404 lines
53 KiB
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404 lines
53 KiB
XML
<document id="1F81503017E206A05199E1C7C42DB2E3" ID-CLB-Dataset="7027" ID-DOI="10.5252/geodiversitas2021v43a10" ID-GBIF-Dataset="bb69ed40-0261-4e0d-8264-ef27d3064148" ID-ISSN="1638-9395" ID-Zenodo-Dep="4746542" ID-ZooBank="urn:lsid:zoobank.org:pub:F9F1091A-DDD1-40C4-81B6-F254151D7D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1620665489502" checkinUser="felipe" docAuthor="Laville, Thomas, Haug, Joachim T. & Haug, Carolin" docDate="2021" docId="03DC0462FFA5D523FEF7FF3EFBC36EF4" docLanguage="en" docName="Geodiversitas.43.10.295-310.pdf" docOrigin="Geodiversitas 43 (10)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Eodollocaris keithflinti Laville, Haug & Haug, 2021, n. gen., n. sp." docType="treatment" docVersion="6" lastPageNumber="303" masterDocId="FFE57C1AFFACD529FFBDFFE9FFB66918" masterDocTitle="New species of Thylacocephala, Eodollocaris keithflinti n. gen., n. sp., from the Mazon Creek Lagerstätte, Illinois, United States (c. 307 Ma) and redescription of other Mazon Creek thylacocephalans" masterLastPageNumber="310" masterPageNumber="295" pageNumber="302" updateTime="1698938558105" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="05E889A61CC7803794FAAA6D8F09C715">New species of Thylacocephala, Eodollocaris keithflinti n. gen., n. sp., from the Mazon Creek Lagerstätte, Illinois, United States (c. 307 Ma) and redescription of other Mazon Creek thylacocephalans</mods:title>
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<mods:name id="82141BF10BC629E3FA07DCFDD2D2DA25" type="personal">
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<mods:role id="0758BA3C438F6A16D058AE9F30419DAA">
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<mods:roleTerm id="64457EFC1B9695851DAB189B3C897C92">Author</mods:roleTerm>
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<mods:namePart id="47E446B8179542C9D98BF9D767E8F1BE">Laville, Thomas</mods:namePart>
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<mods:affiliation id="C51A9ABEBA1E93225EE15C96104081D0">Centre de Recherche en Paléontologie (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, Muséum national d’Histoire naturelle, Département Origines et Évolution, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) and Department of Biology, Ludwig-Maximilians-Universität München, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany)</mods:affiliation>
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<mods:nameIdentifier id="5BBFBC6CD654E447E10601DA54874F37" type="email">thomas.laville1@edu.mnhn.fr</mods:nameIdentifier>
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<mods:roleTerm id="A255BB6F7C04555C515A73F5C5385B2A">Author</mods:roleTerm>
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<mods:namePart id="E4B000205121F89F1FB1F6F812A9D35E">Haug, Joachim T.</mods:namePart>
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<mods:affiliation id="7899A0037155550A8321EA248E6F0634">Department of Biology, Ludwig-Maximilians-Universität München, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) and GeoBio-Center, Ludwig-Maximilians-Universität München, Richard-Wagner-Strasse 10, 80333 München (Germany)</mods:affiliation>
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<mods:nameIdentifier id="24B2EAA803A35F2BCA6328F0166044B9" type="email">joachim.haug@palaeo-evo-devo.info</mods:nameIdentifier>
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<mods:namePart id="D5DB02187FEF77EA352A3F96FAF52A74">Haug, Carolin</mods:namePart>
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<mods:affiliation id="4356CDD640D540F84492B5E73305AC36">Department of Biology, Ludwig-Maximilians-Universität München, Grosshaderner Strasse 2, 82152 Planegg-Martinsried (Germany) and GeoBio-Center, Ludwig-Maximilians-Universität München, Richard-Wagner-Strasse 10, 80333 München (Germany)</mods:affiliation>
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<mods:nameIdentifier id="D1A80143701FFCA16114806771044B85" type="email">carolin.haug@palaeo-evo-devo.info</mods:nameIdentifier>
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<mods:title id="2B226C25A8C52F4D8B31B5F68EE3E3A5">Geodiversitas</mods:title>
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<mods:part id="53CDDC4BC54B4933FB020AD54F1EE88A">
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<mods:date id="C09241DF9E8275F0E7AC87570216B848">2021</mods:date>
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<mods:number id="B01B8FF4AE925C456B6AB0AC0D5A6F06">2021-05-06</mods:number>
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<mods:number id="15D850B3D21BE91B2CAE79B3A6B3AE1B">43</mods:number>
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<treatment id="03DC0462FFA5D523FEF7FF3EFBC36EF4" ID-DOI="http://doi.org/10.5281/zenodo.4746754" ID-Zenodo-Dep="4746754" LSID="urn:lsid:plazi:treatment:03DC0462FFA5D523FEF7FF3EFBC36EF4" httpUri="http://treatment.plazi.org/id/03DC0462FFA5D523FEF7FF3EFBC36EF4" lastPageId="10" lastPageNumber="303" pageId="9" pageNumber="302">
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<subSection id="E2FAAC9FFFA5D520FEF7FF3EFD316CD5" pageId="9" pageNumber="302" type="multiple">
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<subSubSection id="C36FE6FFFFA5D520FEF7FF3EFE4A682A" pageId="9" pageNumber="302" type="nomenclature">
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<paragraph id="8BCAB574FFA5D520FEF7FF3EFE4A682A" blockId="9.[216,692,215,306]" pageId="9" pageNumber="302">
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<heading id="D0820218FFA5D520FEF7FF3EFDF769EA" box="[330,577,215,242]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="302" reason="2">
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<taxonomicName id="4C75CEF7FFA5D520FEF7FF3EFDF769EA" box="[330,577,215,242]" pageId="9" pageNumber="302">
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<emphasis id="B9016966FFA5D520FEF7FF3EFDF769EA" bold="true" box="[330,577,215,242]" italics="true" pageId="9" pageNumber="302">Eodollocaris keithflinti</emphasis>
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</taxonomicName>
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</heading>
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Laville, J.T. Haug & C. Haug
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<taxonomicNameLabel id="A232D41DFFA5D520FD9CFF11FD02680A" box="[545,692,248,274]" pageId="9" pageNumber="302" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="134EA9F1FFA5D520FE25FEFEFE616829" box="[408,471,279,305]" captionStart="FIG" captionStartId="10.[132,143,1542,1559]" captionTargetBox="[133,1455,215,1499]" captionTargetId="figure-214@10.[698,1267,764,1317]" captionTargetPageId="10" captionText="FIG. 6. — Eodollocaris keithflinti Laville,J.T. Haug & C. Haug, n. gen.,n. sp., ROMIP61586: A, B, general view of part and counter-part;C, line drawing of A; D, line drawing of B; E, close-up of the dorsal midline of A; F, close-up of ventral margin of A; G, close-up of prominent appendages of A; H, close-up of prominent appendages of B. Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline; mv, ventral margin; r, rostrum; s, shield; st, stalk; t, trunk; ta, trunk appendage;?, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="9" pageNumber="302">Figs 6</figureCitation>
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;
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<figureCitation id="134EA9F1FFA5D520FE58FEF1FE45682A" box="[485,499,280,306]" captionStart="FIG" captionStartId="11.[132,143,1262,1279]" captionTargetBox="[133,1455,215,1219]" captionTargetId="figure-341@11.[699,1268,213,763]" captionTargetPageId="11" captionText="FIG. 7. — Eodollocaris keithflinti Laville, J.T. Haug & C. Haug, n. gen., n. sp., ROMIP61586: A, B, general view of part and counter-part (color-marked); C, closeup of appendages of A (color-marked); D, close-up of appendages of B (color-marked). Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline;mv, ventral margin;r, rostrum; s, shield;st, stalk;t, trunk;ta, trunk appendage;?, indeterminate oval plate;1-8, spines 1-8. Photographs: C. Haug and J. T. Haug.. Scale bars: A, B, 5 mm; C, D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746556" httpUri="https://zenodo.org/record/4746556/files/figure.png" pageId="9" pageNumber="302">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36FE6FFFFA5D520FF39FEB0FCB16872" box="[132,775,345,362]" pageId="9" pageNumber="302" type="description">
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<paragraph id="8BCAB574FFA5D520FF39FEB0FCB16872" blockId="9.[132,775,345,362]" box="[132,775,345,362]" pageId="9" pageNumber="302">
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<uri id="FFE4B976FFA5D520FF39FEB0FCB16872" box="[132,775,345,362]" pageId="9" pageNumber="302">
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urn:lsid:zoobank.org:act:
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<uuid id="FFD38FA1FFA5D520FEDDFEB0FCB16872" box="[352,775,345,362]" pageId="9" pageNumber="302">1D59F66C-0B63-460B-94AB-F177841EC908</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36FE6FFFFA5D520FF39FE72FDF46B29" pageId="9" pageNumber="302" type="etymology">
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<paragraph id="8BCAB574FFA5D520FF39FE72FD366B1A" blockId="9.[132,777,411,515]" pageId="9" pageNumber="302">ETYMOLOGY. –— In honor of the late Keith Flint from the band ‘The Prodigy’. As the specimen exhibits a certain “aggressiveness” and the fossil is preserved in bright red colour reminding of fire, we herewith tribute to the song ‘Firestarter’ of the band.</paragraph>
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<paragraph id="8BCAB574FFA5D520FF39FDF3FDF46B29" blockId="9.[132,578,537,562]" box="[132,578,537,562]" pageId="9" pageNumber="302">
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TYPE MATERIAL. —
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<typeStatus id="54CE0BD6FFA5D520FEF9FDF3FE156B2A" box="[324,419,538,562]" pageId="9" pageNumber="302" type="holotype">Holotype</typeStatus>
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: ROMIP61586.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36FE6FFFFA5D520FF39FDA3FCBE6BB3" pageId="9" pageNumber="302" type="materials_examined">
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<paragraph id="8BCAB574FFA5D520FF39FDA3FCBE6B64" blockId="9.[132,776,585,636]" pageId="9" pageNumber="302">
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<collectionCode id="ED642DB1FFA5D520FF39FDA3FEA66B78" box="[132,272,586,609]" pageId="9" pageNumber="302">
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<typeStatus id="54CE0BD6FFA5D520FF39FDA3FF056B47" box="[132,179,586,609]" pageId="9" pageNumber="302">TYPE</typeStatus>
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HORIZON
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</collectionCode>
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. — Carbondale Formation, Francis Creek Shale Member,
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<emphasis id="B9016966FFA5D520FF16FD8DFF0C6B63" box="[171,186,612,635]" italics="true" pageId="9" pageNumber="302">c.</emphasis>
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307 Ma, Westphalian D, Middle Pennsylvanian, Carboniferous.
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</paragraph>
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<paragraph id="8BCAB574FFA5D520FF39FD7DFCBE6BB3" blockId="9.[132,776,659,683]" box="[132,776,659,683]" pageId="9" pageNumber="302">
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<collectionCode id="ED642DB1FFA5D520FF39FD7DFEA56BB2" box="[132,275,660,683]" pageId="9" pageNumber="302">
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<typeStatus id="54CE0BD6FFA5D520FF39FD7DFF036BB1" box="[132,181,660,683]" pageId="9" pageNumber="302">TYPE</typeStatus>
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LOCALITY
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</collectionCode>
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. — Mazon Creek area, Pit 11,
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<collectingRegion id="49B17B96FFA5D520FD88FD7AFDCC6BB3" box="[565,634,659,683]" country="United States of America" name="Illinois" pageId="9" pageNumber="302">Illinois</collectingRegion>
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,
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<collectingCountry id="F362F5E4FFA5D520FD3FFD7AFCB26BB3" box="[642,772,659,683]" name="United States of America" pageId="9" pageNumber="302">United States</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36FE6FFFFA5D520FF39FD2AFE4B6A5D" pageId="9" pageNumber="302" type="diagnosis">
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<paragraph id="8BCAB574FFA5D520FF39FD2AFE4B6A5D" blockId="9.[130,775,707,837]" pageId="9" pageNumber="302">DIAGNOSIS. — Trapezoidal shield displaying a rounded rostrum, a tube-shaped dorsal midline, a finely tuberculated ventral margin, a dorsal midline ending posteriorly with a spine and a posterior margin with a notch in its dorsal part. Third prominent appendage bearing seven spines on its most distal element.</paragraph>
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</subSubSection>
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<subSubSection id="C36FE6FFFFA5D520FF39FC9CFD316CD5" pageId="9" pageNumber="302" type="description">
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<paragraph id="8BCAB574FFA5D520FF39FC9CFEA26A95" blockId="9.[130,777,885,1485]" box="[132,276,885,911]" pageId="9" pageNumber="302">DESCRIPTION</paragraph>
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<paragraph id="8BCAB574FFA5D520FF39FC7CFF016AB7" blockId="9.[130,777,885,1485]" box="[132,183,917,943]" pageId="9" pageNumber="302">
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<emphasis id="B9016966FFA5D520FF39FC7CFF016AB7" box="[132,183,917,943]" italics="true" pageId="9" pageNumber="302">Body</emphasis>
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</paragraph>
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<paragraph id="8BCAB574FFA5D520FF39FC5CFD316CD5" blockId="9.[130,777,885,1485]" pageId="9" pageNumber="302">
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Largely enveloped by prominent folded shield (
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<figureCitation id="134EA9F1FFA5D520FD33FC5CFD796AD7" box="[654,719,949,975]" captionStart="FIG" captionStartId="10.[132,143,1542,1559]" captionTargetBox="[133,1455,215,1499]" captionTargetId="figure-214@10.[698,1267,764,1317]" captionTargetPageId="10" captionText="FIG. 6. — Eodollocaris keithflinti Laville,J.T. Haug & C. Haug, n. gen.,n. sp., ROMIP61586: A, B, general view of part and counter-part;C, line drawing of A; D, line drawing of B; E, close-up of the dorsal midline of A; F, close-up of ventral margin of A; G, close-up of prominent appendages of A; H, close-up of prominent appendages of B. Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline; mv, ventral margin; r, rostrum; s, shield; st, stalk; t, trunk; ta, trunk appendage;?, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="9" pageNumber="302">Figs 6</figureCitation>
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A-D; 7A, B). Few structures protruding from it. Exact origin of shield unclear, most likely relatively far anterior, possibly from segments of head. Shield trapezoidal in lateral view; longer than high, about 2.8× (h =
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<quantity id="4C8D1891FFA5D520FE01FBDDFD906D57" box="[444,550,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="6.13" pageId="9" pageNumber="302" unit="mm" value="6.13">6.13 mm</quantity>
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,
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<subScript id="17F1B731FFA5D520FD8EFBDDFDFD6D49" attach="left" box="[563,587,1076,1105]" fontSize="7" pageId="9" pageNumber="302">hp</subScript>
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=
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<quantity id="4C8D1891FFA5D520FDD6FBDDFD626D57" box="[619,724,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="4.97" pageId="9" pageNumber="302" unit="mm" value="4.97">4.97 mm</quantity>
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;
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<subScript id="17F1B731FFA5D520FD5FFBDDFD596D49" attach="left" box="[738,751,1076,1105]" fontSize="7" pageId="9" pageNumber="302">ls</subScript>
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=
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<quantity id="4C8D1891FFA5D520FF3BFBBDFF4C6D77" box="[134,250,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="1.693" pageId="9" pageNumber="302" unit="mm" value="16.93">16.93 mm</quantity>
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;
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<subScript id="17F1B731FFA5D520FEB9FBBDFEA06D69" attach="left" box="[260,278,1108,1137]" fontSize="7" pageId="9" pageNumber="302">lw</subScript>
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=
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<quantity id="4C8D1891FFA5D520FE93FBBDFE156D77" box="[302,419,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="1.523" pageId="9" pageNumber="302" unit="mm" value="15.23">15.23 mm</quantity>
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). Antero-dorsal corner drawn out anteriorly into a rostrum with a rounded tip (l =
|
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<quantity id="4C8D1891FFA5D520FD2EFB9DFD4D6D97" box="[659,763,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="302" unit="mm" value="1.7">1.70 mm</quantity>
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||
). The anterior margin is straight, oriented postero-ventrally (
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||
<emphasis id="B9016966FFA5D520FF30FB5CFF2B6DD6" box="[141,157,1205,1230]" italics="true" pageId="9" pageNumber="302">c.</emphasis>
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||
30°). No clear optical notch distinguishable. Dorsal midline is slightly convex. Postero-dorsal corner drawn out posteriorly into small spine. Ventral margin is convex. Rim in this region is bearing numerous fine tubercles and appears slightly bent in its central part (
|
||
<figureCitation id="134EA9F1FFA5D520FEF4FADAFE2E6C56" box="[329,408,1331,1358]" captionStart="FIG" captionStartId="10.[132,143,1542,1559]" captionTargetBox="[133,1455,215,1499]" captionTargetId="figure-214@10.[698,1267,764,1317]" captionTargetPageId="10" captionText="FIG. 6. — Eodollocaris keithflinti Laville,J.T. Haug & C. Haug, n. gen.,n. sp., ROMIP61586: A, B, general view of part and counter-part;C, line drawing of A; D, line drawing of B; E, close-up of the dorsal midline of A; F, close-up of ventral margin of A; G, close-up of prominent appendages of A; H, close-up of prominent appendages of B. Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline; mv, ventral margin; r, rostrum; s, shield; st, stalk; t, trunk; ta, trunk appendage;?, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="9" pageNumber="302">Fig. 6F</figureCitation>
|
||
). Posterior margin further dorsally concave forming distinct notch, ventral part straight and oriented antero-ventrally (
|
||
<emphasis id="B9016966FFA5D520FEC7FA9DFE3C6C95" box="[378,394,1396,1421]" italics="true" pageId="9" pageNumber="302">c.</emphasis>
|
||
30°). Dorsal midline connecting the two valves tube-shaped, with numerous pores along the entire dorsal line, terminated by the rostrum (
|
||
<figureCitation id="134EA9F1FFA5D520FD97FA5AFDCD6CD5" box="[554,635,1459,1485]" captionStart="FIG" captionStartId="10.[132,143,1542,1559]" captionTargetBox="[133,1455,215,1499]" captionTargetId="figure-214@10.[698,1267,764,1317]" captionTargetPageId="10" captionText="FIG. 6. — Eodollocaris keithflinti Laville,J.T. Haug & C. Haug, n. gen.,n. sp., ROMIP61586: A, B, general view of part and counter-part;C, line drawing of A; D, line drawing of B; E, close-up of the dorsal midline of A; F, close-up of ventral margin of A; G, close-up of prominent appendages of A; H, close-up of prominent appendages of B. Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline; mv, ventral margin; r, rostrum; s, shield; st, stalk; t, trunk; ta, trunk appendage;?, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="9" pageNumber="302">Fig. 6E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
<subSection id="E2FAAC9FFFA5D520FF39FA1AFDC36F94" pageId="9" pageNumber="302" type="multiple">
|
||
<paragraph id="8BCAB574FFA5D520FF39FA1AFF5F6F14" blockId="9.[132,233,1523,1548]" box="[132,233,1523,1548]" pageId="9" pageNumber="302">
|
||
<heading id="D0820218FFA5D520FF39FA1AFF5F6F14" box="[132,233,1523,1548]" fontSize="11" level="2" pageId="9" pageNumber="302" reason="8">
|
||
<emphasis id="B9016966FFA5D520FF39FA1AFF5F6F14" box="[132,233,1523,1548]" italics="true" pageId="9" pageNumber="302">Large eyes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA5D520FF39F9FBFDC36F94" blockId="9.[131,775,1554,1677]" pageId="9" pageNumber="302">
|
||
Anteriorly protruding from under the shield, prominent (l =
|
||
<quantity id="4C8D1891FFA5D520FF39F9DAFF5F6F55" box="[132,233,1587,1613]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="9" pageNumber="302" unit="mm" value="0.94">0.94 mm</quantity>
|
||
;
|
||
<figureCitation id="134EA9F1FFA5D520FF4EF9DBFE996F54" box="[243,303,1586,1612]" captionStart="FIG" captionStartId="10.[132,143,1542,1559]" captionTargetBox="[133,1455,215,1499]" captionTargetId="figure-214@10.[698,1267,764,1317]" captionTargetPageId="10" captionText="FIG. 6. — Eodollocaris keithflinti Laville,J.T. Haug & C. Haug, n. gen.,n. sp., ROMIP61586: A, B, general view of part and counter-part;C, line drawing of A; D, line drawing of B; E, close-up of the dorsal midline of A; F, close-up of ventral margin of A; G, close-up of prominent appendages of A; H, close-up of prominent appendages of B. Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline; mv, ventral margin; r, rostrum; s, shield; st, stalk; t, trunk; ta, trunk appendage;?, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="9" pageNumber="302">Fig. 6</figureCitation>
|
||
A-D). Appear to arise from massive proximal stalks. Posterior to eyes, a massive oval structure emerging ventrally from the shield. Exact origin unclear.
|
||
</paragraph>
|
||
</subSection>
|
||
<subSection id="E2FAAC9FFFA5D520FF39F95AFEF06E73" pageId="9" pageNumber="302" type="multiple">
|
||
<paragraph id="8BCAB574FFA5D520FF39F95AFE2C6FD4" blockId="9.[132,410,1714,1740]" box="[132,410,1714,1740]" pageId="9" pageNumber="302">
|
||
<heading id="D0820218FFA5D520FF39F95AFE2C6FD4" box="[132,410,1714,1740]" fontSize="11" level="2" pageId="9" pageNumber="302" reason="8">
|
||
<emphasis id="B9016966FFA5D520FF39F95AFE2C6FD4" box="[132,410,1714,1740]" italics="true" pageId="9" pageNumber="302">Two prominent appendages</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA5D520FF39F93BFEF06E73" blockId="9.[130,776,1746,1900]" pageId="9" pageNumber="302">
|
||
Protruding from under the shield (
|
||
<figureCitation id="134EA9F1FFA5D520FE4CF93BFDDE6FF4" box="[497,616,1746,1772]" captionStart="FIG" captionStartId="10.[132,143,1542,1559]" captionTargetBox="[133,1455,215,1499]" captionTargetId="figure-214@10.[698,1267,764,1317]" captionTargetPageId="10" captionText="FIG. 6. — Eodollocaris keithflinti Laville,J.T. Haug & C. Haug, n. gen.,n. sp., ROMIP61586: A, B, general view of part and counter-part;C, line drawing of A; D, line drawing of B; E, close-up of the dorsal midline of A; F, close-up of ventral margin of A; G, close-up of prominent appendages of A; H, close-up of prominent appendages of B. Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline; mv, ventral margin; r, rostrum; s, shield; st, stalk; t, trunk; ta, trunk appendage;?, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="9" pageNumber="302">Figs 6G, H</figureCitation>
|
||
;
|
||
<figureCitation id="134EA9F1FFA5D520FDCFF93BFD0F6FF4" box="[626,697,1746,1772]" captionStart="FIG" captionStartId="11.[132,143,1262,1279]" captionTargetBox="[133,1455,215,1219]" captionTargetId="figure-341@11.[699,1268,213,763]" captionTargetPageId="11" captionText="FIG. 7. — Eodollocaris keithflinti Laville, J.T. Haug & C. Haug, n. gen., n. sp., ROMIP61586: A, B, general view of part and counter-part (color-marked); C, closeup of appendages of A (color-marked); D, close-up of appendages of B (color-marked). Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline;mv, ventral margin;r, rostrum; s, shield;st, stalk;t, trunk;ta, trunk appendage;?, indeterminate oval plate;1-8, spines 1-8. Photographs: C. Haug and J. T. Haug.. Scale bars: A, B, 5 mm; C, D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746556" httpUri="https://zenodo.org/record/4746556/files/figure.png" pageId="9" pageNumber="302">7C, D</figureCitation>
|
||
), interpreted as raptorial appendages 2 and 3 (possibly appendages of post-ocular segments 5 and 6 = maxillae? and maxillipeds?). Proximal parts concealed by a tube-shaped structure, probably a part of the body.
|
||
</paragraph>
|
||
</subSection>
|
||
<subSection id="E2FAAC9FFFA5D520FF39F878FA8468E8" pageId="9" pageNumber="302" type="multiple">
|
||
<paragraph id="8BCAB574FFA5D520FF39F878FEDB6EB3" blockId="9.[132,365,1937,1963]" box="[132,365,1937,1963]" pageId="9" pageNumber="302">
|
||
<heading id="D0820218FFA5D520FF39F878FEDB6EB3" box="[132,365,1937,1963]" fontSize="11" level="2" pageId="9" pageNumber="302" reason="8">
|
||
<emphasis id="B9016966FFA5D520FF39F878FEDB6EB3" box="[132,365,1937,1963]" italics="true" pageId="9" pageNumber="302">Raptorial appendage 2</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA5D520FF39F858FCBE6EF4" blockId="9.[132,776,1969,2028]" pageId="9" pageNumber="302">
|
||
Differentiated into four elements. Element 1 rectangular in lateral view, longer than wide, 1.2× (l =
|
||
<quantity id="4C8D1891FFA5D520FDD5F838FD676EF4" box="[616,721,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="9" pageNumber="302" unit="mm" value="1.53">1.53 mm</quantity>
|
||
; w =
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA5D520FC93FF3EFA8468E8" blockId="9.[811,1457,215,497]" pageId="9" pageNumber="302">
|
||
<quantity id="4C8D1891FFA5D520FC93FF3EFC2E69EA" box="[814,920,215,242]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="9" pageNumber="302" unit="mm" value="1.33">1.33 mm</quantity>
|
||
). Element 2 rectangular, longer than wide, about 2.1× (l =
|
||
<quantity id="4C8D1891FFA5D520FC29FF1EFC4A680A" box="[916,1020,247,274]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="302" unit="mm" value="1.85">1.85 mm</quantity>
|
||
; w =
|
||
<quantity id="4C8D1891FFA5D520FB84FF1EFB14680A" box="[1081,1186,247,274]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="302" unit="mm" value="0.87">0.87 mm</quantity>
|
||
), angled in the specimen at 51° downward from the first article. Element 3 not wellpreserved.Width appears similar to that of preceding element. Element 4 longest (
|
||
<figureCitation id="134EA9F1FFA5D520FC44FEBEFBFA6869" box="[1017,1100,343,369]" captionStart="FIG" captionStartId="10.[132,143,1542,1559]" captionTargetBox="[133,1455,215,1499]" captionTargetId="figure-214@10.[698,1267,764,1317]" captionTargetPageId="10" captionText="FIG. 6. — Eodollocaris keithflinti Laville,J.T. Haug & C. Haug, n. gen.,n. sp., ROMIP61586: A, B, general view of part and counter-part;C, line drawing of A; D, line drawing of B; E, close-up of the dorsal midline of A; F, close-up of ventral margin of A; G, close-up of prominent appendages of A; H, close-up of prominent appendages of B. Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline; mv, ventral margin; r, rostrum; s, shield; st, stalk; t, trunk; ta, trunk appendage;?, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="9" pageNumber="302">Fig. 6G</figureCitation>
|
||
), thin, longer than wide, 5.6× (l =
|
||
<quantity id="4C8D1891FFA5D520FC96FE9EFC266889" box="[811,912,375,401]" metricMagnitude="-3" metricUnit="m" metricValue="3.92" pageId="9" pageNumber="302" unit="mm" value="3.92">3.92 mm</quantity>
|
||
; w =
|
||
<quantity id="4C8D1891FFA5D520FC78FE9EFB9A6889" box="[965,1068,375,401]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="302" unit="mm" value="0.7">0.70 mm</quantity>
|
||
), angled at 50° inward from element 3. Distally bearing two spines on its median margin. First spine smaller (l =
|
||
<quantity id="4C8D1891FFA5D520FC51FE5EFBE268C9" box="[1004,1108,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="9" pageNumber="302" unit="mm" value="0.79">0.79 mm</quantity>
|
||
). Second spine very distal. Spine longer than the first one and directed anteriorly.
|
||
</paragraph>
|
||
</subSection>
|
||
<subSection id="E2FAAC9FFFA5D520FC90FDFFFA2A6DF6" pageId="9" pageNumber="302" type="multiple">
|
||
<paragraph id="8BCAB574FFA5D520FC90FDFFFBA36B28" blockId="9.[813,1045,534,560]" box="[813,1045,534,560]" pageId="9" pageNumber="302">
|
||
<heading id="D0820218FFA5D520FC90FDFFFBA36B28" box="[813,1045,534,560]" fontSize="11" level="2" pageId="9" pageNumber="302" reason="8">
|
||
<emphasis id="B9016966FFA5D520FC90FDFFFBA36B28" box="[813,1045,534,560]" italics="true" pageId="9" pageNumber="302">Raptorial appendage 3</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA5D520FC90FDDEFA2A6DF6" blockId="9.[811,1457,566,1262]" pageId="9" pageNumber="302">
|
||
Better preserved, longer than raptorial appendage 2. Differentiated into four elements (
|
||
<figureCitation id="134EA9F1FFA5D520FBDCFDBFFB6F6B68" box="[1121,1241,598,624]" captionStart="FIG" captionStartId="11.[132,143,1262,1279]" captionTargetBox="[133,1455,215,1219]" captionTargetId="figure-341@11.[699,1268,213,763]" captionTargetPageId="11" captionText="FIG. 7. — Eodollocaris keithflinti Laville, J.T. Haug & C. Haug, n. gen., n. sp., ROMIP61586: A, B, general view of part and counter-part (color-marked); C, closeup of appendages of A (color-marked); D, close-up of appendages of B (color-marked). Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline;mv, ventral margin;r, rostrum; s, shield;st, stalk;t, trunk;ta, trunk appendage;?, indeterminate oval plate;1-8, spines 1-8. Photographs: C. Haug and J. T. Haug.. Scale bars: A, B, 5 mm; C, D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746556" httpUri="https://zenodo.org/record/4746556/files/figure.png" pageId="9" pageNumber="302">Fig. 7C, D</figureCitation>
|
||
). Element 1 rectangular, almost square-shaped (l =
|
||
<quantity id="4C8D1891FFA5D520FB2FFD9FFB4C6B88" box="[1170,1274,630,656]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="9" pageNumber="302" unit="mm" value="2.13">2.13 mm</quantity>
|
||
; w =
|
||
<quantity id="4C8D1891FFA5D520FA84FD9FFA156B88" box="[1337,1443,630,656]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="9" pageNumber="302" unit="mm" value="2.03">2.03 mm</quantity>
|
||
); bearing a stout spine in the central part of the lateral margin (l =
|
||
<quantity id="4C8D1891FFA5D520FCE3FD5FFC7A6BC8" box="[862,972,694,720]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="302" unit="mm" value="0.38">0.38 mm</quantity>
|
||
). Element 2 hexagonal in outline, slightly rounded, wider in its central part than at its extremities (
|
||
<subScript id="17F1B731FFA5D520FC8BFD1CFCDE6A0A" attach="left" box="[822,872,757,786]" fontSize="7" pageId="9" pageNumber="302">wmax</subScript>
|
||
=
|
||
<quantity id="4C8D1891FFA5D520FC38FD1CFC5B6A08" box="[901,1005,757,784]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" pageId="9" pageNumber="302" unit="mm" value="2.19">2.19 mm</quantity>
|
||
;
|
||
<subScript id="17F1B731FFA5D520FC44FD1FFB9C6A0B" attach="left" box="[1017,1066,758,787]" fontSize="7" pageId="9" pageNumber="302">wmin</subScript>
|
||
=
|
||
<quantity id="4C8D1891FFA5D520FBF5FD1CFB196A08" box="[1096,1199,757,784]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="302" unit="mm" value="1.05">1.05 mm</quantity>
|
||
; l =
|
||
<quantity id="4C8D1891FFA5D520FB5DFD1CFAFE6A08" box="[1248,1352,757,784]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="9" pageNumber="302" unit="mm" value="1.79">1.79 mm</quantity>
|
||
). Bearing a spine on central part of lateral and median margin. Lateral spine longer (l =
|
||
<quantity id="4C8D1891FFA5D520FC5DFCDCFBF36A48" box="[992,1093,821,848]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="302" unit="mm" value="0.28">0.28 mm</quantity>
|
||
) than median one (l =
|
||
<quantity id="4C8D1891FFA5D520FA86FCDCFA146A48" box="[1339,1442,821,848]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="302" unit="mm" value="0.16">0.16 mm</quantity>
|
||
). Element 3 rectangular, longer than wide, about 3.6× (l =
|
||
<quantity id="4C8D1891FFA5D520FC92FC9CFC216A97" box="[815,919,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="4.27" pageId="9" pageNumber="302" unit="mm" value="4.27">4.27 mm</quantity>
|
||
; w =
|
||
<quantity id="4C8D1891FFA5D520FC68FC9CFB896A97" box="[981,1087,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="9" pageNumber="302" unit="mm" value="1.18">1.18 mm</quantity>
|
||
), angled at 24.3° from element 2. One spine on median margin (l =
|
||
<quantity id="4C8D1891FFA5D520FB09FC7CFA966AB7" box="[1204,1312,917,943]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="302" unit="mm" value="0.37">0.37 mm</quantity>
|
||
). Element 4, distal one, also the longest one (l =
|
||
<quantity id="4C8D1891FFA5D520FB7BFC5CFA856AD7" box="[1222,1331,949,975]" metricMagnitude="-3" metricUnit="m" metricValue="6.32" pageId="9" pageNumber="302" unit="mm" value="6.32">6.32 mm</quantity>
|
||
); rectangular, thin and elongated, longer than wide, about 7.2× (w =
|
||
<quantity id="4C8D1891FFA5D520FC93FC1CFC206D17" box="[814,918,1013,1039]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="9" pageNumber="302" unit="mm" value="0.88">0.88 mm</quantity>
|
||
), angled at 56.6° anteriorly from element 3. Bears seven spines on median margin (
|
||
<figureCitation id="134EA9F1FFA5D520FB39FBFDFB4E6D37" box="[1156,1272,1044,1071]" captionStart="FIG" captionStartId="11.[132,143,1262,1279]" captionTargetBox="[133,1455,215,1219]" captionTargetId="figure-341@11.[699,1268,213,763]" captionTargetPageId="11" captionText="FIG. 7. — Eodollocaris keithflinti Laville, J.T. Haug & C. Haug, n. gen., n. sp., ROMIP61586: A, B, general view of part and counter-part (color-marked); C, closeup of appendages of A (color-marked); D, close-up of appendages of B (color-marked). Abbreviations: a2, a3, second, third raptorial appendages; e, eyes; e1-e4, element 1-4; dm, dorsal midline;mv, ventral margin;r, rostrum; s, shield;st, stalk;t, trunk;ta, trunk appendage;?, indeterminate oval plate;1-8, spines 1-8. Photographs: C. Haug and J. T. Haug.. Scale bars: A, B, 5 mm; C, D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746556" httpUri="https://zenodo.org/record/4746556/files/figure.png" pageId="9" pageNumber="302">Fig. 7C, D</figureCitation>
|
||
). First one is thin (l =
|
||
<quantity id="4C8D1891FFA5D520FCE9FBDDFC0C6D57" box="[852,954,1076,1103]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="302" unit="mm" value="0.65">0.65 mm</quantity>
|
||
); second one also thin (l =
|
||
<quantity id="4C8D1891FFA5D520FB6FFBDDFA8E6D57" box="[1234,1336,1076,1103]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="9" pageNumber="302" unit="mm" value="0.58">0.58 mm</quantity>
|
||
), third also thin, longest of the series (l =
|
||
<quantity id="4C8D1891FFA5D520FBDCFBBDFB7E6D77" box="[1121,1224,1108,1135]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="302" unit="mm" value="0.68">0.68 mm</quantity>
|
||
); fourth spine shorter (l =
|
||
<quantity id="4C8D1891FFA5D520FCE7FB9DFC726D97" box="[858,964,1140,1167]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="302" unit="mm" value="0.44">0.44 mm</quantity>
|
||
); fifth similar to fourth (l =
|
||
<quantity id="4C8D1891FFA5D520FB41FB9DFAD06D97" box="[1276,1382,1140,1167]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="302" unit="mm" value="0.45">0.45 mm</quantity>
|
||
); sixth spine shortest (l =
|
||
<quantity id="4C8D1891FFA5D520FC4DFB7DFBE16DB6" box="[1008,1111,1172,1198]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="302" unit="mm" value="0.2">0.20 mm</quantity>
|
||
), close to seventh spine. Seventh spine corresponds to distal end of appendage; slightly curved, longer than the previous spine, about 3.2× (l =
|
||
<quantity id="4C8D1891FFA5D520FA95FB3DFA266DF6" box="[1320,1424,1236,1262]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="9" pageNumber="302" unit="mm" value="0.64">0.64 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSection>
|
||
<subSection id="E2FAAC9FFFA5D523FC90FAFDFBC36EF4" lastPageId="10" lastPageNumber="303" pageId="9" pageNumber="302" type="multiple">
|
||
<paragraph id="8BCAB574FFA5D520FC90FAFDFBA46C35" blockId="9.[813,1042,1299,1326]" box="[813,1042,1299,1326]" pageId="9" pageNumber="302">
|
||
<heading id="D0820218FFA5D520FC90FAFDFBA46C35" box="[813,1042,1299,1326]" fontSize="11" level="2" pageId="9" pageNumber="302" reason="8">
|
||
<emphasis id="B9016966FFA5D520FC90FAFDFBA46C35" box="[813,1042,1299,1326]" italics="true" pageId="9" pageNumber="302">Posterior part of trunk</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA5D520FC90FADAFAFE6CB5" blockId="9.[813,1458,1331,1453]" pageId="9" pageNumber="302">Only the posterior part of trunk is visible. Carrying three appendages. First posterior trunk appendage paddle-like. Second one not well preserved but bearing a spine distally. Third, also paddle-like, arising from end of trunk.</paragraph>
|
||
<subSubSection id="C36FE6FFFFA5D523FC90FA3AFBC36EF4" lastPageId="10" lastPageNumber="303" pageId="9" pageNumber="302" type="discussion">
|
||
<paragraph id="8BCAB574FFA5D520FC90FA3AFC276CF3" blockId="9.[810,1457,1491,2027]" box="[813,913,1491,1517]" pageId="9" pageNumber="302">REMARKS</paragraph>
|
||
<paragraph id="8BCAB574FFA5D520FC90FA1AFBB36FD4" blockId="9.[810,1457,1491,2027]" pageId="9" pageNumber="302">
|
||
The new specimen possesses a so far unique combination of characters. The rostrum with a rounded tip seems to be a common feature in Palaeozoic species of
|
||
<taxonomicName id="4C75CEF7FFA5D520FAB9F9DBFA1B6F54" authorityName="Pinna, Arduini, Pesarini & Teruzzi" authorityYear="1982" box="[1284,1453,1586,1612]" class="Thylacocephala" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="302" phylum="Arthropoda" rank="class">Thylacocephala</taxonomicName>
|
||
, especially in species of
|
||
<taxonomicName id="4C75CEF7FFA5D520FBABF9BAFAA96F74" authority="Rolfe, 1961" authorityName="Rolfe" authorityYear="1961" box="[1046,1311,1618,1645]" class="Thylacocephala" family="Apidae" genus="Concavicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FBABF9BAFB2E6F75" box="[1046,1176,1619,1645]" italics="true" pageId="9" pageNumber="302">Concavicaris</emphasis>
|
||
<bibRefCitation id="EFE4C885FFA5D520FB22F9BBFAA96F74" author="ROLFE W. D. I." box="[1183,1311,1618,1644]" pageId="9" pageNumber="302" pagination="1243" refId="ref11445" refString="ROLFE W. D. I. 1961. - Concavicaris and Quasicaris, substitute names for Colpocaris Meek, 1872, and Pterocaris Barrande, 1872. Journal of Paleontology 35 (6): 1243." type="journal article" year="1961">Rolfe, 1961</bibRefCitation>
|
||
</taxonomicName>
|
||
. The presence of such a rostrum has also been suggested to be a diagnostic feature of representatives of the group
|
||
<taxonomicName id="4C75CEF7FFA5D520FB13F97BFCD46FD4" authority="Arduini, 1990" authorityName="Arduini" authorityYear="1990" class="Thylacocephala" genus="Ankitokazocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
Ankitokazocaris
|
||
<bibRefCitation id="EFE4C885FFA5D520FAE5F97BFCD46FD4" author="ARDUINI P." pageId="9" pageNumber="302" pagination="197 - 204" refId="ref8495" refString="ARDUINI P. 1990. - Thylacocephala from Lower Trias of Madagascar. Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 131: 197 - 204" type="journal article" year="1990">Arduini, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
of the Triassic.
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA5D523FCFEF93BFE2D6FF4" blockId="9.[810,1457,1491,2027]" lastBlockId="10.[130,777,1714,2027]" lastPageId="10" lastPageNumber="303" pageId="9" pageNumber="302">
|
||
However, it differs from known representatives of both groups,
|
||
<taxonomicName id="4C75CEF7FFA5D520FC39F91BFBBE6E14" authorityName="Rolfe" authorityYear="1961" box="[900,1032,1778,1804]" class="Thylacocephala" family="Apidae" genus="Concavicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FC39F91BFBBE6E14" box="[900,1032,1778,1804]" italics="true" pageId="9" pageNumber="302">Concavicaris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C75CEF7FFA5D520FBFCF91BFB526E14" authorityName="Arduini" authorityYear="1990" box="[1089,1252,1778,1804]" class="Thylacocephala" genus="Ankitokazocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FBFCF91BFB526E14" box="[1089,1252,1778,1804]" italics="true" pageId="9" pageNumber="302">Ankitokazocaris</emphasis>
|
||
</taxonomicName>
|
||
, by the absence of a well-developed optic notch. Also, the new specimen possesses a posterior spine and a posterior notch, both structures are absent in species of
|
||
<taxonomicName id="4C75CEF7FFA5D520FBA2F8BBFB156E74" authorityName="Rolfe" authorityYear="1961" box="[1055,1187,1874,1900]" class="Thylacocephala" family="Apidae" genus="Concavicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FBA2F8BBFB156E74" box="[1055,1187,1874,1900]" italics="true" pageId="9" pageNumber="302">Concavicaris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C75CEF7FFA5D520FB68F8B8FACF6E73" authorityName="Arduini" authorityYear="1990" box="[1237,1401,1873,1899]" class="Thylacocephala" genus="Ankitokazocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FB68F8B8FACF6E73" box="[1237,1401,1873,1899]" italics="true" pageId="9" pageNumber="302">Ankitokazocaris</emphasis>
|
||
</taxonomicName>
|
||
. Yet, they are well known in specimens from younger deposits, such as representatives of
|
||
<taxonomicName id="4C75CEF7FFA5D520FB8BF87BFA1D6EB3" authority="Arduini & Brasca, 1984" authorityName="Arduini & Brasca" authorityYear="1984" box="[1078,1451,1937,1964]" class="Conchyliocarida" family="Microcarididae" genus="Atropicaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FB8BF87BFB136EB4" box="[1078,1189,1938,1964]" italics="true" pageId="9" pageNumber="302">Atropicaris</emphasis>
|
||
<bibRefCitation id="EFE4C885FFA5D520FB11F878FA1D6EB3" author="ARDUINI P. & BRASCA A." box="[1196,1451,1937,1964]" pageId="9" pageNumber="302" pagination="87 - 93" refId="ref8649" refString="ARDUINI P. & BRASCA A. 1984. - Atropicaris: nuovo genere della classe Thylacocephala. Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 125: 87 - 93" type="journal article" year="1984">Arduini & Brasca, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C75CEF7FFA5D520FC96F858FBD86ED3" authority="Van Straelen, 1923" authorityName="Van Straelen" authorityYear="1923" box="[811,1134,1969,1996]" class="Thylacocephala" family="Apidae" genus="Dollocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FC96F858FC256ED3" box="[811,915,1969,1995]" italics="true" pageId="9" pageNumber="302">Dollocaris</emphasis>
|
||
<bibRefCitation id="EFE4C885FFA5D520FC20F85BFBD86ED3" author="VAN STRAELEN V." box="[925,1134,1969,1996]" pageId="9" pageNumber="302" pagination="431 - 439" refId="ref12206" refString="VAN STRAELEN V. 1923. - Les mysidaces du Callovien de la Voultesur-Rhone. Bulletin de la Societe geologique de France 23: 431 - 439" type="journal article" year="1923">Van Straelen, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C75CEF7FFA5D520FB24F85BFA3D6ED3" authority="Polz, 1994" authorityName="Polz" authorityYear="1994" box="[1177,1419,1969,1995]" class="Conchyliocarida" family="Dollocarididae" genus="Mayrocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA5D520FB24F85BFABA6ED3" box="[1177,1292,1970,1995]" italics="true" pageId="9" pageNumber="302">Mayrocaris</emphasis>
|
||
<bibRefCitation id="EFE4C885FFA5D520FAAFF858FA3D6ED3" author="POLZ H." box="[1298,1419,1969,1995]" pageId="9" pageNumber="302" pagination="35 - 44" refId="ref11284" refString="POLZ H. 1994. - Mayrocaris bucculata gen. nov. sp. nov. (Thylacocephala, Conchyliocarida) aus den Solnhofener Plattenkalken. Archaeopteryx 12: 35 - 44" type="journal article" year="1994">Polz, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
. In these species, the posterior notch occupies the entire posterior margin whereas in the new specimen it is restricted to the dorsal part of the margin.
|
||
</paragraph>
|
||
<caption id="DF0AE5FCFFA6D523FF39F9EFFEFC6F69" ID-DOI="http://doi.org/10.5281/zenodo.4746554" ID-Zenodo-Dep="4746554" httpUri="https://zenodo.org/record/4746554/files/figure.png" pageId="10" pageNumber="303" startId="10.[132,143,1542,1559]" targetBox="[133,1455,215,1499]" targetPageId="10">
|
||
<paragraph id="8BCAB574FFA6D523FF39F9EFFEFC6F69" blockId="10.[132,1457,1542,1649]" pageId="10" pageNumber="303">
|
||
FIG. 6. —
|
||
<taxonomicName id="4C75CEF7FFA6D523FF6FF9EFFDC56F0F" authority="Laville,J.T. Haug & C. Haug" authorityName="Laville, J. T. Haug & C. Haug" authorityYear="2021" box="[210,627,1542,1559]" genus="Eodollocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="keithflinti" status="n. gen.,n. sp.">
|
||
<emphasis id="B9016966FFA6D523FF6FF9EFFE316F0F" box="[210,391,1542,1559]" italics="true" pageId="10" pageNumber="303">Eodollocaris keithflinti</emphasis>
|
||
Laville,J.T. Haug & C. Haug
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A232D41DFFA6D523FDC4F9EFFD5A6F0F" box="[633,748,1542,1559]" pageId="10" pageNumber="303" rank="species">n. gen.,n. sp.</taxonomicNameLabel>
|
||
, ROMIP61586:
|
||
<emphasis id="B9016966FFA6D523FCD6F9EFFCCE6F0F" bold="true" box="[875,888,1542,1559]" pageId="10" pageNumber="303">A</emphasis>
|
||
,
|
||
<emphasis id="B9016966FFA6D523FC3CF9EFFC386F0F" bold="true" box="[897,910,1542,1559]" pageId="10" pageNumber="303">B</emphasis>
|
||
, general view of part and counter-part;
|
||
<emphasis id="B9016966FFA6D523FB6FF9EFFB566F0F" bold="true" box="[1234,1248,1542,1559]" pageId="10" pageNumber="303">C</emphasis>
|
||
, line drawing of A;
|
||
<emphasis id="B9016966FFA6D523FAC6F9EFFA3F6F0F" bold="true" box="[1403,1417,1542,1559]" pageId="10" pageNumber="303">D</emphasis>
|
||
, line drawing of B;
|
||
<emphasis id="B9016966FFA6D523FF46F9F5FEB16F35" bold="true" box="[251,263,1564,1581]" pageId="10" pageNumber="303">E</emphasis>
|
||
, close-up of the dorsal midline of A;
|
||
<emphasis id="B9016966FFA6D523FDFCF9F5FDFA6F35" bold="true" box="[577,588,1564,1581]" pageId="10" pageNumber="303">F</emphasis>
|
||
, close-up of ventral margin of A;
|
||
<emphasis id="B9016966FFA6D523FCDAF9F5FCC36F35" bold="true" box="[871,885,1564,1581]" pageId="10" pageNumber="303">G</emphasis>
|
||
, close-up of prominent appendages of A;
|
||
<emphasis id="B9016966FFA6D523FB61F9F5FB5C6F35" bold="true" box="[1244,1258,1564,1581]" pageId="10" pageNumber="303">H</emphasis>
|
||
, close-up of prominent appendages of B. Abbreviations:
|
||
<emphasis id="B9016966FFA6D523FE23F9DAFE056F5C" bold="true" box="[414,435,1587,1604]" pageId="10" pageNumber="303">a2</emphasis>
|
||
,
|
||
<emphasis id="B9016966FFA6D523FE02F9DAFE656F5C" bold="true" box="[447,467,1587,1604]" pageId="10" pageNumber="303">a3</emphasis>
|
||
, second, third raptorial appendages;
|
||
<emphasis id="B9016966FFA6D523FCACF9DAFCAA6F5C" bold="true" box="[785,796,1587,1604]" pageId="10" pageNumber="303">e</emphasis>
|
||
, eyes;
|
||
<emphasis id="B9016966FFA6D523FCE4F9DAFC3C6F5C" bold="true" box="[857,906,1587,1604]" pageId="10" pageNumber="303">e1-e4</emphasis>
|
||
, element 1-4;
|
||
<emphasis id="B9016966FFA6D523FBBBF9DAFB946F5C" bold="true" box="[1030,1058,1587,1604]" pageId="10" pageNumber="303">dm</emphasis>
|
||
, dorsal midline;
|
||
<emphasis id="B9016966FFA6D523FB13F9DAFB7F6F5C" bold="true" box="[1198,1225,1587,1604]" pageId="10" pageNumber="303">mv</emphasis>
|
||
, ventral margin;
|
||
<emphasis id="B9016966FFA6D523FAEAF9DAFAE86F5C" bold="true" box="[1367,1374,1587,1604]" pageId="10" pageNumber="303">r</emphasis>
|
||
, rostrum;
|
||
<emphasis id="B9016966FFA6D523FF39F9A3FF386F43" bold="true" box="[132,142,1610,1627]" pageId="10" pageNumber="303">s</emphasis>
|
||
, shield;
|
||
<emphasis id="B9016966FFA6D523FF68F9A3FF506F43" bold="true" box="[213,230,1610,1627]" pageId="10" pageNumber="303">st</emphasis>
|
||
, stalk;
|
||
<emphasis id="B9016966FFA6D523FE9FF9A3FE9F6F43" bold="true" box="[290,297,1610,1627]" pageId="10" pageNumber="303">t</emphasis>
|
||
, trunk;
|
||
<emphasis id="B9016966FFA6D523FED5F9A3FECF6F43" bold="true" box="[360,377,1610,1627]" pageId="10" pageNumber="303">ta</emphasis>
|
||
, trunk appendage;
|
||
<emphasis id="B9016966FFA6D523FDA0F9A3FD916F43" bold="true" box="[541,551,1610,1627]" pageId="10" pageNumber="303">?</emphasis>
|
||
, indeterminate oval plate. Photographs: C. Haug and J. T. Haug. Line drawings: Thomas Laville. Scale bars: A-D, 5 mm; E-H, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCAB574FFA6D523FF26F91BFB006FF4" blockId="10.[130,777,1714,2027]" lastBlockId="10.[812,1457,1714,2028]" pageId="10" pageNumber="303">
|
||
Among the species from younger deposits, especially
|
||
<emphasis id="B9016966FFA6D523FD70F91AFE8A6E33" italics="true" pageId="10" pageNumber="303">Paradollocaris vannieri</emphasis>
|
||
<bibRefCitation id="EFE4C885FFA6D523FEFFF8FBFDBD6E34" author="CHARBONNIER S. & TERUZZI G. & AUDO D. & LASSERON M. & HAUG C. & HAUG J. T." box="[322,523,1810,1836]" pageId="10" pageNumber="303" pagination="19" refId="ref9596" refString="CHARBONNIER S., TERUZZI G., AUDO D., LASSERON M., HAUG C. & HAUG J. T. 2017. - New thylacocephalans from the Cretaceous Lagerstatten of Lebanon. Bulletin de la Societe geologique de France 188 (3): 19. https: // doi. org / 10.1051 / bsgf / 2017176" type="journal article" year="2017">Charbonnier, 2017</bibRefCitation>
|
||
shows many similarities concerning shield shape with the new specimen. The shield is trapezoidal with a convex dorsal midline. Anteriorly it is drawn out into a rostrum, which is rounded distally. The posterior end is also drawn out into a spine. The posterior margin also has a pronounced notch. Similar to other species from the Mesozoic, the notch occupies the entire posterior margin. The ventral margin of the new specimen seems unique with the anterior part being tuberculated.
|
||
</paragraph>
|
||
<paragraph id="8BCAB574FFA6D523FCFEF91BFBC36EF4" blockId="10.[812,1457,1714,2028]" pageId="10" pageNumber="303">
|
||
Concerning the raptorial appendages, the new specimen exhibits particular differences in form and armature compared to other thylacocephalans from Mazon Creek.
|
||
<taxonomicName id="4C75CEF7FFA6D523FAEFF8DBFC656E73" authorityName="Schram" authorityYear="1990" class="Conchyliocarida" genus="Convexicaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="mazonensis">
|
||
<emphasis id="B9016966FFA6D523FAEFF8DBFC656E73" italics="true" pageId="10" pageNumber="303">Convexicaris mazonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C75CEF7FFA6D523FBB5F8BBFAB56E73" authorityName="Schram" authorityYear="1990" box="[1032,1283,1874,1900]" class="Thylacocephala" family="Apidae" genus="Concavicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Concavicarida" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="georgeorum">
|
||
<emphasis id="B9016966FFA6D523FBB5F8BBFAB56E73" box="[1032,1283,1874,1900]" italics="true" pageId="10" pageNumber="303">Concavicaris georgeorum</emphasis>
|
||
</taxonomicName>
|
||
have less robust appendages, and they do not display any spines. Due to the significant differences of the new specimen we interpret it as a representative of a new, so far unrecognised species:
|
||
<taxonomicName id="4C75CEF7FFA6D523FAD6F858FC416EF4" authority="keithflinti n." authorityName="KEITHFLINTI N." genus="Eodollocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="304" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9016966FFA6D523FAD6F858FC6D6EF3" italics="true" pageId="10" pageNumber="303">Eodollocaris keithflinti</emphasis>
|
||
<taxonomicNameLabel id="A232D41DFFA6D523FC5FF83BFC416EF4" box="[994,1015,2002,2028]" pageId="10" pageNumber="303" rank="species">n.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
gen., n. sp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |