178 lines
22 KiB
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178 lines
22 KiB
XML
<document id="3DC4EF5CF729253D47D8D7B9939E0681" ID-CLB-Dataset="9216" ID-DOI="10.11646/zootaxa.4896.2.10" ID-GBIF-Dataset="9938488f-b51e-426d-8e49-1f5e12985e55" ID-ISSN="1175-5326" ID-Zenodo-Dep="4389263" ID-ZooBank="872E6481-0D12-469D-8581-3498F6F9D556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608541273307" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Mpamnaras, Athanasios & Kalaentzis, Konstantinos" docDate="2020" docId="03FCA61B9A755211FF52FD01FC778DC6" docLanguage="en" docName="zootaxa.4896.2.10.pdf" docOrigin="Zootaxa 4896 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stenohelops Reitter 1922" docType="treatment" docVersion="8" lastPageNumber="297" masterDocId="FFC5DE639A705214FFC5FF8FFF8E887A" masterDocTitle="Contribution to the knowledge of Mediterranean lichen-feeding darkling beetles of the genus Stenohelops Reitter, 1922 (Coleoptera: Tenebrionidae)" masterLastPageNumber="300" masterPageNumber="292" pageNumber="297" updateTime="1698847826758" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0B7B9A86F9FAE8D7F0E82D199C9A6F7A">Contribution to the knowledge of Mediterranean lichen-feeding darkling beetles of the genus Stenohelops Reitter, 1922 (Coleoptera: Tenebrionidae)</mods:title>
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<mods:name id="8AE2DD0D751BF5C932AEEE920C0D686B" type="personal">
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<mods:namePart id="E0178EAF981B70AD0B8DCC9B812CB744">Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation id="71DD5A61FB58DE0E7325695FA4D3D66C">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev Str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. & Dagestan State University, 43 a M. Gadzhiev Str., Makhachkala, Republic of Dagestan, 367000, Russia</mods:affiliation>
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<mods:namePart id="ED580128D2C831C432F3721BA13FBFEC">Mpamnaras, Athanasios</mods:namePart>
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<mods:nameIdentifier id="D58F50981877397FBC5AFBE24EBE8835" type="ORCID">0000-0003-4410-1041</mods:nameIdentifier>
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<mods:affiliation id="8F9AFFB3393E02CEF69C95E8B21E8F5C">Department of Agriculture & Agrotechnology, University of Thessaly, 41500 Larissa, Greece.</mods:affiliation>
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<mods:namePart id="24C017DDFC543BB12E5815A5685A899D">Kalaentzis, Konstantinos</mods:namePart>
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<mods:affiliation id="A99D84E3CA5EFEFBD4EF44A98FD44710">Institute of Biology, Leiden University, Sylvius Laboratory, Sylviusweg 72, 2333 BE, Leiden, The Netherlands. konstakal 95 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4986 - 796 X & Naturalis Biodiversity Center, PO Box 9517, 2300 RA, Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier id="79E5056B55C8D62821C420250368BE3E" type="email">konstakal95@gmail.com</mods:nameIdentifier>
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<mods:title id="43EDFBDDD8FF5B0FF67609F8526579C3">Zootaxa</mods:title>
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<mods:date id="DEDFBE452EF33C8537C3E2F5654A1BF7">2020</mods:date>
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<mods:number id="E6D76AC0181688A0B73B1DDDAB3F837D">2020-12-21</mods:number>
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<mods:identifier id="3205EDEB049887703E329F46045133D5" type="DOI">10.11646/zootaxa.4896.2.10</mods:identifier>
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<treatment id="03FCA61B9A755211FF52FD01FC778DC6" ID-DOI="http://doi.org/10.5281/zenodo.4389283" ID-GBIF-Taxon="176235248" ID-Zenodo-Dep="4389283" LSID="urn:lsid:plazi:treatment:03FCA61B9A755211FF52FD01FC778DC6" httpUri="http://treatment.plazi.org/id/03FCA61B9A755211FF52FD01FC778DC6" lastPageNumber="297" pageId="5" pageNumber="297">
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<subSubSection id="C34F44869A755211FF52FD01FDDE8AD2" box="[151,592,654,681]" pageId="5" pageNumber="297" type="nomenclature">
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<paragraph id="8BEA170D9A755211FF52FD01FDDE8AD2" blockId="5.[151,592,654,681]" box="[151,592,654,681]" pageId="5" pageNumber="297">
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<heading id="D0A2A0619A755211FF52FD01FDDE8AD2" bold="true" box="[151,592,654,681]" fontSize="11" level="1" pageId="5" pageNumber="297" reason="1">
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<emphasis id="B921CB1F9A755211FF52FD01FDDE8AD2" bold="true" box="[151,592,654,681]" pageId="5" pageNumber="297">
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Subgenus
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<taxonomicName id="4C556C8E9A755211FED2FD01FDDE8AD2" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" box="[279,592,654,681]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
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<emphasis id="B921CB1F9A755211FED2FD01FE248AD3" bold="true" box="[279,426,654,681]" italics="true" pageId="5" pageNumber="297">Stenohelops</emphasis>
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<bibRefCitation id="EFC46AFC9A755211FE74FD01FDDE8AD2" author="Reitter, E." box="[433,592,654,680]" pageId="5" pageNumber="297" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F44869A755211FF52FD5BFE778A91" box="[151,505,723,747]" pageId="5" pageNumber="297" type="reference_group">
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<paragraph id="8BEA170D9A755211FF52FD5BFE778A91" blockId="5.[151,505,723,747]" box="[151,505,723,747]" pageId="5" pageNumber="297">
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=
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<taxonomicName id="4C556C8E9A755211FF69FD5CFE278A91" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" baseAuthorityName="Nabozhenko" baseAuthorityYear="2008" box="[172,425,723,747]" class="Insecta" family="Tenebrionidae" genus="Gunarellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
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<emphasis id="B921CB1F9A755211FF69FD5CFE938A91" box="[172,285,723,747]" italics="true" pageId="5" pageNumber="297">Gunarellus</emphasis>
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<bibRefCitation id="EFC46AFC9A755211FEE6FD5CFE278A91" author="Reitter, E." box="[291,425,723,747]" pageId="5" pageNumber="297" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="B921CB1F9A755211FE76FD5CFE778A91" bold="true" box="[435,505,723,747]" pageId="5" pageNumber="297">syn. n.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F44869A755211FF52FC96FC778DC6" pageId="5" pageNumber="297" type="description">
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<paragraph id="8BEA170D9A755211FF52FC96FE378C29" blockId="5.[151,1437,793,1468]" pageId="5" pageNumber="297">
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<emphasis id="B921CB1F9A755211FF52FC96FE418B49" bold="true" box="[151,463,793,819]" pageId="5" pageNumber="297">Taxonomy and synonymy.</emphasis>
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Recently
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<bibRefCitation id="EFC46AFC9A755211FD87FC95FD4E8B49" author="Leo, P." box="[578,704,793,820]" pageId="5" pageNumber="297" pagination="1 - 11" refId="ref5421" refString="Leo, P. (2020) Descrizione di una nuova specie di Stenohelops della Sardegna meridionale con nota sinonimica su Stenohelops berettai Marcuzzi, 2001 (Coleoptera, Tenebrionidae, Helopini). Doriana, 9 (409), 1 - 11." type="journal article" year="2020">Leo (2020)</bibRefCitation>
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discussed about the subgeneric system of
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<taxonomicName id="4C556C8E9A755211FB5AFC96FAA88B49" authorityName="Reitter" authorityYear="1922" box="[1183,1318,793,819]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
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<emphasis id="B921CB1F9A755211FB5AFC96FAA88B49" box="[1183,1318,793,819]" italics="true" pageId="5" pageNumber="297">Stenohelops</emphasis>
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</taxonomicName>
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and noted that the nominotypical subgenus and
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<taxonomicName id="4C556C8E9A755211FDFAFCB2FD328B2D" baseAuthorityName="Nabozhenko" baseAuthorityYear="2008" box="[575,700,829,855]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="subGenus" subGenus="Gunarellus">
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<emphasis id="B921CB1F9A755211FDFAFCB2FD328B2D" box="[575,700,829,855]" italics="true" pageId="5" pageNumber="297">Gunarellus</emphasis>
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</taxonomicName>
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are difficult to distinguish.
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<bibRefCitation id="EFC46AFC9A755211FC3CFCB2FB288B22" author="Espanol, F." box="[1017,1190,829,856]" pageId="5" pageNumber="297" pagination="19 - 38" refId="ref5166" refString="Espanol, F. (1957) Contribucion al estudio de los Stenohelops Ibericos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 33, 19 - 38." type="journal article" year="1957">Español (1957)</bibRefCitation>
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also mentioned these difficulties. Both subgenera are different only by the presence or the absence of the fine rim along the apex of the elytral suture. This unclear character was used by
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<bibRefCitation id="EFC46AFC9A755211FD19FC0AFCF28BE5" author="Reitter, E." box="[732,892,901,927]" pageId="5" pageNumber="297" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter (1922)</bibRefCitation>
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both for species and for subgenera in different groups of
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<taxonomicName id="4C556C8E9A755211FECFFC26FEE18BB9" authorityName="Latreille" authorityYear="1802" box="[266,367,937,963]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="tribe" tribe="Helopini">Helopini</taxonomicName>
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, but it can be variable even in different population of one species. For example,
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<bibRefCitation id="EFC46AFC9A755211FB3BFC26FA128BB9" author="Reitter, E." box="[1278,1436,937,963]" pageId="5" pageNumber="297" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter (1922)</bibRefCitation>
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indicated the apically un-rimmed elytral suture in
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<taxonomicName id="4C556C8E9A755211FD0AFC42FAB08B9D" authority="(J. Frivaldszky, 1894)" baseAuthorityName="J. Frivaldszky" baseAuthorityYear="1894" box="[719,1342,973,1000]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="gratus" subGenus="Gunarellus">
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<emphasis id="B921CB1F9A755211FD0AFC42FCD88B9D" box="[719,854,973,999]" italics="true" pageId="5" pageNumber="297">Stenohelops</emphasis>
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(
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<emphasis id="B921CB1F9A755211FCA2FC42FC6A8B9D" box="[871,996,973,999]" italics="true" pageId="5" pageNumber="297">Gunarellus</emphasis>
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)
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<emphasis id="B921CB1F9A755211FC30FC41FBB38B9D" box="[1013,1085,974,999]" italics="true" pageId="5" pageNumber="297">gratus</emphasis>
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(J. Frivaldszky, 1894)
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</taxonomicName>
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. Kaszab (1937) also mentioned this character, but later (
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<bibRefCitation id="EFC46AFC9A755211FD74FC7EFCC98C76" author="Kaszab, Z." box="[689,839,1009,1036]" pageId="5" pageNumber="297" refId="ref5308" refString="Kaszab, Z. (1957) Magyarorszag Allatvilaga (Fauna Hungariae). IX. Kotet, Coleoptera IV. Felemas labfejizes bogarak I. Heteromera I. Akademiai Kiado, Budapest, 126 pp." type="book" year="1957">Kaszab 1957</bibRefCitation>
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) he wrote, that elytral suture apically is rimmed and transferred this species to the nominotypical subgenus. In fact, this species has left sutural margin rimmed and right one un-rimmed (
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<figureCitation id="136E0B889A755211FE97FBB6FE228C29" box="[338,428,1081,1107]" captionStart="FIGURE 3" captionStartId="6.[151,250,1531,1555]" captionTargetBox="[159,1436,278,1500]" captionTargetId="figure-56@6.[151,1436,273,1508]" captionTargetPageId="6" captionText="FIGURE 3. Stenohelops gratus (J. Frivaldszky, 1894), lectotype, male (© photographs, Hungarian Natural History Museum): A—habitus, B—aedeagus, lateral view, C—labels." figureDoi="http://doi.org/10.5281/zenodo.4389273" httpUri="https://zenodo.org/record/4389273/files/figure.png" pageId="5" pageNumber="297">Fig. 3A</figureCitation>
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).
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</paragraph>
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<paragraph id="8BEA170D9A755211FF02FBD2FB638CC5" blockId="5.[151,1437,793,1468]" pageId="5" pageNumber="297">
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The other question is that
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<taxonomicName id="4C556C8E9A755211FE3DFBD2FD5F8C0D" baseAuthorityName="J. Frivaldszky" baseAuthorityYear="1894" box="[504,721,1117,1143]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="gratus">
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<emphasis id="B921CB1F9A755211FE3DFBD2FD5F8C0D" box="[504,721,1117,1143]" italics="true" pageId="5" pageNumber="297">Stenohelops gratus</emphasis>
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</taxonomicName>
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, which was described from
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<collectingCountry id="F342579D9A755211FBDFFBD2FBEC8C0D" box="[1050,1122,1117,1143]" name="Serbia" pageId="5" pageNumber="297">Serbia</collectingCountry>
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and designated as the
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<typeStatus id="54EEA9AF9A755211FAA9FBD1FA128C02" box="[1388,1436,1118,1144]" pageId="5" pageNumber="297">type</typeStatus>
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species of the subgenus
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<taxonomicName id="4C556C8E9A755211FE6FFB0EFC818CE1" authority="(Nabozhenko 2008)" baseAuthorityName="Nabozhenko" baseAuthorityYear="2008" box="[426,783,1153,1179]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="subGenus" subGenus="Gunarellus">
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<emphasis id="B921CB1F9A755211FE6FFB0EFDA98CE1" box="[426,551,1153,1179]" italics="true" pageId="5" pageNumber="297">Gunarellus</emphasis>
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(
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<bibRefCitation id="EFC46AFC9A755211FDF3FB0EFC868CE1" author="Nabozhenko, M. V." box="[566,776,1153,1179]" pageId="5" pageNumber="297" refId="ref5466" refString="Nabozhenko, M. V. (2008) Tenebrionidae: Helopini. New nomenclatural and taxonomic acts, and comments. In: Lobl, I. & Smetana, A (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Tenebrionoidea. Apollo books, Stenstrup, pp. 36 - 38." type="book" year="2008">Nabozhenko 2008</bibRefCitation>
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)
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</taxonomicName>
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is mentioned as doubtful and little-known species, which occurs far from the main range of this Western Mediterranean subgenus (
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<bibRefCitation id="EFC46AFC9A755211FC0DFB2AFBE88CBA" author="Espanol, F." box="[968,1126,1189,1216]" pageId="5" pageNumber="297" pagination="19 - 38" refId="ref5166" refString="Espanol, F. (1957) Contribucion al estudio de los Stenohelops Ibericos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 33, 19 - 38." type="journal article" year="1957">Español 1957</bibRefCitation>
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,
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<bibRefCitation id="EFC46AFC9A755211FBB4FB29FB6F8CC5" author="Leo, P." box="[1137,1249,1189,1216]" pageId="5" pageNumber="297" pagination="1 - 11" refId="ref5421" refString="Leo, P. (2020) Descrizione di una nuova specie di Stenohelops della Sardegna meridionale con nota sinonimica su Stenohelops berettai Marcuzzi, 2001 (Coleoptera, Tenebrionidae, Helopini). Doriana, 9 (409), 1 - 11." type="journal article" year="2020">Leo 2020</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BEA170D9A755211FF02FB45FD868D09" blockId="5.[151,1437,793,1468]" pageId="5" pageNumber="297">
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We studied one
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<typeStatus id="54EEA9AF9A755211FE44FB45FE568C9E" box="[385,472,1226,1252]" pageId="5" pageNumber="297" type="syntype">syntype</typeStatus>
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(male) of
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<taxonomicName id="4C556C8E9A755211FD91FB46FCA38C99" baseAuthorityName="J. Frivaldszky" baseAuthorityYear="1894" box="[596,813,1225,1251]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="gratus">
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<emphasis id="B921CB1F9A755211FD91FB46FCA38C99" box="[596,813,1225,1251]" italics="true" pageId="5" pageNumber="297">Stenohelops gratus</emphasis>
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</taxonomicName>
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and confirm that this species distinctly belongs to the genus
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<taxonomicName id="4C556C8E9A755211FF24FB62FEE68D7D" authorityName="Reitter" authorityYear="1922" box="[225,360,1261,1287]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
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|
<emphasis id="B921CB1F9A755211FF24FB62FEE68D7D" box="[225,360,1261,1287]" italics="true" pageId="5" pageNumber="297">Stenohelops</emphasis>
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</taxonomicName>
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and has elongated, but distinctly separated deep punctures on prothoracic hypomera as in subgenera
|
|
<taxonomicName id="4C556C8E9A755211FF2FFA9EFEFF8D51" authorityName="Reitter" authorityYear="1922" box="[234,369,1297,1323]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
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|
<emphasis id="B921CB1F9A755211FF2FFA9EFEFF8D51" box="[234,369,1297,1323]" italics="true" pageId="5" pageNumber="297">Stenohelops</emphasis>
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|
</taxonomicName>
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|
and
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|
<taxonomicName id="4C556C8E9A755211FE6CFA9EFDA88D51" baseAuthorityName="Nabozhenko" baseAuthorityYear="2008" box="[425,550,1297,1323]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="subGenus" subGenus="Gunarellus">
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|
<emphasis id="B921CB1F9A755211FE6CFA9EFDA88D51" box="[425,550,1297,1323]" italics="true" pageId="5" pageNumber="297">Gunarellus</emphasis>
|
|
</taxonomicName>
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|
. The character of rimmed elytral suture at apex cannot be used for subgeneric division as indistinct and variable. As a result the following synonymy is proposed:
|
|
<taxonomicName id="4C556C8E9A755211FB9AFABAFA0D8D35" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" box="[1119,1411,1333,1359]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B921CB1F9A755211FB9AFABAFB688D35" box="[1119,1254,1333,1359]" italics="true" pageId="5" pageNumber="297">Stenohelops</emphasis>
|
|
<bibRefCitation id="EFC46AFC9A755211FB35FABAFA0D8D35" author="Reitter, E." box="[1264,1411,1333,1359]" pageId="5" pageNumber="297" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
|
|
</taxonomicName>
|
|
=
|
|
<taxonomicName id="4C556C8E9A755211FF52FAD6FE218D09" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" baseAuthorityName="Nabozhenko" baseAuthorityYear="2008" box="[151,431,1369,1395]" class="Insecta" family="Tenebrionidae" genus="Gunarellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B921CB1F9A755211FF52FAD6FE9A8D09" box="[151,276,1369,1395]" italics="true" pageId="5" pageNumber="297">Gunarellus</emphasis>
|
|
<bibRefCitation id="EFC46AFC9A755211FEDEFAD6FE218D09" author="Reitter, E." box="[283,431,1369,1395]" pageId="5" pageNumber="297" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis id="B921CB1F9A755211FE7FFAD6FD868D09" bold="true" box="[442,520,1369,1395]" pageId="5" pageNumber="297">syn. n.</emphasis>
|
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</paragraph>
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<paragraph id="8BEA170D9A755211FF02FAF1FC778DC6" blockId="5.[151,1437,793,1468]" pageId="5" pageNumber="297">In conclusion, we suggested that the sculpture of prothoracic hypomera is also formal character, and the future molecular phylogeny may show completely different branches for this genus.</paragraph>
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</subSubSection>
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</treatment>
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</document> |