976" baseAuthorityName="Sch. Bip. ex Wedd." class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="12" phylum="Magnoliophyta" rank="species" species="moritziana">
<emphasis italics="true" pageId="3" pageNumber="12">Coespeletia moritziana</emphasis>
</taxonomicName>
(green triangles). Photograph of marshes around Laguna de los Patos, with a population of 
<taxonomicName authorityName="M. Diazgranados &amp; G. Morillo" authorityYear="2013" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="12" phylum="Magnoliophyta" rank="species" species="palustris">
<emphasis italics="true" pageId="3" pageNumber="12">Coespeletia palustris</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="13" type="ecology">
<paragraph pageId="4" pageNumber="13">
<pageBreakToken pageId="4" pageNumber="13" start="start">Ecology</pageBreakToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="13">
Small population of about 200-400 individuals growing in marshes, on very swampy and wet soil. Other 
<taxonomicName lsidName="" pageId="4" pageNumber="13" rank="subFamily" subFamily="Espeletiinae">Espeletiinae</taxonomicName>
found on the proximate drier slopes are: 
<taxonomicName authorityName="Cuatrec" authorityYear="1976" baseAuthorityName="Cuatrec" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="4" pageNumber="13" phylum="Magnoliophyta" rank="species" species="timotensis">
<emphasis italics="true" pageId="4" pageNumber="13">Coespeletia timotensis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="4" pageNumber="13" phylum="Magnoliophyta" rank="species" species="schultzii">
<emphasis italics="true" pageId="4" pageNumber="13">Espeletia schultzii</emphasis>
</taxonomicName>
Wedd., 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="4" pageNumber="13" phylum="Magnoliophyta" rank="species" species="weddellii">
<emphasis italics="true" pageId="4" pageNumber="13">Espeletia weddellii</emphasis>
</taxonomicName>
Sch. Bip. ex Wedd., and 
<taxonomicName authorityName="Ernst" authorityYear="1870" baseAuthorityName="Bonpl. ex Kunth" class="Magnoliopsida" family="Asteraceae" genus="Libanothamnus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="4" pageNumber="13" phylum="Magnoliophyta" rank="species" species="neriifolius">
<emphasis italics="true" pageId="4" pageNumber="13">Libanothamnus neriifolius</emphasis>
</taxonomicName>
(Bonpl. ex Humb.) Ernst.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="14" type="etymology">
<paragraph pageId="5" pageNumber="14">
<pageBreakToken pageId="5" pageNumber="14" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="5" pageNumber="14">
The name 
<normalizedToken originalValue="“palustris”">&quot;palustris&quot;</normalizedToken>
is given because of the boggy habitat in which this species is found.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="14" type="conservation status">
<paragraph pageId="5" pageNumber="14">Conservation status.</paragraph>
<paragraph pageId="5" pageNumber="14">This species may be at a certain risk of extinction, since it is found in a very particular habitat, sensitive to climate change.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="15" pageId="5" pageNumber="14" type="specimens examined">
<paragraph pageId="5" pageNumber="14">Specimens examined.</paragraph>
<paragraph pageId="6" pageNumber="15">
<taxonomicName authorityName="J. Cuatrec" authorityYear="1976" baseAuthorityName="Sch. Bip. ex Wedd." class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="6" pageNumber="15" phylum="Magnoliophyta" rank="species" species="moritziana">
<emphasis italics="true" pageId="6" pageNumber="15">
<pageBreakToken pageId="6" pageNumber="15" start="start">Coespeletia</pageBreakToken>
moritziana
</emphasis>
</taxonomicName>
: VENEZUELA. 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
: Sierra Nevada de 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
: 4000 m, 1842, 
<emphasis italics="true" pageId="6" pageNumber="15">Linden 398</emphasis>
in part (isolectotype; BR, P, US!); Sierra Nevada, flower, Sep-Dec, 
<emphasis italics="true" pageId="6" pageNumber="15">Moritz 1416</emphasis>
(F!); 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de 
<normalizedToken originalValue="Mucuchíes">Mucuchies</normalizedToken>
around Pico del Aguila, 4300 m, in Apartaderos, 
<emphasis italics="true" pageId="6" pageNumber="15">Steyermark 55883</emphasis>
(F!, NY, VEN); id., 4150 m, Sep 1952, 
<emphasis italics="true" pageId="6" pageNumber="15">Humbert 26458</emphasis>
(P, US!); id., Pico del Aguila, 4100 m, Apr 1952, 
<emphasis italics="true" pageId="6" pageNumber="15">Gines 4794</emphasis>
(US!); id., 4200 m, 5 Oct 1969, 
<emphasis italics="true" pageId="6" pageNumber="15">Cuatrecasas</emphasis>
, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; Lopez-Figueiras 28031
</emphasis>
(MERF, US!); id., 4000 m, 5 Dec 1936, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Chardón">Chardon</normalizedToken>
5007
</emphasis>
(COL!, US!); id., 20 Apr 1946, 
<emphasis italics="true" pageId="6" pageNumber="15">Burkart 16802</emphasis>
(US!, VEN); id., Apartaderos towards Timotes, 3 Nov 1976, 
<emphasis italics="true" pageId="6" pageNumber="15">Bernardi</emphasis>
, 
<emphasis italics="true" pageId="6" pageNumber="15">Charpin &amp; Jacquemoud 17064</emphasis>
(G, US!); id., 4118 m, 21-26 Apr 1959, 
<emphasis italics="true" pageId="6" pageNumber="15">Barclay &amp; Juajibioy 9649</emphasis>
(US!); id., 4200 m, 15 Nov 1975, 
<emphasis italics="true" pageId="6" pageNumber="15">
Badillo &amp; 
<normalizedToken originalValue="Páez">Paez</normalizedToken>
6857
</emphasis>
(MY, US!); id., 4000 m, 26 Jan 1939, 
<emphasis italics="true" pageId="6" pageNumber="15">Alston 6631</emphasis>
(GH, NY, P, S, U, US!); Sierra Nevada above 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
, Loma Redonda, 4000-4045 m, 14 Oct 1969, 
<emphasis italics="true" pageId="6" pageNumber="15">Cuatrecasas</emphasis>
, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; 
<normalizedToken originalValue="López-Figueiras">Lopez-Figueiras</normalizedToken>
28093
</emphasis>
(MERF, US!); id., 4050 m, 9 Dec 1983, 
<emphasis italics="true" pageId="6" pageNumber="15">Sobrevila 1552</emphasis>
(US!); id., around Laguna Verde, below Pico Humboldt, 4100 m, 26 Feb 1971, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; Lopez-Palacios 1617
</emphasis>
(MERF, US!); id., Laguna Verde, 4000 m, on rocky slope, 4 Dec 1959, 
<emphasis italics="true" pageId="6" pageNumber="15">Barclay &amp; Juajibioy 10043</emphasis>
(US!); between Laguna Coromoto and Laguna Verde, 3800 m, Oct 1956, 
<emphasis italics="true" pageId="6" pageNumber="15">Aristeguieta 2610</emphasis>
(F, NY, US!, VEN); 
<normalizedToken originalValue="Páramos">Paramos</normalizedToken>
around Pico Bolivar and Pico Espejo on trail to Nevados, 4150 m, 15-18 Dec 1959, 
<emphasis italics="true" pageId="6" pageNumber="15">Barclay &amp; Juajibioy 10224</emphasis>
(US!); id., 
<normalizedToken originalValue="Cañada">Canada</normalizedToken>
of Laguna de Los Anteojos, 3930-3900 m, 22 Feb 1973, 
<emphasis italics="true" pageId="6" pageNumber="15">Cuatrecasas</emphasis>
, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; 
<normalizedToken originalValue="López-Figueiras">Lopez-Figueiras</normalizedToken>
28584
</emphasis>
(MERF, US!); Pico Toro, southern slope on the way to Los Nevados, frequently humid and marshy places, 4000-4200 m, 22 Jan 1972, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
6882
</emphasis>
(MERF, US!); Sierra Nevada de Santo Domingo; NW slopes, 4000-4400 m, 24-26 Sep 1952, 
<emphasis italics="true" pageId="6" pageNumber="15">Humbert 26479</emphasis>
(COL, P, US!); 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Laguna Brava, 3300 m, 22 May 1971, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="López-Figueiras">Lopez-Figueiras</normalizedToken>
8721
</emphasis>
(MERF, US!); 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Timotes, 3800-4200 m, 24 Jan 1928, 
<emphasis italics="true" pageId="6" pageNumber="15">Pittier 12729</emphasis>
(F!, M, MO!, NY, VEN); id., 4200 m, 21 Jan 1922, 
<emphasis italics="true" pageId="6" pageNumber="15">Jahn 869</emphasis>
(GH, K, M, NY!, US!, VEN); id., 3000-3500 m, 6 Dec 1910, 
<emphasis italics="true" pageId="6" pageNumber="15">Jahn 150</emphasis>
(US!, VEN); id., Cabeceras of Quebrada El Turmero, afluent of Rio Motatan, 4260 m, in marshy places, 12 Nov 1984, 
<emphasis italics="true" pageId="6" pageNumber="15">Berry 4402</emphasis>
(US!); between Timotes and Laguna Huacha, 3850 m, 21-26 Nov 1959, 
<emphasis italics="true" pageId="6" pageNumber="15">Barclay &amp; Juajibioy 9853</emphasis>
, 9889 (US!); Sierra del Norte west of La Culata: Alto de Piedras Blancas, vert N, in marshy spots, 17 Jul 1970, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; 
<normalizedToken originalValue="López-Figueiras">Lopez-Figueiras</normalizedToken>
369
</emphasis>
(MERF, US!); id., 4100 m, 23 Nov. 1976, 
<emphasis italics="true" pageId="6" pageNumber="15">Quintero 1697</emphasis>
(MER!), 
<emphasis italics="true" pageId="6" pageNumber="15">1698</emphasis>
(MER!); Llano de Piedras Blancas, 4150-4200 m, 16 Jul 1970, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; 
<normalizedToken originalValue="López-Figueiras">Lopez-Figueiras</normalizedToken>
331
</emphasis>
(MERF, US!); id., 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Los Conejos in Zanjas de Los Castillos, 3800 m, 24 Mar 1972, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
6975
</emphasis>
(MERF, US!); id., in Indio Dormido, 3750 m, 23 Jul 1980, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="López-Figueiras">Lopez-Figueiras</normalizedToken>
23688
</emphasis>
(MERF, US!); id., 3500 m, 1933, 
<emphasis italics="true" pageId="6" pageNumber="15">Jahn 235</emphasis>
(US!); id., 24 Sep 1938, 
<emphasis italics="true" pageId="6" pageNumber="15">Hanbury-Tracy 155</emphasis>
(K, NY, US!). 
<taxonomicName authorityName="M. Diazgranados &amp; G. Morillo" authorityYear="2013" class="Magnoliopsida" family="Asteraceae" genus="Coespeletia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="6" pageNumber="15" phylum="Magnoliophyta" rank="species" species="palustris">
<emphasis italics="true" pageId="6" pageNumber="15">Coespeletia palustris</emphasis>
</taxonomicName>
(paratypes): VENEZUELA. Laguna de Los Patos, 4000 m, 23 Nov 1959, 
<emphasis italics="true" pageId="6" pageNumber="15">Barclay &amp; Juajibioy 9741</emphasis>
(US!); id, 6 Feb 1966, 
<emphasis italics="true" pageId="6" pageNumber="15">Schulz 252</emphasis>
(MER!); 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Los Granates, 
<normalizedToken originalValue="Cañada">Canada</normalizedToken>
del Padre, 3550 m, 10 Oct 1969, 
<emphasis italics="true" pageId="6" pageNumber="15">Cuatrecasas</emphasis>
, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; 
<normalizedToken originalValue="López-Figuieras">Lopez-Figuieras</normalizedToken>
28057
</emphasis>
(MERF, US!); id., 3480 m, 10 Oct 1969, 
<emphasis italics="true" pageId="6" pageNumber="15">Cuatrecasas</emphasis>
, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; 
<normalizedToken originalValue="López-Figuieras">Lopez-Figuieras</normalizedToken>
28055
</emphasis>
(MERF, US!); 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Mucubaji, 6100 m, 6 Jun 1952, 
<emphasis italics="true" pageId="6" pageNumber="15">Gines 4801</emphasis>
(US!); id., 3700 m, Sep 1956, 
<emphasis italics="true" pageId="6" pageNumber="15">Aristeguieta 2445</emphasis>
(MER!, US!, VEN); id., 4670 m, 17 Nov 1958, 
<emphasis italics="true" pageId="6" pageNumber="15">Buza 107</emphasis>
(F!, MER!); id., Laguna Negra, 18 May 1952, 
<emphasis italics="true" pageId="6" pageNumber="15">Vareschi &amp; Pannier 1019</emphasis>
(MER!); id., 
<normalizedToken originalValue="Cañada">Canada</normalizedToken>
of Laguna de Los Anteojos, 3930-3900 m, 22 Feb 1973, 
<emphasis italics="true" pageId="6" pageNumber="15">Cuatrecasas</emphasis>
, 
<emphasis italics="true" pageId="6" pageNumber="15">
<normalizedToken originalValue="Ruiz-Terán">Ruiz-Teran</normalizedToken>
&amp; 
<normalizedToken originalValue="López-Figueiras">Lopez-Figueiras</normalizedToken>
28584
</emphasis>
(MERF, US!); 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de 
<normalizedToken originalValue="Mucuñuque">Mucunuque</normalizedToken>
, 4100 m, 20 June 1958, 
<emphasis italics="true" pageId="6" pageNumber="15">Buza 109</emphasis>
(MER!).
</paragraph>
</subSubSection>
</treatment>
</document>