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<document id="F96EFC259B80416911671A1D12882F93" ID-DOI="10.11646/zootaxa.4032.4.1" ID-GBIF-Dataset="1459e41f-b82d-4c88-be4d-ee86e329f9c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="243485" ID-ZooBank="AFEF8567-B7C6-4F54-AC51-4C990389BF1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461294283733" checkinUser="plazi" docAuthor="Kustov, Semen &amp; Zherebilo, Danil" docDate="2015" docId="03F487D4FF881A70FF04FD84B9E2FA5B" docLanguage="en" docName="zt04032p369.pdf" docOrigin="Zootaxa 4032 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wiedemannia (Philolutra) vaillanti Joost" docType="treatment" docVersion="9" lastPageNumber="366" masterDocId="FFCDFFACFF871A7FFF93FFA0BD01FFCD" masterDocTitle="New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species" masterLastPageNumber="369" masterPageNumber="351" pageNumber="366" updateTime="1698616197258" updateUser="plazi">
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<mods:title id="4A99706971668BB7D4AB8C6540A1CE5F">New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species</mods:title>
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<mods:name id="0B765DE5BCBF0CED73B3AE2550AF2537" type="personal">
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<mods:namePart id="5BE8BA55A5EB686D5B3E68E1C6DBF5CE">Kustov, Semen</mods:namePart>
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<mods:namePart id="D0CD4EEE045FDC9823D6E75ABA5DCFC5">Zherebilo, Danil</mods:namePart>
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<mods:title id="24548944DE0599F098A9DE5D0440B653">Zootaxa</mods:title>
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<mods:date id="BDD68AA89A55DE09FE5061D819A9D94F">2015</mods:date>
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<treatment id="03F487D4FF881A70FF04FD84B9E2FA5B" ID-DOI="http://doi.org/10.5281/zenodo.5691045" ID-GBIF-Taxon="119642277" ID-Zenodo-Dep="5691045" LSID="urn:lsid:plazi:treatment:03F487D4FF881A70FF04FD84B9E2FA5B" httpUri="http://treatment.plazi.org/id/03F487D4FF881A70FF04FD84B9E2FA5B" lastPageNumber="366" pageId="15" pageNumber="366">
<subSubSection id="C3476549FF881A70FF04FD84BDF8FD92" pageId="15" pageNumber="366" type="nomenclature">
<paragraph id="8BE236C2FF881A70FF04FD84BF86FDF3" blockId="15.[151,647,548,607]" box="[151,647,548,574]" pageId="15" pageNumber="366">
<heading id="D0AA81AEFF881A70FF04FD84BF86FDF3" bold="true" box="[151,647,548,574]" fontSize="11" level="1" pageId="15" pageNumber="366" reason="1">
<taxonomicName id="4C5D4D41FF881A70FF04FD84BF86FDF3" ID-CoL="8FM83" authority="Joost" authorityName="Joost" box="[151,647,548,574]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="vaillanti" subGenus="Philolutra">
<emphasis id="B929EAD0FF881A70FF04FD84BF86FDF3" bold="true" box="[151,647,548,574]" pageId="15" pageNumber="366">
<emphasis id="B929EAD0FF881A70FF04FD84BC3FFDF3" bold="true" box="[151,318,548,574]" italics="true" pageId="15" pageNumber="366">Wiedemannia</emphasis>
(
<emphasis id="B929EAD0FF881A70FEDDFD84BCCBFDF3" bold="true" box="[334,458,548,574]" italics="true" pageId="15" pageNumber="366">Philolutra</emphasis>
)
<emphasis id="B929EAD0FF881A70FE48FD84BF3FFDF3" bold="true" box="[475,574,548,574]" italics="true" pageId="15" pageNumber="366">vaillanti</emphasis>
Joost
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE236C2FF881A70FF04FDE6BDF8FD92" blockId="15.[151,647,548,607]" box="[151,249,582,607]" pageId="15" pageNumber="366">
(
<figureCitation id="13662A47FF881A70FF0CFDE6BDF0FD92" box="[159,241,582,607]" captionStart="FIGURES 12 17" captionStartId="12.[151,263,1937,1959]" captionTargetBox="[247,1322,214,1871]" captionTargetId="figure@12.[229,1337,193,1891]" captionTargetPageId="12" captionText="FIGURES 12 17. Male terminalia of Wiedemannia, lateral view. 12, W. pseguashae sp. nov.; 13, W. pseudovaillanti Joost; 14, W. shamshevi Kustov &amp; Zherebilo; 15, W. sinclairi Kustov &amp; Zherebilo; 16, W. vaillanti Joost; 17, W. zetterstedti (Fallén). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/243488/files/figure.png" pageId="15" pageNumber="366">Fig. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF881A70FF04FD2EBFAFFD69" box="[151,686,654,676]" pageId="15" pageNumber="366" type="reference_group">
<paragraph id="8BE236C2FF881A70FF04FD2EBFAFFD69" blockId="15.[151,686,654,676]" box="[151,686,654,676]" pageId="15" pageNumber="366">
<treatmentCitationGroup id="AB4D11ECFF881A70FF04FD2EBFAFFD69" box="[151,686,654,676]" pageId="15" pageNumber="366">
<taxonomicName id="4C5D4D41FF881A70FF04FD2EBFAAFD69" ID-CoL="8FM83" authority="Joost, 1981: 189" authorityName="Joost" authorityPageNumber="189" authorityYear="1981" box="[151,683,654,676]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="vaillanti" subGenus="Philolutra">
<emphasis id="B929EAD0FF881A70FF04FD2EBC20FD69" box="[151,289,654,676]" italics="true" pageId="15" pageNumber="366">Wiedemannia</emphasis>
(
<emphasis id="B929EAD0FF881A70FEBCFD2EBC96FD69" box="[303,407,654,676]" italics="true" pageId="15" pageNumber="366">Philolutra</emphasis>
)
<emphasis id="B929EAD0FF881A70FE35FD2EBCFBFD69" box="[422,506,654,676]" italics="true" pageId="15" pageNumber="366">vaillanti</emphasis>
<treatmentCitation id="0AFC10D3FF881A70FD92FD2FBFAAFD69" author="Joost" box="[513,683,654,676]" page="189" pageId="15" pageNumber="366" year="1981">
<bibRefCitation id="EFCC4B33FF881A70FD92FD2FBF79FD69" author="Joost" box="[513,632,654,676]" pageId="15" pageNumber="366" refString="Joost, W. (1981) Beitrag zur Kenntnis der Hemerodromiiniae des Kaukasus (I) (Diptera, Empididae). Reichenbachia, 19 (31), 183 - 191." type="journal article" year="1981">Joost, 1981</bibRefCitation>
: 189
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF881A70FF04FD71BCA3FBC7" pageId="15" pageNumber="366" type="materials_examined">
<paragraph id="8BE236C2FF881A70FF04FD71BCA3FBC7" blockId="15.[151,1437,721,1430]" pageId="15" pageNumber="366">
<emphasis id="B929EAD0FF881A70FF04FD71BC7DFD27" bold="true" box="[151,380,721,746]" pageId="15" pageNumber="366">Material examined</emphasis>
.
<emphasis id="B929EAD0FF881A70FE1FFD71BCF3FD27" bold="true" box="[396,498,721,746]" pageId="15" pageNumber="366">
<collectingCountry id="F34A7652FF881A70FE1FFD71BCF3FD27" box="[396,498,721,746]" name="Russia" pageId="15" pageNumber="366">RUSSIA</collectingCountry>
</emphasis>
: Krasnodarskiy Territory, Nature Reserve “Kamyshanova Polyana” (7),
<date id="FFE31002FF881A70FAA6FD72B896FD27" box="[1333,1431,722,746]" pageId="15" pageNumber="366" value="2014-05-02">2.v.2014</date>
, S.Yu. Kustov (
<specimenCount id="9D5BFD4BFF881A70FEA9FD56BC64FCC0" box="[314,357,758,782]" pageId="15" pageNumber="366" type="generic">2 ♂</specimenCount>
, 1 ♀); Nature Reserve “Kamyshanova Polyana” (8),
<date id="FFE31002FF881A70FC2BFD56B91BFCC3" box="[952,1050,758,782]" pageId="15" pageNumber="366" value="2014-05-04">4.v.2014</date>
, S.Yu. Kustov (
<specimenCount id="9D5BFD4BFF881A70FB5AFD56B801FCC0" box="[1225,1280,758,782]" pageId="15" pageNumber="366" type="generic">11 ♂</specimenCount>
, 4 ♀); Nature Reserve “Kamyshanova Polyana” (6),
<date id="FFE31002FF881A70FDD4FCB9BFC5FCFF" box="[583,708,793,818]" pageId="15" pageNumber="366" value="2014-06-13">13.vi.2014</date>
, S.Yu. Kustov (
<specimenCount id="9D5BFD4BFF881A70FCEFFCBABEADFCFC" box="[892,940,794,818]" pageId="15" pageNumber="366" type="generic">4 ♂</specimenCount>
, 3 ♀). Karachay-Cherkess Republic, River Phiya (16),
<date id="FFE31002FF881A70FE89FC9DBC94FC9B" box="[282,405,829,854]" pageId="15" pageNumber="366" value="2013-06-10">10.vi.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FDACFC9DBF79FC98" box="[575,632,829,854]" pageId="15" pageNumber="366" type="generic">59 ♂</specimenCount>
, 47 ♀); env. Rozhkao (13),
<date id="FFE31002FF881A70FC21FC9DB91EFC9B" box="[946,1055,829,854]" pageId="15" pageNumber="366" value="2013-06-08">8.vi.2013</date>
, D. Zherebilo (1 ♀); watershed of River Bolshaya Laba (12),
<date id="FFE31002FF881A70FE59FCC1BF37FCB7" box="[458,566,865,890]" pageId="15" pageNumber="366" value="2013-06-09">9.vi.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FD71FCC2BE11FCB4" box="[738,784,866,890]" pageId="15" pageNumber="366" type="generic">2 ♂</specimenCount>
, 7 ♀).
<collectingCountry id="F34A7652FF881A70FCCCFCC1BEE7FCB7" box="[863,998,865,890]" name="Georgia" pageId="15" pageNumber="366">
<emphasis id="B929EAD0FF881A70FCCCFCC1BEE7FCB7" bold="true" box="[863,998,865,890]" pageId="15" pageNumber="366">GEORGIA</emphasis>
</collectingCountry>
: Lechkhumi ridge (41),
<date id="FFE31002FF881A70FB6FFCC1B874FCB7" box="[1276,1397,865,890]" pageId="15" pageNumber="366" value="2014-04-28">28.iv.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FE8BFC26BC49FC50" box="[280,328,902,926]" pageId="15" pageNumber="366" type="generic">2 ♂</specimenCount>
, 1 ♀); Mingrelia, ridge Ofitsari (31),
<date id="FFE31002FF881A70FC80FC26BE77FC53" box="[787,886,902,926]" pageId="15" pageNumber="366" value="2014-05-03">3.v.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FBBCFC26B95EFC50" box="[1071,1119,902,926]" pageId="15" pageNumber="366" type="generic">2 ♂</specimenCount>
, 1 ♀); ridge Ofitsari (30),
<date id="FFE31002FF881A70FF04FC0ABDF8FC0F" box="[151,249,938,962]" pageId="15" pageNumber="366" value="2014-05-04">4.v.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FE20FC0ABCE3FC0F" box="[435,482,938,962]" pageId="15" pageNumber="366" type="generic">1 ♂</specimenCount>
); same data,
<date id="FFE31002FF881A70FD11FC09BFE4FC0F" box="[642,741,937,962]" pageId="15" pageNumber="366" value="2014-05-06">6.v.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FC08FC0ABECAFC0C" box="[923,971,938,962]" pageId="15" pageNumber="366" type="generic">4 ♂</specimenCount>
, 2 ♀); ridge Ofitsari (32),
<date id="FFE31002FF881A70FA9DFC09B870FC0F" box="[1294,1393,937,962]" pageId="15" pageNumber="366" value="2014-05-05">5.v.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FE8BFC6EBC44FC2B" box="[280,325,974,998]" pageId="15" pageNumber="366" type="generic">1 ♂</specimenCount>
).
<emphasis id="B929EAD0FF881A70FECFFC6DBCF4FC2B" bold="true" box="[348,501,973,998]" pageId="15" pageNumber="366">ABKHAZIA</emphasis>
: River Bzyb (20),
<date id="FFE31002FF881A70FD5CFC6DBE4BFC2B" box="[719,842,973,998]" pageId="15" pageNumber="366" value="2013-06-30">30.vi.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FC68FC6EB926FC28" box="[1019,1063,974,998]" pageId="15" pageNumber="366" type="generic">3 ♂</specimenCount>
, 2 ♀); same data,
<date id="FFE31002FF881A70FB6CFC6DB872FC2B" box="[1279,1395,973,998]" pageId="15" pageNumber="366" value="2013-07-01">1.vii.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF881A70FE80FC52BC4FFBC4" box="[275,334,1010,1034]" pageId="15" pageNumber="366" type="generic">41 ♂</specimenCount>
, 32 ♀).
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF881A70FF54FBB5B955FB54" pageId="15" pageNumber="366" type="diagnosis">
<paragraph id="8BE236C2FF881A70FF54FBB5B955FB54" blockId="15.[151,1437,721,1430]" pageId="15" pageNumber="366">
<emphasis id="B929EAD0FF881A70FF54FBB5BC3BFBE3" bold="true" box="[199,314,1045,1070]" pageId="15" pageNumber="366">Diagnosis</emphasis>
.
<taxonomicName id="4C5D4D41FF881A70FEDEFBB5BFE0FBE0" box="[333,737,1045,1070]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="vaillanti" subGenus="Philolutra">
<emphasis id="B929EAD0FF881A70FEDEFBB5BCE7FBE0" box="[333,486,1045,1069]" italics="true" pageId="15" pageNumber="366">Wiedemannia</emphasis>
(
<emphasis id="B929EAD0FF881A70FE68FBB5BF6FFBE0" box="[507,622,1045,1069]" italics="true" pageId="15" pageNumber="366">Philolutra</emphasis>
)
<emphasis id="B929EAD0FF881A70FD17FBB5BFE0FBE0" box="[644,737,1045,1069]" italics="true" pageId="15" pageNumber="366">vaillanti</emphasis>
</taxonomicName>
differs by the completely dark brown scutum, acrostichals divergent around prescutellar depression, 5 pairs of dorsocentrals without interspersed shorter setae, fore femur apically without strong anterior setae; clasping cercus with anterior lobe prolonged, digitiform and slender, without cluster of apical setae; dorsal process anvil-shaped, inner margin without dense setae.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF881A70FF54FB05BE89FB2F" pageId="15" pageNumber="366" type="distribution">
<paragraph id="8BE236C2FF881A70FF54FB05BE89FB2F" blockId="15.[151,1437,721,1430]" pageId="15" pageNumber="366">
<emphasis id="B929EAD0FF881A70FF54FB05BC58FB73" bold="true" box="[199,345,1189,1214]" pageId="15" pageNumber="366">Distribution</emphasis>
. PALAEARCTIC: Caucasus:
<collectingCountry id="F34A7652FF881A70FD2CFB05BE0BFB73" box="[703,778,1189,1214]" name="Russia" pageId="15" pageNumber="366">Russia</collectingCountry>
(Krasnodarsky Territory, Karachay-Cherkess Republic),
<collectingCountry id="F34A7652FF881A70FF04FB69BDF5FB2F" box="[151,244,1225,1250]" name="Georgia" pageId="15" pageNumber="366">Georgia</collectingCountry>
, Abkhazia; Asia:
<collectingCountry id="F34A7652FF881A70FE28FB69BF0AFB2F" box="[443,523,1225,1250]" name="Turkey" pageId="15" pageNumber="366">Turkey</collectingCountry>
(Bolu: Yedigöller National Park).
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF881A70FF54FB4DB9E2FA5B" pageId="15" pageNumber="366" type="discussion">
<paragraph id="8BE236C2FF881A70FF54FB4DB9E2FA5B" blockId="15.[151,1437,721,1430]" pageId="15" pageNumber="366">
<emphasis id="B929EAD0FF881A70FF54FB4DBC35FACB" bold="true" box="[199,308,1261,1286]" pageId="15" pageNumber="366">Remarks</emphasis>
. This species was described by
<bibRefCitation id="EFCC4B33FF881A70FD40FB4EBE66FACB" author="Joost" box="[723,871,1262,1286]" pageId="15" pageNumber="366" refString="Joost, W. (1981) Beitrag zur Kenntnis der Hemerodromiiniae des Kaukasus (I) (Diptera, Empididae). Reichenbachia, 19 (31), 183 - 191." type="journal article" year="1981">Joost (1981)</bibRefCitation>
from Tscheget, Terskol and Dombai (
<collectingCountry id="F34A7652FF881A70FADBFB4DB899FACB" box="[1352,1432,1261,1286]" name="Russia" pageId="15" pageNumber="366">Russia</collectingCountry>
: Kabardino-Balkaria Republic and Karachay-Cherkessia Republic), but
<bibRefCitation id="EFCC4B33FF881A70FC2FFAB1B9DBFAE7" author="Chvala" box="[956,1242,1297,1322]" pageId="15" pageNumber="366" refString="Chvala, M. &amp; Wagner, R. (1989) Hybotidae. In: Catalogue of Palaearctic Diptera. Volume 6. Therevidae-Empididae. Elsevier Science Publishing, pp. 228 - 336." type="book" year="1989">Chvála &amp; Wagner (1989)</bibRefCitation>
in the Catalogue of Palaearctic
<taxonomicName id="4C5D4D41FF881A70FEABFA95BC8DFA83" box="[312,396,1333,1358]" kingdom="Plantae" order="Diptera" pageId="15" pageNumber="366" rank="order">Diptera</taxonomicName>
mistakenly indicated the species from the Georgian territory. Therefore
<taxonomicName id="4C5D4D41FF881A70FB28FA96B83DFA80" box="[1211,1340,1333,1357]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="vaillanti">
<emphasis id="B929EAD0FF881A70FB28FA96B83DFA80" box="[1211,1340,1333,1357]" italics="true" pageId="15" pageNumber="366">W. vaillanti</emphasis>
</taxonomicName>
is noted from the territories of
<collectingCountry id="F34A7652FF881A70FE0EFAF9BCF6FABF" box="[413,503,1369,1394]" name="Georgia" pageId="15" pageNumber="366">Georgia</collectingCountry>
and Abkhazia for the first time. This common species is known from mountain rivers of the Caucasus from elevations between
<quantity id="4CA59B27FF881A70FD23FADEBE3CFA58" box="[688,829,1406,1430]" metricMagnitude="3" metricUnit="m" metricValue="1.44" metricValueMax="2.1" metricValueMin="0.78" pageId="15" pageNumber="366" unit="m" value="1440.0" valueMax="2100.0" valueMin="780.0">7802100 m</quantity>
a.s.l. and is active from April to July.
</paragraph>
</subSubSection>
</treatment>
</document>