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<document id="A2A055C12270A308A82A9DE3445C2D26" ID-CLB-Dataset="8924" ID-DOI="10.11646/zootaxa.4819.2.2" ID-GBIF-Dataset="8e35f5ad-f97e-432a-9005-3da1fd0bb897" ID-ISSN="1175-5326" ID-Zenodo-Dep="4396828" ID-ZooBank="2459A542-6CF2-4545-9E6F-262C68838D99" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609201221636" checkinUser="plazi" docAuthor="Chamutiová, Tímea, Hamerlík, Ladislav &amp; Bitušík, Peter" docDate="2020" docId="03F487F1FFB3FFAAFF22FCA9AA03FA82" docLanguage="en" docName="zootaxa.4819.2.2.pdf" docOrigin="Zootaxa 4819 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chaetocladius Kieffer 1911" docType="treatment" docVersion="5" lastPageNumber="230" masterDocId="FFCDFF89FFBDFFA4FFB5FFB7AE49FFFB" masterDocTitle="Subfossil chironomids (Diptera, Chironomidae) of lakes in the Tatra Mountains an illustrated guide" masterLastPageNumber="264" masterPageNumber="216" pageNumber="230" updateTime="1698848074966" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3B589ECE6C237F97054181AF34B834F4">Subfossil chironomids (Diptera, Chironomidae) of lakes in the Tatra Mountains an illustrated guide</mods:title>
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<mods:namePart id="88D7C5204CA4DA02E4FAB95E7DA3B5F4">Chamutiová, Tímea</mods:namePart>
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<mods:affiliation id="F3F2D5D89FA69692FFC9C92CEFB31E55">peter. bitusik @ umb. sk; https: // orcid. org / 0000 - 0002 - 8439 - 4582</mods:affiliation>
<mods:nameIdentifier id="464156C605A3A05032A68F97C4C7B8C4" type="email">peter.bitusik@umb.sk</mods:nameIdentifier>
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<mods:namePart id="DBCE29886ABFE345D8179EE8A4D32307">Hamerlík, Ladislav</mods:namePart>
<mods:nameIdentifier id="DB8357C9F44CB69AAF88DBC5C6B86E1D" type="ORCID">0000-0002-0803-8981</mods:nameIdentifier>
<mods:affiliation id="3A3C147FB782014ABB5B2D5B4F79B5B6">ladislav. hamerlik @ umb. sk; https: // orcid. org / 0000 - 0002 - 0803 - 8981</mods:affiliation>
<mods:nameIdentifier id="61400C37BC5403559FF6B1D84B7E8AA6" type="email">ladislav.hamerlik@umb.sk</mods:nameIdentifier>
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<mods:namePart id="E4EAD5C2B0ACCE3EF5AC5F4577B96FBA">Bitušík, Peter</mods:namePart>
<mods:nameIdentifier id="B8B376A78F4BE829553432B7C3306FCE" type="ORCID">0000-0003-4375-7727</mods:nameIdentifier>
<mods:affiliation id="CCA392E7348A0A967B618C07D436D77F">timea. chamutiova @ umb. sk; https: // orcid. org / 0000 - 0003 - 4375 - 7727</mods:affiliation>
<mods:nameIdentifier id="33BD59C208CA087A573694184AD891FE" type="email">timea.chamutiova@umb.sk</mods:nameIdentifier>
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<mods:date id="7AD20B6C10A1761107C2380927328DF7">2020</mods:date>
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<treatment id="03F487F1FFB3FFAAFF22FCA9AA03FA82" ID-DOI="http://doi.org/10.5281/zenodo.4437258" ID-GBIF-Taxon="176583958" ID-Zenodo-Dep="4437258" LSID="urn:lsid:plazi:treatment:03F487F1FFB3FFAAFF22FCA9AA03FA82" httpUri="http://treatment.plazi.org/id/03F487F1FFB3FFAAFF22FCA9AA03FA82" lastPageNumber="230" pageId="14" pageNumber="230">
<subSubSection id="C347656CFFB3FFAAFF22FCA9AC0DFCC3" box="[151,580,798,825]" pageId="14" pageNumber="230" type="nomenclature">
<paragraph id="8BE236E7FFB3FFAAFF22FCA9AC0DFCC3" blockId="14.[151,580,798,825]" box="[151,580,798,825]" pageId="14" pageNumber="230">
<heading id="D0AA818BFFB3FFAAFF22FCA9AC0DFCC3" bold="true" box="[151,580,798,825]" fontSize="11" level="1" pageId="14" pageNumber="230" reason="1">
<emphasis id="B929EAF5FFB3FFAAFF22FCA9AC0DFCC3" bold="true" box="[151,580,798,825]" pageId="14" pageNumber="230">
<taxonomicName id="4C5D4D64FFB3FFAAFF22FCA9AFEDFCC3" ID-CoL="3LKN" ID-ENA="611341" authority="Kieffer" authorityName="Kieffer" authorityYear="1911" box="[151,420,798,825]" class="Insecta" family="Chironomidae" genus="Chaetocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFF22FCA9AF0DFCC2" bold="true" box="[151,324,798,825]" italics="true" pageId="14" pageNumber="230">Chaetocladius</emphasis>
Kieffer
</taxonomicName>
(
<figureCitation id="13662A62FFB3FFAAFE01FCA9AC75FCC3" box="[436,572,798,824]" captionStart="FIGURES 28–31" captionStartId="13.[151,264,1907,1931]" captionTargetBox="[177,1411,669,1884]" captionTargetId="figure-170@13.[177,1411,669,1884]" captionTargetPageId="13" captionText="FIGURES 28–31. Brillia bifida-type: 28—head capsule (arrows indicate fourth and fifth lateral teeth of the mentum and position of seta submenti), A—antenna (arrow indicates second segment). Chaetocladius piger-type: 29—bifid premandible, A—antenna. C. piger-type: 30—labral lamellae (arrow indicates its position). C. piger-type: 31—head capsule (arrow indicates seta submenti), A—detail of mentum (arrows indicate minute last lateral tooth and bulbous ventromental plate)." figureDoi="http://doi.org/10.5281/zenodo.4396844" httpUri="https://zenodo.org/record/4396844/files/figure.png" pageId="14" pageNumber="230">Figs 29–31</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C347656CFFB3FFAAFF22FCD0AAA7FBEA" pageId="14" pageNumber="230" type="description">
<paragraph id="8BE236E7FFB3FFAAFF22FCD0AAA7FBEA" blockId="14.[151,1437,870,1401]" pageId="14" pageNumber="230">
Head yellow to light brown with darkened occipital margin. Antenna 5 segmented, never strongly reduced (
<figureCitation id="13662A62FFB3FFAAFADAFCD0AE82FC5E" captionStart="FIGURES 28–31" captionStartId="13.[151,264,1907,1931]" captionTargetBox="[177,1411,669,1884]" captionTargetId="figure-170@13.[177,1411,669,1884]" captionTargetPageId="13" captionText="FIGURES 28–31. Brillia bifida-type: 28—head capsule (arrows indicate fourth and fifth lateral teeth of the mentum and position of seta submenti), A—antenna (arrow indicates second segment). Chaetocladius piger-type: 29—bifid premandible, A—antenna. C. piger-type: 30—labral lamellae (arrow indicates its position). C. piger-type: 31—head capsule (arrow indicates seta submenti), A—detail of mentum (arrows indicate minute last lateral tooth and bulbous ventromental plate)." figureDoi="http://doi.org/10.5281/zenodo.4396844" httpUri="https://zenodo.org/record/4396844/files/figure.png" pageId="14" pageNumber="230">Fig. 29A</figureCitation>
). Premandible bifid with 1–2 blunt inner teeth (
<figureCitation id="13662A62FFB3FFAAFD50FC3CAD73FC5E" box="[741,826,907,933]" captionStart="FIGURES 28–31" captionStartId="13.[151,264,1907,1931]" captionTargetBox="[177,1411,669,1884]" captionTargetId="figure-170@13.[177,1411,669,1884]" captionTargetPageId="13" captionText="FIGURES 28–31. Brillia bifida-type: 28—head capsule (arrows indicate fourth and fifth lateral teeth of the mentum and position of seta submenti), A—antenna (arrow indicates second segment). Chaetocladius piger-type: 29—bifid premandible, A—antenna. C. piger-type: 30—labral lamellae (arrow indicates its position). C. piger-type: 31—head capsule (arrow indicates seta submenti), A—detail of mentum (arrows indicate minute last lateral tooth and bulbous ventromental plate)." figureDoi="http://doi.org/10.5281/zenodo.4396844" httpUri="https://zenodo.org/record/4396844/files/figure.png" pageId="14" pageNumber="230">Fig. 29</figureCitation>
). Labral lamellae of various shape usually well developed between labral setae SI, but can be vestigial or absent (
<figureCitation id="13662A62FFB3FFAAFC9EFC18AD37FC32" box="[811,894,943,969]" captionStart="FIGURES 28–31" captionStartId="13.[151,264,1907,1931]" captionTargetBox="[177,1411,669,1884]" captionTargetId="figure-170@13.[177,1411,669,1884]" captionTargetPageId="13" captionText="FIGURES 28–31. Brillia bifida-type: 28—head capsule (arrows indicate fourth and fifth lateral teeth of the mentum and position of seta submenti), A—antenna (arrow indicates second segment). Chaetocladius piger-type: 29—bifid premandible, A—antenna. C. piger-type: 30—labral lamellae (arrow indicates its position). C. piger-type: 31—head capsule (arrow indicates seta submenti), A—detail of mentum (arrows indicate minute last lateral tooth and bulbous ventromental plate)." figureDoi="http://doi.org/10.5281/zenodo.4396844" httpUri="https://zenodo.org/record/4396844/files/figure.png" pageId="14" pageNumber="230">Fig. 30</figureCitation>
). Apical mandibular tooth shorter than combined width of 3 inner teeth. Mentum with 1–2 median teeth and 5 pairs of lateral teeth, the outermost tooth may be reduced. Ventromental plates usually large, rounded and extending beyond outer lateral mental tooth.
</paragraph>
</subSubSection>
<subSubSection id="C347656CFFB3FFAAFF72FBACAA03FA82" pageId="14" pageNumber="230" type="discussion">
<paragraph id="8BE236E7FFB3FFAAFF72FBACAC99FAF6" blockId="14.[151,1437,870,1401]" pageId="14" pageNumber="230">
Remarks: The shape of mentum is often the only character that can be used for the identification of subfossils.
<emphasis id="B929EAF5FFB3FFAAFF22FB88AD0BFBA2" box="[151,834,1087,1113]" italics="true" pageId="14" pageNumber="230">
<taxonomicName id="4C5D4D64FFB3FFAAFF22FB88AF26FBA3" authorityName="Goetghebuer" authorityYear="1932" box="[151,367,1087,1112]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">Parametriocnemus</taxonomicName>
,
<taxonomicName id="4C5D4D64FFB3FFAAFEC9FB88AC13FBA2" authorityName="Thienemann" authorityYear="1924" box="[380,602,1087,1113]" class="Insecta" family="Chironomidae" genus="Paraphaenocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">Paraphaenocladius</taxonomicName>
,
<taxonomicName id="4C5D4D64FFB3FFAAFDD3FB88AD0BFBA2" box="[614,834,1087,1113]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">Heterotrissocladius</taxonomicName>
</emphasis>
have mentum with double median tooth, and 5 pairs of laterals, combined with large ventromental plates extending beyond the outer lateral teeth without beard. Setae submenti of
<taxonomicName id="4C5D4D64FFB3FFAAFE95FB30AF8BFB5A" authorityName="Kieffer" authorityYear="1911" box="[288,450,1159,1185]" class="Insecta" family="Chironomidae" genus="Chaetocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFE95FB30AF8BFB5A" box="[288,450,1159,1185]" italics="true" pageId="14" pageNumber="230">Chaetocladius</emphasis>
</taxonomicName>
are arising under the ventromental plates (
<figureCitation id="13662A62FFB3FFAAFC2DFB30ADA1FB5A" box="[920,1000,1159,1185]" captionStart="FIGURES 28–31" captionStartId="13.[151,264,1907,1931]" captionTargetBox="[177,1411,669,1884]" captionTargetId="figure-170@13.[177,1411,669,1884]" captionTargetPageId="13" captionText="FIGURES 28–31. Brillia bifida-type: 28—head capsule (arrows indicate fourth and fifth lateral teeth of the mentum and position of seta submenti), A—antenna (arrow indicates second segment). Chaetocladius piger-type: 29—bifid premandible, A—antenna. C. piger-type: 30—labral lamellae (arrow indicates its position). C. piger-type: 31—head capsule (arrow indicates seta submenti), A—detail of mentum (arrows indicate minute last lateral tooth and bulbous ventromental plate)." figureDoi="http://doi.org/10.5281/zenodo.4396844" httpUri="https://zenodo.org/record/4396844/files/figure.png" pageId="14" pageNumber="230">Fig. 31</figureCitation>
) while there are located much higher in
<taxonomicName id="4C5D4D64FFB3FFAAFF22FB1CAF25FB3F" authorityName="Goetghebuer" authorityYear="1932" box="[151,364,1195,1220]" class="Insecta" family="Chironomidae" genus="Parametriocnemus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFF22FB1CAF25FB3F" box="[151,364,1195,1220]" italics="true" pageId="14" pageNumber="230">Parametriocnemus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D4D64FFB3FFAAFE16FB1CAC37FB3E" authorityName="Thienemann" authorityYear="1924" box="[419,638,1195,1221]" class="Insecta" family="Chironomidae" genus="Paraphaenocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFE16FB1CAC37FB3E" box="[419,638,1195,1221]" italics="true" pageId="14" pageNumber="230">Paraphaenocladius</emphasis>
</taxonomicName>
(see
<figureCitation id="13662A62FFB3FFAAFD0FFB1CAD47FB3E" box="[698,782,1194,1221]" captionStart="FIGURES 54–59" captionStartId="24.[151,264,1873,1897]" captionTargetBox="[177,1411,309,1849]" captionTargetId="figure-55@24.[177,1411,309,1849]" captionTargetPageId="24" captionText="FIGURES 54–59. Metriocnemus fuscipes-type: 54—head capsule. Orthocladius (Orthocladius) dentifer-type: 55—head capsule. Orthocladius (Mesorthocladius) frigidus: 56—head capsule (arrow indicates seta submenti). Orthocladius (Euorthocladius): 57—head capsule. Parametriocnemus/Paraphaenocladius: 58 (arrow indicates seta submenti). Parorthocladius: 59—head capsule." figureDoi="http://doi.org/10.5281/zenodo.4396858" httpUri="https://zenodo.org/record/4396858/files/figure.png" pageId="14" pageNumber="230">Fig. 58</figureCitation>
). The outermost mental tooth is not reduced in
<taxonomicName id="4C5D4D64FFB3FFAAFA94FB1CAF48FB12" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFA94FB1CAF48FB12" italics="true" pageId="14" pageNumber="230">Heterotrissocladius</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5D4D64FFB3FFAAFEBBFB78AC18FB12" authority=", Hydrobeanus" authorityName="Hydrobeanus" box="[270,593,1231,1257]" class="Insecta" family="Chironomidae" genus="Metriocnemus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFEBBFB78AC18FB12" box="[270,593,1231,1257]" italics="true" pageId="14" pageNumber="230">Metriocnemus, Hydrobeanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D4D64FFB3FFAAFD31FB78AD6DFB12" box="[644,804,1231,1257]" class="Insecta" family="Chironomidae" genus="Thienemannia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFD31FB78AD6DFB12" box="[644,804,1231,1257]" italics="true" pageId="14" pageNumber="230">Thienemannia</emphasis>
</taxonomicName>
have similar menta, however without large ventromental plates and obviously narrower median teeth/ tooth.
</paragraph>
<paragraph id="8BE236E7FFB3FFAAFF72FAA0AFD6FAAE" blockId="14.[151,1437,870,1401]" pageId="14" pageNumber="230">
The only
<taxonomicName id="4C5D4D64FFB3FFAAFE83FAA0AF91FACA" authorityName="Kieffer" authorityYear="1911" box="[310,472,1303,1329]" class="Insecta" family="Chironomidae" genus="Chaetocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF5FFB3FFAAFE83FAA0AF91FACA" box="[310,472,1303,1329]" italics="true" pageId="14" pageNumber="230">Chaetocladius</emphasis>
</taxonomicName>
identified to species level from the Tatra Mts. is
<taxonomicName id="4C5D4D64FFB3FFAAFBBAFAA0AB05FACA" authority="Moubayed, 2019" authorityName="Moubayed" authorityYear="2019" box="[1039,1356,1303,1329]" class="Insecta" family="Chironomidae" genus="Chaetocladius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="230" phylum="Arthropoda" rank="species" species="bitusiki">
<emphasis id="B929EAF5FFB3FFAAFBBAFAA0AACAFACA" box="[1039,1155,1303,1329]" italics="true" pageId="14" pageNumber="230">C. bitusiki</emphasis>
Moubayed, 2019
</taxonomicName>
(Moubayed &amp; Bitusik 2019).
</paragraph>
<paragraph id="8BE236E7FFB3FFAAFF72FAE8AA03FA82" blockId="14.[151,1437,870,1401]" box="[199,1098,1375,1401]" pageId="14" pageNumber="230">
The morphotype recognized in our material corresponds to
<bibRefCitation id="EFCC4B16FFB3FFAAFCD4FAE8AA0CFA82" author="Brooks, S. J. &amp; Langdon, P. G. &amp; Heiri, O." box="[865,1093,1375,1401]" pageId="14" pageNumber="230" refId="ref23398" refString="Brooks, S. J., Langdon, P. G. &amp; Heiri, O. (2007) The Identification and Use of Palaearctic Chironomidae Larvae in Palaeoecology. QRA Technical Guide No. 10. QRA, London, 276 pp." type="book" year="2007">
Brooks
<emphasis id="B929EAF5FFB3FFAAFC0CFAE8ADA3FA82" box="[953,1002,1375,1401]" italics="true" pageId="14" pageNumber="230">et al</emphasis>
. (2007)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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