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<document id="0288E23D0835E393454C115F17935C6B" ID-DOI="10.11646/phytotaxa.188.1.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="5147105" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627609653251" checkinUser="felipe" docAuthor="Niissalo, Matti A., Wijedasa, Lahiru S., Boyce, Peter C. &amp; Leong-Škorničková, Jana" docDate="2014" docId="15704A5FFFEE8465FF3FFC7FF2E2F8E6" docLanguage="en" docName="Phytotaxa.188.1.014-020.pdf" docOrigin="Phytotaxa 188 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.188.1.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Hanguana Blume 1827" docType="key" docVersion="3" lastPageNumber="18" masterDocId="E9493227FFEA8461FFB7FFB3F749FFF5" masterDocTitle="Hanguana neglecta (Hanguanaceae): a new plant species from a heavily collected and visited reserve in Singapore" masterLastPageNumber="20" masterPageNumber="14" pageNumber="18" updateTime="1699030861252" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="58B2CEC835C9351F8E5A04B5DD33C481">Hanguana neglecta (Hanguanaceae): a new plant species from a heavily collected and visited reserve in Singapore</mods:title>
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<mods:namePart id="F08EDF6BCE5B6B583D27321833F08133">Niissalo, Matti A.</mods:namePart>
<mods:affiliation id="F0E3EF326B669CD894A94E2C24C2A56E">Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543.</mods:affiliation>
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<mods:namePart id="D6090C4A8CB4A5244EE801EC074902A2">Wijedasa, Lahiru S.</mods:namePart>
<mods:affiliation id="DD62CE8A99C6AF34B7980B5DF6E76ACB">Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543. &amp; Herbarium, Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569. &amp; Rimba, 18 E Kampung Basung, Kuala Berang 21700, Terengganu, Malaysia.</mods:affiliation>
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<mods:namePart id="849E5FE5B03D11B5AF562CA7F185FDFB">Boyce, Peter C.</mods:namePart>
<mods:affiliation id="627D50E345A72CC938598E0720F37442">Institute of Biodiversity and Environmental Conservation (IBEC), Universiti Malaysia Sarawak, 94300 Samarahan, Sarawak, Malaysia. * Corresponding author</mods:affiliation>
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<mods:namePart id="BA4EDA80018B776E37E0510941C50350">Leong-Škorničková, Jana</mods:namePart>
<mods:affiliation id="AD7325738CC81F0B3A1668140F171F95">Herbarium, Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569.</mods:affiliation>
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<mods:title id="73FE5C7D435D2C8AB075C1BB22061C2D">Phytotaxa</mods:title>
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<mods:date id="D54656E6A769569D3112DF5BC38603E9">2014</mods:date>
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<heading id="C62E4C25FFEE8465FF3FFC7FF401FC13" bold="true" box="[136,840,972,998]" fontSize="11" level="1" pageId="4" pageNumber="18" reason="1">
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Key to
<taxonomicName id="5AD980CAFFEE8465FF55FC7FF62DFC13" ID-CoL="8VY3X" authorityName="Blume" authorityYear="1827" box="[226,356,972,998]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="AFAD275BFFEE8465FF55FC7FF62DFC13" bold="true" box="[226,356,972,998]" italics="true" pageId="4" pageNumber="18">Hanguana</emphasis>
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in Peninsular
<collectingCountry id="E5CEBBD9FFEE8465FDAEFC7FF5C2FC13" box="[537,651,972,998]" name="Malaysia" pageId="4" pageNumber="18">Malaysia</collectingCountry>
and
<collectingCountry id="E5CEBBD9FFEE8465FD7EFC7FF401FC13" box="[713,840,972,998]" name="Singapore" pageId="4" pageNumber="18">Singapore</collectingCountry>
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<subSubSection id="D5C3A8C2FFEE8465FF3FFBA2F2E2F8E6" pageId="4" pageNumber="18" type="key">
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<paragraph id="9D66FB49FFEE8465FF3FFBA2F2E2FBD3" blockId="4.[136,1451,1040,1091]" box="[136,1451,1040,1062]" pageId="4" pageNumber="18">
<keyLead id="2628587CFFEE8465FF3FFBA2F2E2FBD3" box="[136,1451,1040,1062]" pageId="4" pageNumber="18">1. Stoloniferous colonial helophytes ................................................................................................................................................… 2</keyLead>
</paragraph>
<paragraph id="9D66FB49FFEE8465FF3FFB9DF2E2FBB7" blockId="4.[136,1451,1040,1091]" box="[136,1451,1069,1091]" pageId="4" pageNumber="18">
<keyLead id="2628587CFFEE8465FF3FFB9DF2E2FBB7" box="[136,1451,1069,1091]" pageId="4" pageNumber="18"> Clumping mesophytes lacking stolons .........................................................................................................................................… 3</keyLead>
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<paragraph id="9D66FB49FFEE8465FF3FFBD4F2E3FB43" blockId="4.[136,1452,1127,1264]" pageId="4" pageNumber="18">
<keyLead id="2628587CFFEE8465FF3FFBD4F2E3FB43" pageId="4" pageNumber="18">
2. Leaves stiffly erect, acute; fruits ellipsoid, ripening glossy purple-red; stigma lobes flat, connate at base, the whole
<quantity id="5A2156ACFFEE8465FA97FBD4F226FB89" box="[1312,1391,1127,1148]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="18" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
diam. and almost obscuring the end of the fruit. Plants of open situations along muddy banks of large rivers, margins of freshwater bodies, and of freshwater swamp forest ...........................................................................................................................
<taxonomicName id="5AD980CAFFEE8465FA9CFB12F2E3FB43" ID-CoL="3JJTD" authorityName="Merrill" authorityYear="1915" baseAuthorityName="Jack" baseAuthorityYear="1820" box="[1323,1450,1184,1206]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="species" species="malayana">
<emphasis id="AFAD275BFFEE8465FA9CFB12F2E3FB43" bold="true" box="[1323,1450,1184,1206]" italics="true" pageId="4" pageNumber="18">H. malayana</emphasis>
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Leaves arching, long-attenuate; fruits globose, ripening semiglossy black; stigma lobes small, separate, erect, pointed. Plants of shaded peatswamp mires .....................................................................................................................................................
<taxonomicName id="5AD980CAFFEE8465FAE7FB69F2E2FB1A" ID-CoL="3JJTF" box="[1360,1451,1242,1263]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="species" species="nitens">
<emphasis id="AFAD275BFFEE8465FAE7FB69F2E2FB1A" bold="true" box="[1360,1451,1242,1263]" italics="true" pageId="4" pageNumber="18">H. nitens</emphasis>
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<paragraph id="9D66FB49FFEE8465FF3FFAA7F2E2FADC" blockId="4.[136,1451,1300,1350]" box="[136,1451,1300,1322]" pageId="4" pageNumber="18">
<keyLead id="2628587CFFEE8465FF3FFAA7F2E2FADC" box="[136,1451,1300,1322]" pageId="4" pageNumber="18">3. Stigma inserted obliquely ................................................................................................................................................................. 4</keyLead>
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<paragraph id="9D66FB49FFEE8465FF3FFA82F2E2FAB3" blockId="4.[136,1451,1300,1350]" box="[136,1451,1328,1350]" pageId="4" pageNumber="18">
<keyLead id="2628587CFFEE8465FF3FFA82F2E2FAB3" box="[136,1451,1328,1350]" pageId="4" pageNumber="18"> Stigma terminal ................................................................................................................................................................................ 5</keyLead>
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<paragraph id="9D66FB49FFEE8465FF3FFAD9F2E2FA69" blockId="4.[136,1452,1386,1494]" pageId="4" pageNumber="18">
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4. Fertile portion of infructescence not exceeding leaves, panicle dense, branches of the partial inflorescences ascending in fruit; plants sessile even in old age ....................................................................................................................................…
<taxonomicName id="5AD980CAFFEE8465FA9FFA34F2E2FA69" ID-CoL="3JJTG" box="[1320,1451,1415,1436]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="species" species="pantiensis">
<emphasis id="AFAD275BFFEE8465FA9FFA34F2E2FA69" bold="true" box="[1320,1451,1415,1436]" italics="true" pageId="4" pageNumber="18">H. pantiensis</emphasis>
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<keyLead id="2628587CFFEE8465FF3FFA17F2E2FA23" pageId="4" pageNumber="18">
Fertile portion of infructescence far exceeding leaves, panicle very open, branches of the partial inflorescences spreading and forming regular tiers; plants developing an erect, leafless stem up to
<quantity id="5A2156ACFFEE8465FCD2FA73F4D5FA23" box="[869,924,1472,1494]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="18" unit="m" value="1.5">1.5 m</quantity>
tall … ...................................................
<taxonomicName id="5AD980CAFFEE8465FAABFA72F2E2FA23" ID-CoL="8XQ8H" authorityName="Siti Nurfazilah, Mohd Fahmi, Sofiman Othman &amp; P.C.Boyce" authorityYear="2010" box="[1308,1451,1472,1494]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="species" species="podzolicola">
<emphasis id="AFAD275BFFEE8465FAABFA72F2E2FA23" bold="true" box="[1308,1451,1472,1494]" italics="true" pageId="4" pageNumber="18">H. podzolicola</emphasis>
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<keyLead id="2628587CFFEE8465FF3FFA49F2E2F9E5" box="[136,1451,1530,1552]" pageId="4" pageNumber="18">
5. Fruits ripening black ....................................................................................................................................................…
<taxonomicName id="5AD980CAFFEE8465FA8EFA49F2E2F9E5" ID-CoL="6LBV3" authorityName="Skornick. &amp; Niissalo" authorityYear="2014" box="[1337,1451,1530,1552]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="species" species="neglecta">
<emphasis id="AFAD275BFFEE8465FA8EFA49F2E2F9E5" bold="true" box="[1337,1451,1530,1552]" italics="true" pageId="4" pageNumber="18">H. neglecta</emphasis>
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</paragraph>
<paragraph id="9D66FB49FFEE8465FF3FF9A4F2E2F9D8" blockId="4.[136,1451,1530,1581]" box="[136,1451,1559,1581]" pageId="4" pageNumber="18">
<keyLead id="2628587CFFEE8465FF3FF9A4F2E2F9D8" box="[136,1451,1559,1581]" pageId="4" pageNumber="18"> Fruits ripening glossy white to pale yellow...................................................................................................................................… 6</keyLead>
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<paragraph id="9D66FB49FFEE8465FF3FF9E3F2E2F949" blockId="4.[136,1452,1616,1811]" pageId="4" pageNumber="18">
<keyLead id="2628587CFFEE8465FF3FF9E3F2E2F949" pageId="4" pageNumber="18">
6. Infructescence with caducous foliaceous bracts; partial inflorescences each with 2 or rarely 3 branches, spreading in fruit; fruits globose with a briefly stipitate stigma, lobes connate basally, deep chocolate brown; fruit ripening white with conspicuous black speckles; old plants developing a short (to ca.
<quantity id="5A2156ACFFEE8465FD32F939F589F955" box="[645,704,1674,1696]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="18" unit="cm" value="25.0">25 cm</quantity>
) slender, leafless stem. Plants of well-drained slopes and low ridges in lowland humid, moist mixed dipterocarp forest on yellow clay soils ....................................................................................…
<taxonomicName id="5AD980CAFFEE8465FA9FF914F2E2F949" ID-CoL="3JJTK" box="[1320,1451,1702,1724]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="species" species="stenopoda">
<emphasis id="AFAD275BFFEE8465FA9FF914F2E2F949" bold="true" box="[1320,1451,1702,1724]" italics="true" pageId="4" pageNumber="18">H. stenopoda</emphasis>
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<keyLead id="2628587CFFEE8465FF3FF977F2E2F8E6" pageId="4" pageNumber="18">
Infructescence with persistent foliaceous bracts; partial inflorescences each with 45 branches, these rather sharply ascending in fruit; fruits ventrally gibbose-ellipsoid, stigma sessile, comprising 3 free orange brown lobes; fruit ripening pale yellow without conspicuous black speckles. Plants of low-lying wet podzols in peatforest .................................................................
<taxonomicName id="5AD980CAFFEE8465FA8FF94DF2E2F8E6" ID-CoL="3JJT9" box="[1336,1451,1789,1811]" class="Liliopsida" family="Hanguanaceae" genus="Hanguana" kingdom="Plantae" order="Commelinales" pageId="4" pageNumber="18" phylum="Tracheophyta" rank="species" species="exultans">
<emphasis id="AFAD275BFFEE8465FA8FF94DF2E2F8E6" bold="true" box="[1336,1451,1789,1811]" italics="true" pageId="4" pageNumber="18">H. exultans</emphasis>
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