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<document id="8B3D1F3F31D57A20220BD7CD77A2011E" ID-CLB-Dataset="9216" ID-DOI="10.11646/zootaxa.4896.2.10" ID-GBIF-Dataset="9938488f-b51e-426d-8e49-1f5e12985e55" ID-ISSN="1175-5326" ID-Zenodo-Dep="4389263" ID-ZooBank="872E6481-0D12-469D-8581-3498F6F9D556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608541273307" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Mpamnaras, Athanasios &amp; Kalaentzis, Konstantinos" docDate="2020" docId="03FCA61B9A715217FF52F94EFC7A8D03" docLanguage="en" docName="zootaxa.4896.2.10.pdf" docOrigin="Zootaxa 4896 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helopelius Reitter 1922" docType="treatment" docVersion="8" lastPageNumber="295" masterDocId="FFC5DE639A705214FFC5FF8FFF8E887A" masterDocTitle="Contribution to the knowledge of Mediterranean lichen-feeding darkling beetles of the genus Stenohelops Reitter, 1922 (Coleoptera: Tenebrionidae)" masterLastPageNumber="300" masterPageNumber="292" pageNumber="293" updateTime="1698847826758" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8CC95D0ECB7AEA4731F3CDC9B90DA531">Contribution to the knowledge of Mediterranean lichen-feeding darkling beetles of the genus Stenohelops Reitter, 1922 (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="D3200A12D91A7BB2C81FA252CBED52BB">Nabozhenko, Maxim V.</mods:namePart>
<mods:affiliation id="1C4116920644794174830F52EA59B18C">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev Str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. &amp; Dagestan State University, 43 a M. Gadzhiev Str., Makhachkala, Republic of Dagestan, 367000, Russia</mods:affiliation>
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<mods:namePart id="BBFD01889BC99A67CB1AB88871AEF483">Mpamnaras, Athanasios</mods:namePart>
<mods:nameIdentifier id="5AE355921AB5C8F5CF863D788DDDDD30" type="ORCID">0000-0003-4410-1041</mods:nameIdentifier>
<mods:affiliation id="2D4C0A7C9F480453E184301356F22A3D">Department of Agriculture &amp; Agrotechnology, University of Thessaly, 41500 Larissa, Greece.</mods:affiliation>
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<mods:namePart id="C02471D322980AC0BAADC3A7887A37A0">Kalaentzis, Konstantinos</mods:namePart>
<mods:nameIdentifier id="02B865623FC23A9937C69FBE0E4F0197" type="ORCID">0000-0003-4986-796X</mods:nameIdentifier>
<mods:affiliation id="EC4033BD06C7CB163D67A7C16B7DE3CA">Institute of Biology, Leiden University, Sylvius Laboratory, Sylviusweg 72, 2333 BE, Leiden, The Netherlands. konstakal 95 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4986 - 796 X &amp; Naturalis Biodiversity Center, PO Box 9517, 2300 RA, Leiden, The Netherlands</mods:affiliation>
<mods:nameIdentifier id="1E0006165B6BA17813A13766CBCA80A3" type="email">konstakal95@gmail.com</mods:nameIdentifier>
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<mods:title id="173BAE1149AF343A208CDC0583F6E832">Zootaxa</mods:title>
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<mods:date id="8D278856F381CA6666543BCE4256790D">2020</mods:date>
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<treatment id="03FCA61B9A715217FF52F94EFC7A8D03" ID-DOI="http://doi.org/10.5281/zenodo.4389271" ID-GBIF-Taxon="176235249" ID-Zenodo-Dep="4389271" LSID="urn:lsid:plazi:treatment:03FCA61B9A715217FF52F94EFC7A8D03" httpUri="http://treatment.plazi.org/id/03FCA61B9A715217FF52F94EFC7A8D03" lastPageId="3" lastPageNumber="295" pageId="1" pageNumber="293">
<subSubSection id="C34F44869A715215FF52F94EFDCE8EA1" box="[151,576,1729,1756]" pageId="1" pageNumber="293" type="nomenclature">
<paragraph id="8BEA170D9A715215FF52F94EFDCE8EA1" blockId="1.[151,576,1729,1756]" box="[151,576,1729,1756]" pageId="1" pageNumber="293">
<heading id="D0A2A0619A715215FF52F94EFDCE8EA1" bold="true" box="[151,576,1729,1756]" fontSize="11" level="1" pageId="1" pageNumber="293" reason="1">
<emphasis id="B921CB1F9A715215FF52F94EFDCE8EA1" bold="true" box="[151,576,1729,1756]" pageId="1" pageNumber="293">
Subgenus
<taxonomicName id="4C556C8E9A715215FED2F94EFDCE8EA1" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" box="[279,576,1729,1756]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A715215FED2F94EFE178EA6" bold="true" box="[279,409,1729,1756]" italics="true" pageId="1" pageNumber="293">Helopelius</emphasis>
<bibRefCitation id="EFC46AFC9A715215FE64F94EFDCE8EA1" author="Reitter, E." box="[417,576,1729,1755]" pageId="1" pageNumber="293" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C34F44869A715215FF52F889FBF88F64" box="[151,1142,1798,1822]" pageId="1" pageNumber="293" type="materials_examined">
<paragraph id="8BEA170D9A715215FF52F889FBF88F64" blockId="1.[151,1325,1798,1853]" box="[151,1142,1798,1822]" pageId="1" pageNumber="293">
<typeStatus id="54EEA9AF9A715215FF52F889FF478F64" box="[151,201,1798,1822]" pageId="1" pageNumber="293">Type</typeStatus>
species
<taxonomicName id="4C556C8E9A715215FEDAF889FD068F64" authority="Allard, 1876" authorityName="Allard" authorityYear="1876" box="[287,648,1798,1822]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B921CB1F9A715215FEDAF889FD8F8F67" box="[287,513,1798,1821]" italics="true" pageId="1" pageNumber="293">Stenomax aeneipennis</emphasis>
<bibRefCitation id="EFC46AFC9A715215FDC3F889FD068F64" author="Allard, E." box="[518,648,1798,1822]" pageId="1" pageNumber="293" pagination="1 - 80" refId="ref5031" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard, 1876</bibRefCitation>
</taxonomicName>
, by subsequent designation (
<bibRefCitation id="EFC46AFC9A715215FC6EF889FBE48F64" author="Nabozhenko, M. V." box="[939,1130,1798,1822]" pageId="1" pageNumber="293" refId="ref5466" refString="Nabozhenko, M. V. (2008) Tenebrionidae: Helopini. New nomenclatural and taxonomic acts, and comments. In: Lobl, I. &amp; Smetana, A (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Tenebrionoidea. Apollo books, Stenstrup, pp. 36 - 38." type="book" year="2008">Nabozhenko 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F44869A715216FF52F8AAFDE5893B" lastPageId="2" lastPageNumber="294" pageId="1" pageNumber="293" type="reference_group">
<paragraph id="8BEA170D9A715215FF52F8AAFAA38F47" blockId="1.[151,1325,1798,1853]" box="[151,1325,1828,1853]" pageId="1" pageNumber="293">
=
<taxonomicName id="4C556C8E9A715215FF69F8ABFE308F46" authority="Espanol, 1957" authorityName="Espanol" authorityYear="1957" box="[172,446,1828,1852]" class="Insecta" family="Tenebrionidae" genus="Stenomaleis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A715215FF69F8ABFEA88F46" box="[172,294,1828,1852]" italics="true" pageId="1" pageNumber="293">Stenomaleis</emphasis>
<bibRefCitation id="EFC46AFC9A715215FEE8F8ABFE308F46" author="Espanol, F." box="[301,446,1828,1852]" pageId="1" pageNumber="293" pagination="19 - 38" refId="ref5166" refString="Espanol, F. (1957) Contribucion al estudio de los Stenohelops Ibericos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 33, 19 - 38." type="journal article" year="1957">Español, 1957</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54EEA9AF9A715215FE0EF8AAFE768F47" box="[459,504,1829,1853]" pageId="1" pageNumber="293">type</typeStatus>
species:
<taxonomicName id="4C556C8E9A715215FD90F8ABFC378F46" authority="Espanol, 1957" authorityName="Espanol" authorityYear="1957" box="[597,953,1828,1852]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B921CB1F9A715215FD90F8ABFCAE8F46" box="[597,800,1828,1852]" italics="true" pageId="1" pageNumber="293">Stenohelops ardoini</emphasis>
<bibRefCitation id="EFC46AFC9A715215FCE3F8ABFC378F46" author="Espanol, F." box="[806,953,1828,1852]" pageId="1" pageNumber="293" pagination="19 - 38" refId="ref5166" refString="Espanol, F. (1957) Contribucion al estudio de los Stenohelops Ibericos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 33, 19 - 38." type="journal article" year="1957">Español, 1957</bibRefCitation>
</taxonomicName>
, by the original designation),
<emphasis id="B921CB1F9A715215FB22F8AAFAA38F47" bold="true" box="[1255,1325,1829,1853]" pageId="1" pageNumber="293">syn. n.</emphasis>
</paragraph>
<paragraph id="8BEA170D9A715216FF52F8E4FDE5893B" blockId="1.[151,1437,1898,2033]" lastBlockId="2.[151,1437,151,321]" lastPageId="2" lastPageNumber="294" pageId="1" pageNumber="293">
<emphasis id="B921CB1F9A715215FF52F8E4FD0F8FFF" bold="true" box="[151,641,1899,1925]" pageId="1" pageNumber="293">Taxonomy, nomenclature and synonymy.</emphasis>
<bibRefCitation id="EFC46AFC9A715215FD4CF8E4FCB88FFF" author="Espanol, F." box="[649,822,1898,1925]" pageId="1" pageNumber="293" pagination="19 - 38" refId="ref5166" refString="Espanol, F. (1957) Contribucion al estudio de los Stenohelops Ibericos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 33, 19 - 38." type="journal article" year="1957">Español (1957)</bibRefCitation>
erected the subgenus
<taxonomicName id="4C556C8E9A715215FBF5F8E4FB398FFF" authority="Español, 1957" box="[1072,1207,1899,1925]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="subGenus" subGenus="Stenomaleis">
<emphasis id="B921CB1F9A715215FBF5F8E4FB398FFF" box="[1072,1207,1899,1925]" italics="true" pageId="1" pageNumber="293">Stenomaleis</emphasis>
</taxonomicName>
for three species,
<taxonomicName id="4C556C8E9A715215FA42F8E4FE408FD3" authority="(Seidlitz, 1895)" baseAuthorityName="Seidlitz" baseAuthorityYear="1895" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="protensulus">
<emphasis id="B921CB1F9A715215FA42F8E4FE978FD3" italics="true" pageId="1" pageNumber="293">S. protensulus</emphasis>
(Seidlitz, 1895)
</taxonomicName>
,
<taxonomicName id="4C556C8E9A715215FE1CF800FD6D8FD3" authority="Espanol 1957" authorityName="Espanol" authorityYear="1957" box="[473,739,1934,1961]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B921CB1F9A715215FE1CF800FDCB8FD3" box="[473,581,1935,1961]" italics="true" pageId="1" pageNumber="293">S. ardoini</emphasis>
<bibRefCitation id="EFC46AFC9A715215FD8EF800FD6D8FD3" author="Espanol, F." box="[587,739,1934,1961]" pageId="1" pageNumber="293" pagination="19 - 38" refId="ref5166" refString="Espanol, F. (1957) Contribucion al estudio de los Stenohelops Ibericos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 33, 19 - 38." type="journal article" year="1957">Español 1957</bibRefCitation>
</taxonomicName>
(now a junior synonym of
<taxonomicName id="4C556C8E9A715215FBC8F800FB248FD3" baseAuthorityName="Seidlitz" baseAuthorityYear="1895" box="[1037,1194,1935,1961]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="protensulus">
<emphasis id="B921CB1F9A715215FBC8F800FB248FD3" box="[1037,1194,1935,1961]" italics="true" pageId="1" pageNumber="293">S. protensulus</emphasis>
</taxonomicName>
according to
<bibRefCitation id="EFC46AFC9A715215FAFAF800FED88FB7" author="Lagar, A. &amp; Fresneda, J." pageId="1" pageNumber="293" pagination="93 - 99" refId="ref5345" refString="Lagar, A. &amp; Fresneda, J. (2014) Stenohelops barcinensis sp. n. de Barcelona (Espana) (Coleoptera, Tenebrionidae, Helopini). Boletin de la Sociedad Entomologica Aragonesa, 55, 93 - 99." type="journal article" year="2014">Lagar &amp; Fresneda (2014))</bibRefCitation>
and
<taxonomicName id="4C556C8E9A715215FE4FF83CFD5E8FB7" authority="(Mulsant, 1854)" baseAuthorityName="Mulsant" baseAuthorityYear="1854" box="[394,720,1970,1997]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="293" phylum="Arthropoda" rank="species" species="pyrenaeus">
<emphasis id="B921CB1F9A715215FE4FF83CFD998FB6" box="[394,535,1971,1997]" italics="true" pageId="1" pageNumber="293">S. pyrenaeus</emphasis>
(Mulsant, 1854)
</taxonomicName>
(
<typeStatus id="54EEA9AF9A715215FD18F83CFC818FB7" box="[733,783,1971,1997]" pageId="1" pageNumber="293">type</typeStatus>
species of the subgenus) on the basis of rugose prothoracic hypomera and glabrous dorsal side of the body, in contrast with other two subgenera with punctured prohypomera, glabrous (s. str.) or pubescent (
<taxonomicName id="4C556C8E9A725216FE37FF18FDE188CB" baseAuthorityName="Nabozhenko" baseAuthorityYear="2008" box="[498,623,151,177]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="subGenus" subGenus="Gunarellus">
<emphasis id="B921CB1F9A725216FE37FF18FDE188CB" box="[498,623,151,177]" italics="true" pageId="2" pageNumber="294">Gunarellus</emphasis>
</taxonomicName>
) body. Later, he described
<taxonomicName id="4C556C8E9A725216FC65FF18FBB388CB" authorityName="Espanol" authorityYear="1963" box="[928,1085,151,177]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="effeminatus">
<emphasis id="B921CB1F9A725216FC65FF18FBB388CB" box="[928,1085,151,177]" italics="true" pageId="2" pageNumber="294">S. effeminatus</emphasis>
</taxonomicName>
within this subgenus (
<bibRefCitation id="EFC46AFC9A725216FAFCFF18FF5D88AC" author="Espanol, F." pageId="2" pageNumber="294" pagination="389 - 396" refId="ref5199" refString="Espanol, F. (1963) Mas datos sobre los Helopinae de Marruecos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 39, 389 - 396." type="journal article" year="1963">Español, 1963</bibRefCitation>
).
<bibRefCitation id="EFC46AFC9A725216FF22FF34FDB488AF" author="Nabozhenko, M. V. &amp; Keskin, B." box="[231,570,187,213]" pageId="2" pageNumber="294" pagination="53 - 60" refId="ref5749" refString="Nabozhenko, M. V. &amp; Keskin, B. (2009) Two new species of the genus Gunarus Des Gozis, 1886 (Coleoptera: Tenebrionidae: Helopini) from Southern Turkey. Zootaxa, 2170 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 2170.1.6" type="journal article" year="2009">Nabozhenko &amp; Keskin (2009)</bibRefCitation>
described two Anatolian species within the genus
<taxonomicName id="4C556C8E9A725216FBB7FF34FA1688AF" authority="Des Gozis, 1886" authorityName="Des Gozis" authorityYear="1886" box="[1138,1432,187,213]" class="Insecta" family="Tenebrionidae" genus="Gunarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A725216FBB7FF34FB5A88AF" box="[1138,1236,187,213]" italics="true" pageId="2" pageNumber="294">Gunarus</emphasis>
Des Gozis, 1886
</taxonomicName>
, but later transferred these species to the genus
<taxonomicName id="4C556C8E9A725216FD76FF50FCB48883" authorityName="Reitter" authorityYear="1922" box="[691,826,223,249]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A725216FD76FF50FCB48883" box="[691,826,223,249]" italics="true" pageId="2" pageNumber="294">Stenohelops</emphasis>
</taxonomicName>
within the subgenus
<taxonomicName id="4C556C8E9A725216FBF4FF50FF648964" authority="(Nabozhenko et al. 2016 b)" baseAuthorityName="Nabozhenko" baseAuthorityYear="2016" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="subGenus" subGenus="Stenomaleis">
<emphasis id="B921CB1F9A725216FBF4FF50FB368883" box="[1073,1208,223,249]" italics="true" pageId="2" pageNumber="294">Stenomaleis</emphasis>
(
<bibRefCitation id="EFC46AFC9A725216FB0CFF50FF6C8964" author="Nabozhenko, M. V. &amp; Keskin, B. &amp; Dvadnenko, K. D. &amp; Alpagut Keskin N." pageId="2" pageNumber="294" pagination="111 - 116" refId="ref6006" refString="Nabozhenko, M. V., Keskin, B., Dvadnenko, K. D. &amp; Alpagut Keskin N. (2016 b) The genus Gunarus Des Gozis, 1886 belongs to the subtribe Cylindrinotina (Coleoptera: Tenebrionidae: Helopini). Caucasian Entomological Bulletin, 12, 111 - 116. https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 1 - 111 - 116" type="journal article" year="2016">
Nabozhenko
<emphasis id="B921CB1F9A725216FAA7FF50FA138883" box="[1378,1437,223,249]" italics="true" pageId="2" pageNumber="294">et al.</emphasis>
2016b
</bibRefCitation>
)
</taxonomicName>
. Two species of this subgenus were recently described from
<collectingCountry id="F342579D9A725216FC5BFE8CFC508967" box="[926,990,259,285]" name="Spain" pageId="2" pageNumber="294">Spain</collectingCountry>
(
<bibRefCitation id="EFC46AFC9A725216FC2BFE8CFA978967" author="Lagar, A. &amp; Fresneda, J." box="[1006,1305,259,285]" pageId="2" pageNumber="294" pagination="93 - 99" refId="ref5345" refString="Lagar, A. &amp; Fresneda, J. (2014) Stenohelops barcinensis sp. n. de Barcelona (Espana) (Coleoptera, Tenebrionidae, Helopini). Boletin de la Sociedad Entomologica Aragonesa, 55, 93 - 99." type="journal article" year="2014">Lagar and Fresneda, 2014</bibRefCitation>
) and
<collectingCountry id="F342579D9A725216FA9CFE8CFA128967" box="[1369,1436,259,285]" name="China" pageId="2" pageNumber="294">China</collectingCountry>
(
<bibRefCitation id="EFC46AFC9A725216FF5AFEA8FE41893B" author="Nabozhenko, M. V. &amp; Ando, K." box="[159,463,295,321]" pageId="2" pageNumber="294" pagination="277 - 327" refId="ref5688" refString="Nabozhenko, M. V. &amp; Ando, K. (2018) Subtribal, generic and subgeneric composition of darkling beetles of the tribe Helopini (Coleoptera: Tenebrionidae) in the Eastern Palaearctic. Acta zoologica Academiae Scientiarum Hungaricae, 64, 277 - 327. https: // doi. org / 10.17109 / AZH. 64.4.277.2018" type="journal article" year="2018">Nabozhenko &amp; Ando 2018</bibRefCitation>
) respectively.
</paragraph>
</subSubSection>
<caption id="DF2A47859A725216FF52F93AFEF38F27" ID-DOI="http://doi.org/10.5281/zenodo.4389267" ID-Zenodo-Dep="4389267" httpUri="https://zenodo.org/record/4389267/files/figure.png" pageId="2" pageNumber="294" startId="2.[151,250,1717,1741]" targetBox="[151,1432,348,1686]" targetPageId="2">
<paragraph id="8BEA170D9A725216FF52F93AFEF38F27" blockId="2.[151,1437,1717,1886]" pageId="2" pageNumber="294">
<emphasis id="B921CB1F9A725216FF52F93AFE988EB4" bold="true" box="[151,278,1717,1742]" pageId="2" pageNumber="294">FIGURE 1.</emphasis>
<taxonomicName id="4C556C8E9A725216FEDBF939FCCE8EB4" authority="(Allard, 1876)" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Allard" baseAuthorityYear="1876" box="[286,832,1718,1742]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="aeneipennis" subGenus="Helopelius">
<emphasis id="B921CB1F9A725216FEDBF939FE168EB4" box="[286,408,1718,1742]" italics="true" pageId="2" pageNumber="294">Stenohelops</emphasis>
(
<emphasis id="B921CB1F9A725216FE6FF939FD968EB4" box="[426,536,1718,1742]" italics="true" pageId="2" pageNumber="294">Helopelius</emphasis>
)
<emphasis id="B921CB1F9A725216FDECF939FD2C8EB7" box="[553,674,1718,1741]" italics="true" pageId="2" pageNumber="294">aeneipennis</emphasis>
(
<bibRefCitation id="EFC46AFC9A725216FD77F939FCB78EB4" author="Allard, E." box="[690,825,1718,1742]" pageId="2" pageNumber="294" pagination="1 - 80" refId="ref5031" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard, 1876</bibRefCitation>
)
</taxonomicName>
, habitus (A–D—© photographs, Natural History Museum London; E–H—© photographs, Naturhistorische Museum, Basel): A—lectotype of
<taxonomicName id="4C556C8E9A725216FC1AF955FB4E8E8B" authorityName="Allard" authorityYear="1876" box="[991,1216,1754,1777]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B921CB1F9A725216FC1AF955FB4E8E8B" box="[991,1216,1754,1777]" italics="true" pageId="2" pageNumber="294">Stenomax aeneipennis</emphasis>
</taxonomicName>
, female; B—labels of the lectotype of
<taxonomicName id="4C556C8E9A725216FEFEF971FE438F6F" authorityName="Allard" authorityYear="1876" box="[315,461,1790,1813]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B921CB1F9A725216FEFEF971FE438F6F" box="[315,461,1790,1813]" italics="true" pageId="2" pageNumber="294">S. aeneipennis</emphasis>
</taxonomicName>
; C—labels of non-type male from Rhodes (NHML); D—non-type male from Rhodes (NHML); E—lectotype of
<taxonomicName id="4C556C8E9A725216FE85F8ADFDBB8F40" authorityName="Reitter" authorityYear="1922" box="[320,565,1826,1850]" class="Insecta" family="Tenebrionidae" genus="Cylindrinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="disgregus">
<emphasis id="B921CB1F9A725216FE85F8ADFDBB8F40" box="[320,565,1826,1850]" italics="true" pageId="2" pageNumber="294">Cylindrinotus disgregus</emphasis>
</taxonomicName>
, male dorsally; F—the same, laterally; G—the same, ventrally; H—labels of the lectotype of
<taxonomicName id="4C556C8E9A725216FF3DF8C9FEF68F24" authorityName="Reitter" authorityYear="1922" box="[248,376,1862,1886]" class="Insecta" family="Tenebrionidae" genus="Cylindrinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="disgregus">
<emphasis id="B921CB1F9A725216FF3DF8C9FEF68F24" box="[248,376,1862,1886]" italics="true" pageId="2" pageNumber="294">C. disgregus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C34F44869A725217FF02F803FC4B8BDF" lastPageId="3" lastPageNumber="295" pageId="2" pageNumber="294" type="description">
<paragraph id="8BEA170D9A725217FF02F803FA0A8AFF" blockId="2.[151,1436,1932,2030]" lastBlockId="3.[151,1437,151,1401]" lastPageId="3" lastPageNumber="295" pageId="2" pageNumber="294">
The second two authors of this paper collected
<taxonomicName id="4C556C8E9A725216FD1AF803FC688FDC" baseAuthorityName="Allard" baseAuthorityYear="1876" box="[735,998,1932,1958]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B921CB1F9A725216FD1AF803FC688FDC" box="[735,998,1932,1958]" italics="true" pageId="2" pageNumber="294">Helopelius aeneipennis</emphasis>
</taxonomicName>
(
<typeStatus id="54EEA9AF9A725216FC30F803FBA98FDC" box="[1013,1063,1932,1958]" pageId="2" pageNumber="294">type</typeStatus>
species of the genus
<taxonomicName id="4C556C8E9A725216FADFF803FA1D8FDC" authorityName="Reitter" authorityYear="1922" box="[1306,1427,1932,1958]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A725216FADFF803FA1D8FDC" box="[1306,1427,1932,1958]" italics="true" pageId="2" pageNumber="294">Helopelius</emphasis>
</taxonomicName>
) in Rhodes Island,
<collectingCountry id="F342579D9A725216FEA6F83FFE388FB0" box="[355,438,1968,1994]" name="Greece" pageId="2" pageNumber="294">Greece</collectingCountry>
. After examination of
<typeStatus id="54EEA9AF9A725216FD76F83FFD6D8FB0" box="[691,739,1968,1994]" pageId="2" pageNumber="294">type</typeStatus>
and non-type material we found, that this species,
<taxonomicName id="4C556C8E9A725216FAE6F83FFE148F94" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="disgregus">
<emphasis id="B921CB1F9A725216FAE6F83FFE8D8F94" italics="true" pageId="2" pageNumber="294">Helopelius disgregus</emphasis>
<bibRefCitation id="EFC46AFC9A725216FECFF85BFE148F94" author="Reitter, E." box="[266,410,2004,2030]" pageId="2" pageNumber="294" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C556C8E9A725216FE0AF85BFBA98F94" authority="(Nabozhenko &amp; Keskin, 2009)" baseAuthorityName="Nabozhenko &amp; Keskin" baseAuthorityYear="2009" box="[463,1063,2004,2030]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="gayirbegi">
<emphasis id="B921CB1F9A725216FE0AF85BFD468F94" box="[463,712,2004,2030]" italics="true" pageId="2" pageNumber="294">Stenohelops gayirbegi</emphasis>
(
<bibRefCitation id="EFC46AFC9A725216FD13F85BFB918F94" author="Nabozhenko, M. V. &amp; Keskin, B." box="[726,1055,2004,2030]" pageId="2" pageNumber="294" pagination="53 - 60" refId="ref5749" refString="Nabozhenko, M. V. &amp; Keskin, B. (2009) Two new species of the genus Gunarus Des Gozis, 1886 (Coleoptera: Tenebrionidae: Helopini) from Southern Turkey. Zootaxa, 2170 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 2170.1.6" type="journal article" year="2009">Nabozhenko &amp; Keskin, 2009</bibRefCitation>
)
</taxonomicName>
are conspecific. A systematic position of two Anatolian species
<taxonomicName id="4C556C8E9A735217FE37FF18FDF688CB" baseAuthorityName="Nabozhenko &amp; Keskin" baseAuthorityYear="2009" box="[498,632,151,177]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="gayirbegi">
<emphasis id="B921CB1F9A735217FE37FF18FDF688CB" box="[498,632,151,177]" italics="true" pageId="3" pageNumber="295">S. gayirbegi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C556C8E9A735217FD68FF18FB3788CB" authority="(Nabozhenko &amp; Keskin, 2009)" baseAuthorityName="Nabozhenko &amp; Keskin" baseAuthorityYear="2009" box="[685,1209,151,177]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="korkutelensis">
<emphasis id="B921CB1F9A735217FD68FF18FCD288CB" box="[685,860,151,177]" italics="true" pageId="3" pageNumber="295">S. korkutelensis</emphasis>
(
<bibRefCitation id="EFC46AFC9A735217FCACFF18FB3F88CB" author="Nabozhenko, M. V. &amp; Keskin, B." box="[873,1201,151,177]" pageId="3" pageNumber="295" pagination="53 - 60" refId="ref5749" refString="Nabozhenko, M. V. &amp; Keskin, B. (2009) Two new species of the genus Gunarus Des Gozis, 1886 (Coleoptera: Tenebrionidae: Helopini) from Southern Turkey. Zootaxa, 2170 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 2170.1.6" type="journal article" year="2009">Nabozhenko &amp; Keskin, 2009</bibRefCitation>
)
</taxonomicName>
within the subgenus
<taxonomicName id="4C556C8E9A735217FF52FF34FE9088AF" box="[151,286,187,213]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="subGenus" subGenus="Stenomaleis">
<emphasis id="B921CB1F9A735217FF52FF34FE9088AF" box="[151,286,187,213]" italics="true" pageId="3" pageNumber="295">Stenomaleis</emphasis>
</taxonomicName>
of the genus
<taxonomicName id="4C556C8E9A735217FE7DFF34FDB188AF" authorityName="Reitter" authorityYear="1922" box="[440,575,187,213]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FE7DFF34FDB188AF" box="[440,575,187,213]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
</taxonomicName>
has been argued earlier (
<bibRefCitation id="EFC46AFC9A735217FC9CFF34FB0F88AC" author="Nabozhenko, M. V. &amp; Perkovsky, E. E. &amp; Chernei, L. S." box="[857,1153,187,214]" pageId="3" pageNumber="295" pagination="947 - 952" refId="ref5942" refString="Nabozhenko, M. V., Perkovsky, E. E. &amp; Chernei, L. S. (2016 a) A new species of the genus Nalassus Mulsant (Coleoptera: Tenebrionidae: Helopini) from the Baltic amber. Paleontological Journal, 50, 947 - 952. https: // doi. org / 10.1134 / S 0031030116090082" type="journal article" year="2016">
Nabozhenko
<emphasis id="B921CB1F9A735217FC35FF34FBAC88AF" box="[1008,1058,187,213]" italics="true" pageId="3" pageNumber="295">et al</emphasis>
., 2016a
</bibRefCitation>
). Respectively, the genus
<taxonomicName id="4C556C8E9A735217FF52FF50FE238883" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" box="[151,429,223,249]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FF52FF50FE9E8883" box="[151,272,223,249]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
<bibRefCitation id="EFC46AFC9A735217FEDFFF50FE238883" author="Reitter, E." box="[282,429,223,249]" pageId="3" pageNumber="295" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
</taxonomicName>
and the subgenus
<taxonomicName id="4C556C8E9A735217FD40FF50FC828883" box="[645,780,223,249]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="subGenus" subGenus="Stenomaleis">
<emphasis id="B921CB1F9A735217FD40FF50FC828883" box="[645,780,223,249]" italics="true" pageId="3" pageNumber="295">Stenomaleis</emphasis>
</taxonomicName>
are congeneric, but Reitter’s taxon has a priority.
<bibRefCitation id="EFC46AFC9A735217FA95FF50FF6C8967" author="Reitter, E." pageId="3" pageNumber="295" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter (1922)</bibRefCitation>
described
<taxonomicName id="4C556C8E9A735217FE9BFE8CFE6B8967" authorityName="Reitter" authorityYear="1922" box="[350,485,259,285]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FE9BFE8CFE6B8967" box="[350,485,259,285]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
</taxonomicName>
and
<taxonomicName id="4C556C8E9A735217FDDAFE8CFD168967" authorityName="Reitter" authorityYear="1922" box="[543,664,259,285]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FDDAFE8CFD168967" box="[543,664,259,285]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
</taxonomicName>
in the same work, but the first one was described a little earlier, on the page 22, Teil I, and the latter was indicated on the page 123, Teil II. We conserve the priority of
<taxonomicName id="4C556C8E9A735217FB3DFEA8FAF1893B" authorityName="Reitter" authorityYear="1922" box="[1272,1407,295,321]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FB3DFEA8FAF1893B" box="[1272,1407,295,321]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
</taxonomicName>
in order to the stability of the nomenclature, guided by the article 23.2 of
<bibRefCitation id="EFC46AFC9A735217FC6BFEC4FBCA891F" author="ICZN" box="[942,1092,331,357]" pageId="3" pageNumber="295" refId="ref5273" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth edition. Adopted by the International Union of Biological Sciences. International Trust of Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
. It should be noted, that
<bibRefCitation id="EFC46AFC9A735217FA90FEC4FF5589F3" author="Allard, E." pageId="3" pageNumber="295" pagination="1 - 80" refId="ref5031" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard (1876</bibRefCitation>
,
<bibRefCitation id="EFC46AFC9A735217FF22FEE0FEAE89F3" author="Allard, E." box="[231,288,367,393]" pageId="3" pageNumber="295" pagination="13 - 268" refId="ref5056" refString="Allard, E. (1877) Revision des Helopides vrais. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 13 - 268." type="journal article" year="1877">1877</bibRefCitation>
) quite correctly compared his “
<taxonomicName id="4C556C8E9A735217FD40FEE0FD7C89F3" authorityName="Allard" authorityYear="1876" box="[645,754,367,393]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FD40FEE0FD7C89F3" box="[645,754,367,393]" italics="true" pageId="3" pageNumber="295">Stenomax</emphasis>
</taxonomicName>
”
<taxonomicName id="4C556C8E9A735217FCC0FEE0FC0589F2" baseAuthorityName="Allard" baseAuthorityYear="1876" box="[773,907,367,392]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B921CB1F9A735217FCC0FEE0FC0589F2" box="[773,907,367,392]" italics="true" pageId="3" pageNumber="295">aeneipennis</emphasis>
</taxonomicName>
with “
<taxonomicName id="4C556C8E9A735217FC1DFEE0FBCB89F3" authorityName="Allard" authorityYear="1876" box="[984,1093,367,393]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FC1DFEE0FBCB89F3" box="[984,1093,367,393]" italics="true" pageId="3" pageNumber="295">Stenomax</emphasis>
</taxonomicName>
”
<taxonomicName id="4C556C8E9A735217FB9CFEE0FB4589F2" baseAuthorityName="Mulsant" baseAuthorityYear="1854" box="[1113,1227,367,392]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="pyrenaeus">
<emphasis id="B921CB1F9A735217FB9CFEE0FB4589F2" box="[1113,1227,367,392]" italics="true" pageId="3" pageNumber="295">pyrenaeus</emphasis>
</taxonomicName>
. So, the following new combination, synonymy and status are proposed:
<taxonomicName id="4C556C8E9A735217FD3AFE1CFB9889D7" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" box="[767,1046,403,429]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FD3AFE1CFCF689D7" box="[767,888,403,429]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
<bibRefCitation id="EFC46AFC9A735217FC44FE1CFB9889D7" author="Reitter, E." box="[897,1046,403,429]" pageId="3" pageNumber="295" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FBE7FE1CFBFA89D7" bold="true" box="[1058,1140,403,429]" pageId="3" pageNumber="295">stat. n.</emphasis>
(from genus to subgenus) =
<taxonomicName id="4C556C8E9A735217FF6BFE38FE5789AB" authority="Espanol 1957" authorityName="Espanol" authorityYear="1957" box="[174,473,438,465]" class="Insecta" family="Tenebrionidae" genus="Stenomaleis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FF6BFE38FEBB89AB" box="[174,309,439,465]" italics="true" pageId="3" pageNumber="295">Stenomaleis</emphasis>
<bibRefCitation id="EFC46AFC9A735217FEF8FE38FE5789AB" author="Espanol, F." box="[317,473,438,465]" pageId="3" pageNumber="295" pagination="19 - 38" refId="ref5166" refString="Espanol, F. (1957) Contribucion al estudio de los Stenohelops Ibericos (Col. Tenebrionidae). Eos, Revista Espanola de Entomologia, 33, 19 - 38." type="journal article" year="1957">Español 1957</bibRefCitation>
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FE20FE38FDBC89AB" bold="true" box="[485,562,439,465]" pageId="3" pageNumber="295">syn. n.</emphasis>
;
<taxonomicName id="4C556C8E9A735217FD84FE38FB1C89A8" authority="(Allard, 1876)" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Allard" baseAuthorityYear="1876" box="[577,1170,439,466]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="aeneipennis" status="comb. nov." subGenus="Helopelius">
<emphasis id="B921CB1F9A735217FD84FE38FD4689AB" box="[577,712,439,465]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
(
<emphasis id="B921CB1F9A735217FD1CFE38FCDC89AB" box="[729,850,439,465]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
)
<emphasis id="B921CB1F9A735217FCA6FE38FC6789AA" box="[867,1001,439,464]" italics="true" pageId="3" pageNumber="295">aeneipennis</emphasis>
(
<bibRefCitation id="EFC46AFC9A735217FC32FE38FB0589A8" author="Allard, E." box="[1015,1163,439,466]" pageId="3" pageNumber="295" pagination="1 - 80" refId="ref5031" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard, 1876</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FB5BFE38FA8B89AB" bold="true" box="[1182,1285,439,465]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FB5BFE38FA8B89AB" box="[1182,1285,439,465]" pageId="3" pageNumber="295" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
=
<taxonomicName id="4C556C8E9A735217FAE6FE38FE10898F" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="disgregus">
<emphasis id="B921CB1F9A735217FAE6FE38FE8D898F" italics="true" pageId="3" pageNumber="295">Helopelius disgregus</emphasis>
<bibRefCitation id="EFC46AFC9A735217FECFFE54FE10898F" author="Reitter, E." box="[266,414,475,501]" pageId="3" pageNumber="295" pagination="1 - 44" refId="ref6085" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FE6CFE54FE79898F" bold="true" box="[425,503,475,501]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FE6CFE54FE79898F" box="[425,503,475,501]" pageId="3" pageNumber="295">syn. n.</taxonomicNameLabel>
</emphasis>
, =
<taxonomicName id="4C556C8E9A735217FDD9FE54FBCF898F" authority="Nabozhenko &amp; Keskin, 2009" authorityName="Nabozhenko &amp; Keskin" authorityYear="2009" box="[540,1089,475,501]" class="Insecta" family="Tenebrionidae" genus="Gunarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="gayirbegi">
<emphasis id="B921CB1F9A735217FDD9FE54FD7E898F" box="[540,752,475,501]" italics="true" pageId="3" pageNumber="295">Gunarus gayirbegi</emphasis>
<bibRefCitation id="EFC46AFC9A735217FD32FE54FBCF898F" author="Nabozhenko, M. V. &amp; Keskin, B." box="[759,1089,475,501]" pageId="3" pageNumber="295" pagination="53 - 60" refId="ref5749" refString="Nabozhenko, M. V. &amp; Keskin, B. (2009) Two new species of the genus Gunarus Des Gozis, 1886 (Coleoptera: Tenebrionidae: Helopini) from Southern Turkey. Zootaxa, 2170 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 2170.1.6" type="journal article" year="2009">Nabozhenko &amp; Keskin, 2009</bibRefCitation>
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FB89FE54FB14898F" bold="true" box="[1100,1178,475,501]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FB89FE54FB14898F" box="[1100,1178,475,501]" pageId="3" pageNumber="295">syn. n.</taxonomicNameLabel>
</emphasis>
As a result of this synonymy, the following new combinations (from
<taxonomicName id="4C556C8E9A735217FD75FE70FCA78A63" authorityName="Reitter" authorityYear="1922" box="[688,809,511,537]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FD75FE70FCA78A63" box="[688,809,511,537]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
</taxonomicName>
to
<taxonomicName id="4C556C8E9A735217FC95FE70FC598A63" authorityName="Reitter" authorityYear="1922" box="[848,983,511,537]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FC95FE70FC598A63" box="[848,983,511,537]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
</taxonomicName>
) are established for five Western Mediterranean species:
<taxonomicName id="4C556C8E9A735217FEA2FDACFC068A47" authority="(Kraatz, 1880)" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Kraatz" baseAuthorityYear="1880" box="[359,904,547,573]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="nodifer" status="comb. nov." subGenus="Helopelius">
<emphasis id="B921CB1F9A735217FEA2FDACFE608A47" box="[359,494,547,573]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
(
<emphasis id="B921CB1F9A735217FE3AFDACFDF68A47" box="[511,632,547,573]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
)
<emphasis id="B921CB1F9A735217FD4CFDACFD548A47" box="[649,730,547,573]" italics="true" pageId="3" pageNumber="295">nodifer</emphasis>
(Kraatz, 1880)
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FC51FDACFC758A47" bold="true" box="[916,1019,547,573]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FC51FDACFC758A47" box="[916,1019,547,573]" pageId="3" pageNumber="295" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4C556C8E9A735217FBCFFDACFEAF8A1B" authority="(Antoine, 1949)" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Antoine" baseAuthorityYear="1949" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="otini" status="comb. nov." subGenus="Helopelius">
<emphasis id="B921CB1F9A735217FBCFFDACFB1F8A47" box="[1034,1169,547,573]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
(
<emphasis id="B921CB1F9A735217FB67FDACFA958A47" box="[1186,1307,547,573]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
)
<emphasis id="B921CB1F9A735217FAE9FDACFAD18A46" box="[1324,1375,547,572]" italics="true" pageId="3" pageNumber="295">otini</emphasis>
(
<bibRefCitation id="EFC46AFC9A735217FAAAFDACFE978A1B" author="Antoine, M." pageId="3" pageNumber="295" pagination="123 - 162" refId="ref5080" refString="Antoine, M. (1949) Note d'Entomologie Marocaine. XLIV. Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides). Bulletin de la Societe des Sciences Naturelles du Maroc, 25 - 27, 123 - 162. [1945 - 1947]" type="journal article" year="1949">Antoine, 1949</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FEE9FDC8FE1D8A1B" bold="true" box="[300,403,583,609]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FEE9FDC8FE1D8A1B" box="[300,403,583,609]" pageId="3" pageNumber="295" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4C556C8E9A735217FE64FDC8FB8C8A1B" authority="(Antoine, 1951)" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Antoine" baseAuthorityYear="1951" box="[417,1026,582,609]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="subsinuatus" status="comb. nov." subGenus="Helopelius">
<emphasis id="B921CB1F9A735217FE64FDC8FDA68A1B" box="[417,552,583,609]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
(
<emphasis id="B921CB1F9A735217FDFDFDC8FD3F8A1B" box="[568,689,583,609]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
)
<emphasis id="B921CB1F9A735217FD07FDC8FCC88A1B" box="[706,838,583,609]" italics="true" pageId="3" pageNumber="295">subsinuatus</emphasis>
(
<bibRefCitation id="EFC46AFC9A735217FC90FDC8FC778A1B" author="Antoine, M." box="[853,1017,582,609]" pageId="3" pageNumber="295" pagination="311 - 321" refId="ref5129" refString="Antoine, M. (1951) Notes d'entomologie marocaine LII. - Coleopteres nouveaux ou interessants. Bulletin de la Societe des Sciences Naturelles du Maroc, 29, 311 - 321. [1949]" type="journal article" year="1951">Antoine, 1951</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FBCBFDC8FBFA8A1B" bold="true" box="[1038,1140,583,609]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FBCBFDC8FBFA8A1B" box="[1038,1140,583,609]" pageId="3" pageNumber="295" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4C556C8E9A735217FB47FDC8FDD18AFF" authority="(Vauloger de Beaupre, 1900)" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Vauloger de Beaupre" baseAuthorityYear="1900" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="verrucosus" status="comb. nov." subGenus="Helopelius">
<emphasis id="B921CB1F9A735217FB47FDC8FA878A1B" box="[1154,1289,583,609]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
(
<emphasis id="B921CB1F9A735217FADFFDC8FA1D8A1B" box="[1306,1427,583,609]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
)
<emphasis id="B921CB1F9A735217FF52FDE4FE9C8AFE" box="[151,274,619,644]" italics="true" pageId="3" pageNumber="295">verrucosus</emphasis>
(Vauloger de Beaupré, 1900)
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FDAFFDE4FD5F8AFF" bold="true" box="[618,721,619,645]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FDAFFDE4FD5F8AFF" box="[618,721,619,645]" pageId="3" pageNumber="295" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4C556C8E9A735217FD1AFDE4FA9C8AFF" authority="(Antoine, 1949)" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Antoine" baseAuthorityYear="1949" box="[735,1298,619,645]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="zaianus" status="comb. nov." subGenus="Helopelius">
<emphasis id="B921CB1F9A735217FD1AFDE4FCE88AFF" box="[735,870,619,645]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
(
<emphasis id="B921CB1F9A735217FCB2FDE4FC7E8AFF" box="[887,1008,619,645]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
)
<emphasis id="B921CB1F9A735217FBC5FDE4FBD88AFE" box="[1024,1110,619,644]" italics="true" pageId="3" pageNumber="295">zaianus</emphasis>
(
<bibRefCitation id="EFC46AFC9A735217FBA0FDE4FA858AFF" author="Antoine, M." box="[1125,1291,619,645]" pageId="3" pageNumber="295" pagination="123 - 162" refId="ref5080" refString="Antoine, M. (1949) Note d'Entomologie Marocaine. XLIV. Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides). Bulletin de la Societe des Sciences Naturelles du Maroc, 25 - 27, 123 - 162. [1945 - 1947]" type="journal article" year="1949">Antoine, 1949</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B921CB1F9A735217FADBFDE4FA0A8AFF" bold="true" box="[1310,1412,619,645]" pageId="3" pageNumber="295">
<taxonomicNameLabel id="A21276649A735217FADBFDE4FA0A8AFF" box="[1310,1412,619,645]" pageId="3" pageNumber="295" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BEA170D9A735217FF02FD00FC4B8BDF" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="295">
The mentioned five species from North Africa and
<collectingCountry id="F342579D9A735217FCCAFD00FCE08AD3" box="[783,878,655,681]" name="Portugal" pageId="3" pageNumber="295">Portugal</collectingCountry>
are different from other
<taxonomicName id="4C556C8E9A735217FB4FFD00FA8D8AD3" authorityName="Reitter" authorityYear="1922" box="[1162,1283,655,681]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FB4FFD00FA8D8AD3" box="[1162,1283,655,681]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
</taxonomicName>
by acute and detached apex of apical piece of aedeagus.
<bibRefCitation id="EFC46AFC9A735217FD41FD3CFCBF8AB7" author="Antoine, M." box="[644,817,691,717]" pageId="3" pageNumber="295" pagination="123 - 162" refId="ref5080" refString="Antoine, M. (1949) Note d'Entomologie Marocaine. XLIV. Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides). Bulletin de la Societe des Sciences Naturelles du Maroc, 25 - 27, 123 - 162. [1945 - 1947]" type="journal article" year="1949">Antoine (1949)</bibRefCitation>
specially noted this character and it was the reason to include these species in a genus other than
<taxonomicName id="4C556C8E9A735217FDAAFD58FD788A8B" authorityName="Reitter" authorityYear="1922" box="[623,758,727,753]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FDAAFD58FD788A8B" box="[623,758,727,753]" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
</taxonomicName>
. The form of apex of aedeagus is diverse within the subgenus
<taxonomicName id="4C556C8E9A735217FF52FD74FE9E8B6F" authorityName="Reitter" authorityYear="1922" box="[151,272,763,789]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FF52FD74FE9E8B6F" box="[151,272,763,789]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
</taxonomicName>
and cannot be used in the super-species taxonomy of this subgenus and generally of the genus
<taxonomicName id="4C556C8E9A735217FA90FD74FF6E8B43" authorityName="Reitter" authorityYear="1922" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FA90FD74FF6E8B43" italics="true" pageId="3" pageNumber="295">Stenohelops</emphasis>
</taxonomicName>
: acute and detached (most of North African—Portuguese species), narrowly rounded and weakly detached (eastern Mediterranean species), acute and not detached (Iberian
<taxonomicName id="4C556C8E9A735217FCABFCCCFB828B27" baseAuthorityName="Seidlitz" baseAuthorityYear="1895" box="[878,1036,835,861]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="protensulus">
<emphasis id="B921CB1F9A735217FCABFCCCFB828B27" box="[878,1036,835,861]" italics="true" pageId="3" pageNumber="295">S. protensulus</emphasis>
</taxonomicName>
), widely rounded (European, North African and Chinese species). Here we interpret five mentioned North African species (
<taxonomicName id="4C556C8E9A735217FBBEFCE8FF658BDF" authority="sensu Antoine (1949))" authorityName="sensu Antoine" authorityYear="1949" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FBBEFCE8FB7A8BFB" box="[1147,1268,871,897]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
sensu
<bibRefCitation id="EFC46AFC9A735217FA87FCE8FF658BDF" author="Antoine, M." pageId="3" pageNumber="295" pagination="123 - 162" refId="ref5080" refString="Antoine, M. (1949) Note d'Entomologie Marocaine. XLIV. Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides). Bulletin de la Societe des Sciences Naturelles du Maroc, 25 - 27, 123 - 162. [1945 - 1947]" type="journal article" year="1949">Antoine (1949))</bibRefCitation>
</taxonomicName>
as the
<taxonomicName id="4C556C8E9A735217FEFFFC04FE3B8BDE" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Vauloger de Beaupre" baseAuthorityYear="1900" box="[314,437,907,932]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B921CB1F9A735217FEFFFC04FE3B8BDE" box="[314,437,907,932]" italics="true" pageId="3" pageNumber="295">verrucosus</emphasis>
</taxonomicName>
species-group within the subgenus
<taxonomicName id="4C556C8E9A735217FC80FC04FC308BDF" authorityName="sensu Antoine" authorityYear="1949" box="[837,958,907,933]" class="Insecta" family="Tenebrionidae" genus="Helopelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B921CB1F9A735217FC80FC04FC308BDF" box="[837,958,907,933]" italics="true" pageId="3" pageNumber="295">Helopelius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F44869A735217FF02FC21FC7A8D03" pageId="3" pageNumber="295" type="distribution">
<paragraph id="8BEA170D9A735217FF02FC21FB2F8C6B" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="295">
<emphasis id="B921CB1F9A735217FF02FC21FD918BB3" bold="true" box="[199,543,942,969]" pageId="3" pageNumber="295">Distribution of the subgenus.</emphasis>
Eastern Mediterranean region: Western
<collectingCountry id="F342579D9A735217FC1BFC20FBCC8BB3" box="[990,1090,943,969]" name="Turkey" pageId="3" pageNumber="295">Anatolia</collectingCountry>
;
<collectingCountry id="F342579D9A735217FB88FC20FB5A8BB3" box="[1101,1236,943,969]" name="Greece" pageId="3" pageNumber="295">Dodecanese</collectingCountry>
islands (Rhodes). Western Mediterranean region: Maghreb (
<collectingCountry id="F342579D9A735217FDB9FC5CFD5A8B97" box="[636,724,979,1005]" name="Algeria" pageId="3" pageNumber="295">Algeria</collectingCountry>
,
<collectingCountry id="F342579D9A735217FD21FC5CFCC08B97" box="[740,846,979,1005]" name="Morocco" pageId="3" pageNumber="295">Morocco</collectingCountry>
); Iberian Peninsula; Southern
<collectingCountry id="F342579D9A735217FB6BFC5CFB748B97" box="[1198,1274,979,1005]" name="France" pageId="3" pageNumber="295">France</collectingCountry>
(Eastern Pyrenees, Montagne Noire) (
<bibRefCitation id="EFC46AFC9A735217FE71FC78FDC68C6B" author="Soldati, F." box="[436,584,1014,1041]" pageId="3" pageNumber="295" pagination="261 - 275" refId="ref6130" refString="Soldati, F. (1995) Les Tenebrionidae de l'Herault: inventaire et ecologie (Coleoptera). Bulletin de la Societe entomologique de France, 100, 261 - 275." type="journal article" year="1995">Soldati 1995</bibRefCitation>
).
<collectingCountry id="F342579D9A735217FD9EFC78FD108C6B" box="[603,670,1015,1041]" name="China" pageId="3" pageNumber="295">China</collectingCountry>
(
<collectingRegion id="4991D9EF9A735217FD69FC78FCC28C6B" box="[684,844,1015,1041]" country="China" name="Sichuan" pageId="3" pageNumber="295">Sichuan Prov.</collectingRegion>
) (
<bibRefCitation id="EFC46AFC9A735217FCA4FC78FB1B8C6B" author="Nabozhenko, M. V. &amp; Ando, K." box="[865,1173,1015,1041]" pageId="3" pageNumber="295" pagination="277 - 327" refId="ref5688" refString="Nabozhenko, M. V. &amp; Ando, K. (2018) Subtribal, generic and subgeneric composition of darkling beetles of the tribe Helopini (Coleoptera: Tenebrionidae) in the Eastern Palaearctic. Acta zoologica Academiae Scientiarum Hungaricae, 64, 277 - 327. https: // doi. org / 10.17109 / AZH. 64.4.277.2018" type="journal article" year="2018">Nabozhenko &amp; Ando 2018</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA170D9A735217FF02FB95FC7A8D03" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="295">
<emphasis id="B921CB1F9A735217FF02FB95FEC58C4E" bold="true" box="[199,331,1050,1076]" pageId="3" pageNumber="295">Bionomics.</emphasis>
The below studied species feed on foliose lichens associated with different trees.
<taxonomicName id="4C556C8E9A735217FAD0FB94FE938C22" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Allard" baseAuthorityYear="1876" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="B921CB1F9A735217FAD0FB94FE938C22" italics="true" pageId="3" pageNumber="295">Stenohelops aeneipennis</emphasis>
</taxonomicName>
was found in
<collectingCountry id="F342579D9A735217FE00FBB0FD9B8C23" box="[453,533,1087,1113]" name="Turkey" pageId="3" pageNumber="295">Turkey</collectingCountry>
at night, mainly on old
<taxonomicName id="4C556C8E9A735217FCEAFBB0FC778C23" box="[815,1017,1087,1113]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="B921CB1F9A735217FCEAFBB0FC778C23" box="[815,1017,1087,1113]" italics="true" pageId="3" pageNumber="295">Juniperus excelsa</emphasis>
</taxonomicName>
; only two specimens were collected in alpine zone of Akdağ ridge (
<collectingRegion id="4991D9EF9A735217FE3FFBECFD1D8C07" box="[506,659,1123,1149]" country="Turkey" name="Denizli" pageId="3" pageNumber="295">Denizli Prov.</collectingRegion>
, Babadag Distr.) under stone, but near the slopes were cut down
<taxonomicName id="4C556C8E9A735217FABDFBECFF658CDA" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="genus">
<emphasis id="B921CB1F9A735217FABDFBECFF658CDA" italics="true" pageId="3" pageNumber="295">Juniperus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C556C8E9A735217FEE6FB08FE0F8CDB" box="[291,385,1159,1185]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="genus">
<emphasis id="B921CB1F9A735217FEE6FB08FE0F8CDB" box="[291,385,1159,1185]" italics="true" pageId="3" pageNumber="295">Quercus</emphasis>
</taxonomicName>
. The species was collected in Rhodes at night near
<taxonomicName id="4C556C8E9A735217FC0EFB08FB178CDB" box="[971,1177,1159,1185]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="species" species="coccifera">
<emphasis id="B921CB1F9A735217FC0EFB08FB178CDB" box="[971,1177,1159,1185]" italics="true" pageId="3" pageNumber="295">Quercus coccifera</emphasis>
</taxonomicName>
and an old trunk of
<taxonomicName id="4C556C8E9A735217FA44FB08FE9C8CBF" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="species" species="macrolepis">
<emphasis id="B921CB1F9A735217FA44FB08FE9C8CBF" italics="true" pageId="3" pageNumber="295">Q. macrolepis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C556C8E9A735217FEE5FB24FDCC8CBF" baseAuthorityName="Nabozhenko &amp; Keskin" baseAuthorityYear="2009" box="[288,578,1195,1221]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="korkutelensis">
<emphasis id="B921CB1F9A735217FEE5FB24FDCC8CBF" box="[288,578,1195,1221]" italics="true" pageId="3" pageNumber="295">Stenohelops korkutelensis</emphasis>
</taxonomicName>
feeds on lichens exclusively on trunks of old
<taxonomicName id="4C556C8E9A735217FB85FB24FA828CBF" box="[1088,1292,1195,1221]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="species" species="coccifera">
<emphasis id="B921CB1F9A735217FB85FB24FA828CBF" box="[1088,1292,1195,1221]" italics="true" pageId="3" pageNumber="295">Quercus coccifera</emphasis>
</taxonomicName>
. The species
<taxonomicName id="4C556C8E9A735217FF52FB40FE2F8C92" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Vauloger de Beaupre" baseAuthorityYear="1900" box="[151,417,1231,1257]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B921CB1F9A735217FF52FB40FE2F8C92" box="[151,417,1231,1257]" italics="true" pageId="3" pageNumber="295">Stenohelops verrucosus</emphasis>
</taxonomicName>
was collected on trunks of old
<taxonomicName id="4C556C8E9A735217FCC0FB40FCEF8C92" box="[773,865,1231,1256]" class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="genus">
<emphasis id="B921CB1F9A735217FCC0FB40FCEF8C92" box="[773,865,1231,1256]" italics="true" pageId="3" pageNumber="295">Tamarix</emphasis>
</taxonomicName>
sp. in
<collectingCountry id="F342579D9A735217FC68FB40FC758C93" box="[941,1019,1231,1257]" name="Morocco" pageId="3" pageNumber="295">Agadir</collectingCountry>
(
<collectingCountry id="F342579D9A735217FBCFFB40FBF88C93" box="[1034,1142,1231,1257]" name="Morocco" pageId="3" pageNumber="295">Morocco</collectingCountry>
).
<taxonomicName id="4C556C8E9A735217FB4CFB40FAC58C92" authorityName="Nabozhenko &amp; Mpamnaras &amp; Kalaentzis" authorityYear="2020" baseAuthorityName="Antoine" baseAuthorityYear="1949" box="[1161,1355,1231,1257]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="otini">
<emphasis id="B921CB1F9A735217FB4CFB40FAC58C92" box="[1161,1355,1231,1257]" italics="true" pageId="3" pageNumber="295">Stenohelops otini</emphasis>
</taxonomicName>
occurs in several localities in Middle Atlas with forests of
<taxonomicName id="4C556C8E9A735217FD22FB7CFC2B8D77" box="[743,933,1267,1293]" class="Pinopsida" family="Pinaceae" genus="Cedrus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="species" species="atlantica">
<emphasis id="B921CB1F9A735217FD22FB7CFC2B8D77" box="[743,933,1267,1293]" italics="true" pageId="3" pageNumber="295">Cedrus atlantica</emphasis>
</taxonomicName>
and probably is associated with these trees.
<taxonomicName id="4C556C8E9A735217FF52FA98FDD78D4B" authority="(Mulsant, 1854)" baseAuthorityName="Mulsant" baseAuthorityYear="1854" box="[151,601,1302,1329]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="pyrenaeus">
<emphasis id="B921CB1F9A735217FF52FA98FE148D4A" box="[151,410,1303,1329]" italics="true" pageId="3" pageNumber="295">Stenohelops pyrenaeus</emphasis>
(Mulsant, 1854)
</taxonomicName>
is also arboreal species and was collected under the bark of
<taxonomicName id="4C556C8E9A735217FAD2FA98FE9E8D2F" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="subSpecies" species="nigra" subSpecies="salzmannii">
<emphasis id="B921CB1F9A735217FAD2FA98FE9E8D2F" italics="true" pageId="3" pageNumber="295">Pinus nigra salzmannii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C556C8E9A735217FEDAFAB4FE228D2F" box="[287,428,1339,1365]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B921CB1F9A735217FEDAFAB4FE228D2F" box="[287,428,1339,1365]" italics="true" pageId="3" pageNumber="295">Quercus ilex</emphasis>
</taxonomicName>
,
<taxonomicName id="4C556C8E9A735217FE7FFAB4FDAA8D2F" box="[442,548,1339,1365]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="genus">
<emphasis id="B921CB1F9A735217FE7FFAB4FDAA8D2F" box="[442,548,1339,1365]" italics="true" pageId="3" pageNumber="295">Castanea</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="4C556C8E9A735217FD47FAB4FD508D2F" box="[642,734,1339,1365]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="295" phylum="Tracheophyta" rank="genus">
<emphasis id="B921CB1F9A735217FD47FAB4FD508D2F" box="[642,734,1339,1365]" italics="true" pageId="3" pageNumber="295">Populus</emphasis>
</taxonomicName>
sp. (
<bibRefCitation id="EFC46AFC9A735217FCD1FAB4FC268D2F" author="Soldati, F." box="[788,936,1338,1365]" pageId="3" pageNumber="295" pagination="261 - 275" refId="ref6130" refString="Soldati, F. (1995) Les Tenebrionidae de l'Herault: inventaire et ecologie (Coleoptera). Bulletin de la Societe entomologique de France, 100, 261 - 275." type="journal article" year="1995">Soldati 1995</bibRefCitation>
).
<taxonomicName id="4C556C8E9A735217FC79FAB4FA128D2F" authority="Lagar &amp; Fresneda" authorityName="Lagar &amp; Fresneda" box="[956,1436,1339,1365]" class="Insecta" family="Tenebrionidae" genus="Stenohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="barcinensis">
<emphasis id="B921CB1F9A735217FC79FAB4FB448D2F" box="[956,1226,1339,1365]" italics="true" pageId="3" pageNumber="295">Stenohelops barcinensis</emphasis>
Lagar &amp; Fresneda
</taxonomicName>
was collected in rotten stump of figs
<taxonomicName id="4C556C8E9A735217FDF3FAD0FC7E8D03" authority="(Lagar &amp; Fresneda, 2014)" baseAuthorityName="Lagar &amp; Fresneda" baseAuthorityYear="2014" box="[566,1008,1375,1401]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="295" phylum="Mollusca" rank="species" species="carica">
<emphasis id="B921CB1F9A735217FDF3FAD0FD4D8D02" box="[566,707,1375,1400]" italics="true" pageId="3" pageNumber="295">Ficus carica</emphasis>
(
<bibRefCitation id="EFC46AFC9A735217FD14FAD0FC668D03" author="Lagar, A. &amp; Fresneda, J." box="[721,1000,1375,1401]" pageId="3" pageNumber="295" pagination="93 - 99" refId="ref5345" refString="Lagar, A. &amp; Fresneda, J. (2014) Stenohelops barcinensis sp. n. de Barcelona (Espana) (Coleoptera, Tenebrionidae, Helopini). Boletin de la Sociedad Entomologica Aragonesa, 55, 93 - 99." type="journal article" year="2014">Lagar &amp; Fresneda, 2014</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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