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<document id="4A91D736E507D957973BCE7BDBAF0973" ID-CLB-Dataset="26916" ID-DOI="10.5281/zenodo.2669927" ID-GBIF-Dataset="f32a6f5a-0c9f-43c4-a273-4bc325ece96e" ID-ISSN="1175-5326" ID-Zenodo-Dep="2669927" ID-ZooBank="F066A962-743F-4899-AFAE-485C5A51EF2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557202401381" checkinUser="plazi" docAuthor="Williams, Kevin A., Lelej, Arkady S., Okayasu, Juriya, Borkent, Christopher J., Malee, Rufeah, Thoawan, Kodeeyah &amp; Thaochan, Narit" docDate="2019" docId="03DCF504FFEF9864FF392BCA4AFDC7FB" docLanguage="en" docName="zootaxa.4602.1.1.pdf" docOrigin="Zootaxa 4602 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Smicromyrme helarctos Williams &amp; Lelej &amp; Okayasu &amp; Borkent &amp; Malee &amp; Thoawan &amp; Thaochan 2019, sp. nov." docType="treatment" docUuid="35C77EAE-646A-4EB6-8E58-6E583FAD8478" docUuidSource="ZooBank" docVersion="20" lastPageNumber="28" masterDocId="FFE58D7CFFF5987FFFAE2F624960C60C" masterDocTitle="The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia" masterLastPageNumber="69" masterPageNumber="1" pageNumber="27" updateTime="1698726454309" updateUser="ExternalLinkService">
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<mods:title id="7F930474EDD7B7D0D6D586F1564B7A7F">The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia</mods:title>
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<mods:namePart id="AC3C5A417F36B14FD1FC919AF3BAE40F">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="223D3621A7CC5DC957B87B21B9E167C8">Lelej, Arkady S.</mods:namePart>
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<mods:namePart id="B1D23C93BC62AE39B9DD853D1591BB87">Okayasu, Juriya</mods:namePart>
</mods:name>
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<mods:namePart id="7563CDC1EBF4BC2F69CE1A5B8A61071B">Borkent, Christopher J.</mods:namePart>
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<mods:roleTerm id="CC912F2615A0230D14E12B170F84A507">Author</mods:roleTerm>
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<mods:namePart id="E6388DE25D6F8E0A6DC93A24B20A7995">Malee, Rufeah</mods:namePart>
</mods:name>
<mods:name id="6235F1DCF51BAE9202B86C8910076CE4" type="personal">
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<mods:roleTerm id="F581DE5C7830A54AE7C17C4EE0C1FDEB">Author</mods:roleTerm>
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<mods:namePart id="8E1AC69EB26CDC2443304F00F0C3EF9E">Thoawan, Kodeeyah</mods:namePart>
</mods:name>
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<mods:namePart id="D605216B6EE75709ADD5EC955E942AC6">Thaochan, Narit</mods:namePart>
</mods:name>
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<mods:title id="71CC3AF5C5FBEEA6F43FF74B65BA19D9">Zootaxa</mods:title>
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<mods:date id="E1C40EBEC2938FD7D7E5A600F1142F48">2019</mods:date>
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<mods:number id="E8C58F7A4B999A7B08FCB4CC95F7B22B">2019-05-07</mods:number>
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<mods:number id="DD12F7A2B9891453617E2127FFB6F343">4602</mods:number>
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<mods:classification id="A63431AE83DA6AE03663073BD391E392">journal article</mods:classification>
<mods:identifier id="5BD418269F24799237F4ED50855CE8E0" type="CLB-Dataset">26916</mods:identifier>
<mods:identifier id="6DD7A16A318BDDDEE65C9B99E336B707" type="DOI">10.5281/zenodo.2669927</mods:identifier>
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<treatment id="03DCF504FFEF9864FF392BCA4AFDC7FB" ID-DOI="http://doi.org/10.5281/zenodo.5923601" ID-GBIF-Taxon="157062794" ID-Zenodo-Dep="5923601" LSID="urn:lsid:zoobank.org:act:35C77EAE-646A-4EB6-8E58-6E583FAD8478" httpUri="http://treatment.plazi.org/id/03DCF504FFEF9864FF392BCA4AFDC7FB" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="C36F1799FFEF9865FF392BCA4BB6C2CE" box="[151,726,1192,1218]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BCA4412FFEF9865FF392BCA4BB6C2CE" blockId="26.[151,726,1192,1251]" box="[151,726,1192,1218]" pageId="26" pageNumber="27">
<heading id="D082F37EFFEF9865FF392BCA4BB6C2CE" bold="true" box="[151,726,1192,1218]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B9019800FFEF9865FF392BCA4BB6C2CE" bold="true" box="[151,726,1192,1218]" pageId="26" pageNumber="27">
15b.
<taxonomicName id="4C753F91FFEF9865FF7D2BCA4B0CC2CE" authority="Williams" authorityName="Williams &amp; Lelej &amp; Okayasu &amp; Borkent &amp; Malee &amp; Thoawan &amp; Thaochan" authorityYear="2019" box="[211,620,1192,1218]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="helarctos" status="sp. nov.">
<emphasis id="B9019800FFEF9865FF7D2BCA4892C2CE" bold="true" box="[211,498,1192,1218]" italics="true" pageId="26" pageNumber="27">Smicromyrme helarctos</emphasis>
Williams
</taxonomicName>
,
<taxonomicNameLabel id="A232257BFFEF9865FDD62BCA4BB6C2CE" box="[632,726,1192,1218]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFEF9865FF392BA84823C2EF" box="[151,323,1226,1251]" pageId="26" pageNumber="27" type="description">
<paragraph id="8BCA4412FFEF9865FF392BA84823C2EF" blockId="26.[151,726,1192,1251]" box="[151,323,1226,1251]" pageId="26" pageNumber="27">
(
<figureCitation id="134E5897FFEF9865FF312BA8485BC2EF" box="[159,315,1226,1251]" captionStart="FIGURES 95106" captionStartId="59.[151,264,1809,1832]" captionTargetBox="[169,1417,199,1788]" captionTargetId="figure@59.[169,1417,193,1788]" captionTargetPageId="59" captionText="FIGURES 95106. Mutillinae: Smicromyrmini females V. 9597. Nemka conjungenda; 98100. Smicromyrme borkenti, sp. nov.; 101103. S. lochius; 104106. S. helarctos, sp. nov. 95, 98, 101, 104. Habitus, dorsal view; 96, 99, 102, 105. Habitus, lateral view; 97, 100, 103, 106. T6." httpUri="https://zenodo.org/record/2669943/files/figure.png" pageId="26" pageNumber="27">Figs 104106</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFEF9865FF392A704DDAC39B" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BCA4412FFEF9865FF392A704DDAC39B" blockId="26.[151,1437,1298,2007]" pageId="26" pageNumber="27">
<emphasis id="B9019800FFEF9865FF392A704871C327" bold="true" box="[151,273,1298,1323]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
FEMALE. This species has unique coloration in Southeast Asia, wherein the mesosoma is black except for the orange-brown dorsal face. The head is entirely black, the legs are entirely orange, the metasoma is mostly black with a single basomedial spot on the T2 disc. The scutellar scale is broadly arcuate and the pygidium is elongate oval with sparse striae and the apex smooth. Body length
<quantity id="4C8DE9F7FFEF9865FC2D2A1D4ABCC39B" box="[899,988,1407,1431]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="26" pageNumber="27" unit="mm" value="3.4">3.4 mm</quantity>
. MALE. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFEF9864FF692AC04865C6D9" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BCA4412FFEF9864FF692AC04865C6D9" blockId="26.[151,1437,1298,2007]" lastBlockId="27.[151,1436,151,504]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="B9019800FFEF9865FF692AC04839C3B7" bold="true" box="[199,345,1442,1467]" pageId="26" pageNumber="27">Description.</emphasis>
FEMALE. Body length
<quantity id="4C8DE9F7FFEF9865FDC22AC14BA4C3B6" box="[620,708,1443,1467]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="26" pageNumber="27" unit="mm" value="3.4">3.4 mm</quantity>
.
<emphasis id="B9019800FFEF9865FD602AC04A31C3B7" box="[718,849,1442,1467]" italics="true" pageId="26" pageNumber="27">Coloration.</emphasis>
Head black-brown, except frons, mandible, clypeus, and antenna largely orange-brown. Mesosoma black-brown, except dorsum orange-brown and legs pale orange. Metasoma black, except T1, S1, T2 apically, T3, and S26 red-brown to yellow-brown. Body setae generally sparse and silvery, except vertex, mesosomal dorsum, T2, and T45 with blackish setae; and T1 apex, T2 basomedial spot, T2 apex, T3 entirely, and T6 basolaterally with dense whitish silver setae.
<emphasis id="B9019800FFEF9865FB1B29504D91C047" box="[1205,1265,1586,1611]" italics="true" pageId="26" pageNumber="27">Head</emphasis>
. Width behind eye 0.95 × mesosoma width. Frons, vertex, and gena with separated punctures. Mandible apex obscurely tridentate. Clypeus with narrow transverse medially-emarginate lamella; basomedial portion with carinae forming acute triangle. Antennal scrobe without dorsal carina. Antennal tubercle smooth. Genal carina obliterated. F1 1.2 × pedicel length, F2 1.2 × pedicel length.
<emphasis id="B9019800FFEF9865FDD629A64B90C0D7" box="[632,752,1732,1755]" italics="true" pageId="26" pageNumber="27">Mesosoma</emphasis>
. Length 0.95 × width. Dorsum of mesosoma with coarse confluent punctures. Side of mesosoma with scattered moderate punctures with long setae, interspaces mostly smooth, propodeal side with distinct reticulations posteriorly. Mesopleural lamella absent. Humeral carina simple, rounded dorsally. Ratio of width of humeral angle, anterior spiracle, narrowest point of mesonotum, propodeal spiracle, and midpoint of propodeum 42:46:45:46:45. Scutellar scale transverse arculate, ~9 punctures wide, with transverse wavy carina anterior and posterior to scale. Posterior propodeal face reticulate. Lateral and posterior propodeal faces not separated by carina. Metatibio-tarsal ratio 45:25:18:15:12:9.
<emphasis id="B9019800FFEF9865FB8428FE4DC6C1BF" box="[1066,1190,1948,1971]" italics="true" pageId="26" pageNumber="27">Metasoma.</emphasis>
Terga 15 with small separated punctures, sparser on T1. S1 with simple longitudinal carina. T2 felt line 0.4 × T2 total length. T6 with long ovate pygidium; with parallel striae (16 at widest point), posterior quarter smooth. S6 posterior margin bidentate.
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFEE9864FF692FBD4BBCC713" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BCA4412FFEE9864FF692FBD4BBCC713" blockId="27.[151,1436,151,504]" pageId="27" pageNumber="28">
<emphasis id="B9019800FFEE9864FF692FBD4B38C6F4" bold="true" box="[199,600,223,248]" pageId="27" pageNumber="28">
Material examined.
<materialsCitation id="3B1D4E4FFFEE9864FE6D2FBD4B38C6F4" ID-GBIF-Occurrence="2238655927" box="[451,600,223,248]" collectingDate="2018-05-21" collectionCode="CSCA" country="Thailand" elevation="180" latitude="14.24" location="Phalad" longLatPrecision="773" longitude="99.34" municipality="Mueang" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" stateProvince="Kanchanaburi" typeStatus="holotype">
<typeStatus id="54CEFAB0FFEE9864FE6D2FBD4B50C6F4" box="[451,560,223,248]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
<specimenCount id="9D738F9BFFEE9864FD922FBD4B31C6F4" box="[572,593,223,248]" pageId="27" pageNumber="28" type="female"></specimenCount>
,
</materialsCitation>
</emphasis>
<collectingCountry id="F3620482FFEE9864FDCB2F824B99C6F4" box="[613,761,224,248]" name="Thailand" pageId="27" pageNumber="28">THAILAND</collectingCountry>
,
<collectingRegion id="49B18AF0FFEE9864FCA72FBD4ACBC6F4" box="[777,939,223,248]" country="Thailand" name="Kanchanaburi" pageId="27" pageNumber="28">
<emphasis id="B9019800FFEE9864FCA72FBD4ACBC6F4" box="[777,939,223,248]" italics="true" pageId="27" pageNumber="28">Kanchanaburi</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BAEDE68FFEE9864FC112F824D7BC6F4" box="[959,1051,224,248]" pageId="27" pageNumber="28">Mueang</collectingMunicipality>
<collectingRegion id="49B18AF0FFEE9864FB862FBD4DA8C6F4" box="[1064,1224,223,248]" country="Thailand" name="Kanchanaburi" pageId="27" pageNumber="28">Kanchanaburi</collectingRegion>
,
<location id="8EAA12C9FFEE9864FB762FBD4C44C6F4" LSID="urn:lsid:plazi:treatment:03DCF504FFEF9864FF392BCA4AFDC7FB:8EAA12C9FFEE9864FB762FBD4C44C6F4" box="[1240,1316,223,248]" country="Thailand" latitude="14.24" longLatPrecision="773" longitude="99.34" municipality="Mueang" name="Phalad" pageId="27" pageNumber="28" stateProvince="Kanchanaburi">Phalad</location>
waterfall,
<geoCoordinate id="EE4122D5FFEE9864FF392E654991C713" box="[151,241,259,287]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="14.24">
14.24
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N
</geoCoordinate>
<geoCoordinate id="EE4122D5FFEE9864FF572E654832C713" box="[249,338,259,287]" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="555" value="99.34">
99.34
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E
</geoCoordinate>
,
<elevation id="0058A321FFEE9864FEF02E6548C7C713" box="[350,423,263,287]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="180.0">
<quantity id="4C8DE9F7FFEE9864FEF02E6548C7C713" box="[350,423,263,287]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="180.0">180 m</quantity>
</elevation>
,
<collectingDate id="EF8F9B3AFFEE9864FE1D2E654B48C713" box="[435,552,263,287]" pageId="27" pageNumber="28" value="2018-05-21">
<date id="FFCB62D2FFEE9864FE1D2E654B48C713" box="[435,552,263,287]" pageId="27" pageNumber="28" value="2018-05-21">21.V.2018</date>
</collectingDate>
, MKT (CSCA).
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFEE9864FF692E484D51C74F" box="[199,1073,298,323]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BCA4412FFEE9864FF692E484D51C74F" blockId="27.[151,1436,151,504]" box="[199,1073,298,323]" pageId="27" pageNumber="28">
<emphasis id="B9019800FFEE9864FF692E484800C74F" bold="true" box="[199,352,298,323]" pageId="27" pageNumber="28">Distribution.</emphasis>
Known only from the
<typeStatus id="54CEFAB0FFEE9864FDCE2E494BF0C74F" box="[608,656,299,323]" pageId="27" pageNumber="28">type</typeStatus>
locality in
<collectingCountry id="F3620482FFEE9864FCA12E484A13C74F" box="[783,883,298,323]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
(
<collectingRegion id="49B18AF0FFEE9864FC2F2E484D45C74F" box="[897,1061,298,323]" country="Thailand" name="Kanchanaburi" pageId="27" pageNumber="28">Kanchanaburi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFEE9864FF692E2C4C2EC787" pageId="27" pageNumber="28" type="etymology">
<paragraph id="8BCA4412FFEE9864FF692E2C4C2EC787" blockId="27.[151,1436,151,504]" pageId="27" pageNumber="28">
<emphasis id="B9019800FFEE9864FF692E2C482EC76B" bold="true" box="[199,334,334,359]" pageId="27" pageNumber="28">Etymology.</emphasis>
This species is named after the Sun Bear,
<emphasis id="B9019800FFEE9864FC982E2C4D45C76B" box="[822,1061,334,359]" italics="true" pageId="27" pageNumber="28">Helarctos malayanus</emphasis>
(
<bibRefCitation id="EFE439E3FFEE9864FB982E2C4DB2C76B" author="Raffles, T. S." box="[1078,1234,334,359]" pageId="27" pageNumber="28" pagination="239 - 274" refId="ref60739" refString="Raffles, T. S. (1821) XVII. Descriptive Catalogue of a Zoological Collection, made on account of the Honourable East India Company, in the Island of Sumatra and its Vicinity, under the Direction of Sir Thomas Stamford Raffles, Lieutenant- Governor of Fort Marlborough; with additional Notices illustrative of the Natural History of those Countries. Transactions of the Linnean Society of London, 1, 239 - 274." type="journal article" year="1821">Raffles, 1821</bibRefCitation>
), one of two bear species native to Southeast Asia, and for all the other bears around the World. Treat as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36F1799FFEE9864FF692EF44AFDC7FB" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BCA4412FFEE9864FF692EF44AFDC7FB" blockId="27.[151,1436,151,504]" pageId="27" pageNumber="28">
<emphasis id="B9019800FFEE9864FF692EF4485BC7A3" bold="true" box="[199,315,406,431]" pageId="27" pageNumber="28">Remarks.</emphasis>
With its broad scutellar scale and pygidium shape and sculpture, this species is structurally similar to
<emphasis id="B9019800FFEE9864FF1A2ED8483AC7DF" box="[180,346,442,467]" italics="true" pageId="27" pageNumber="28">Mickelomyrme</emphasis>
. We treat it as a member of
<taxonomicName id="4C753F91FFEE9864FD322ED94A57C7DE" authorityName="Thomson" authorityYear="1870" box="[668,823,443,466]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9019800FFEE9864FD322ED94A57C7DE" box="[668,823,443,466]" italics="true" pageId="27" pageNumber="28">Smicromyrme</emphasis>
</taxonomicName>
because T2 has only one medial setal spot. Discovery of the male could provide greater confidence in the genus placement.
</paragraph>
</subSubSection>
</treatment>
</document>