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<document id="BE3F95B63253D30BCD7364AE5EA74938" ID-CLB-Dataset="39302" ID-DOI="10.11646/zootaxa.4057.1.10" ID-GBIF-Dataset="a9c2ada1-02a9-4f30-b82b-211faa62748e" ID-ISSN="1175-5326" ID-Zenodo-Dep="238911" ID-ZooBank="BCC393E4-ACA9-40FD-99EA-774604D0EBC9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461354831214" checkinUser="plazi" docAuthor="Häckel, Martin &amp; Anichtchenko, Alexandr" docDate="2015" docId="03BD7017FFE1506EFF7B8243FBA2EFE5" docLanguage="en" docName="zt04057p150.pdf" docOrigin="Zootaxa 4057 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudotetracha timberensis Häckel &amp; Anichtchenko, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="144" masterDocId="FF84086FFFE0506FFFEC8102FFF1E83C" masterDocTitle="Pseudotetracha timberensis: a new species from the Northern Territory, Australia (Coleoptera: Carabidae, Cicindelinae, Megacephalini) and considerations on the taxonomy of Pseudotetracha" masterLastPageNumber="150" masterPageNumber="143" pageNumber="144" updateTime="1698661470441" updateUser="ExternalLinkService">
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<mods:title id="5D31E4CE553F8C2208CD3196DA481EA5">Pseudotetracha timberensis: a new species from the Northern Territory, Australia (Coleoptera: Carabidae, Cicindelinae, Megacephalini) and considerations on the taxonomy of Pseudotetracha</mods:title>
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<mods:namePart id="5EB458EBC3ADDC313EE6D0CA4806BD93">Häckel, Martin</mods:namePart>
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<mods:namePart id="C204E3C867F3FE44ED81F9D1CC97C567">Anichtchenko, Alexandr</mods:namePart>
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<mods:date id="69AEC42E5AA7A8E23180C97515E0DE84">2015</mods:date>
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<treatment id="03BD7017FFE1506EFF7B8243FBA2EFE5" ID-DOI="http://doi.org/10.5281/zenodo.6109387" ID-GBIF-Taxon="119647207" ID-Zenodo-Dep="6109387" LSID="urn:lsid:plazi:treatment:03BD7017FFE1506EFF7B8243FBA2EFE5" httpUri="http://treatment.plazi.org/id/03BD7017FFE1506EFF7B8243FBA2EFE5" lastPageNumber="144" pageId="1" pageNumber="144">
<subSubSection id="C30E928AFFE1506EFF7B8243FDB8EB67" box="[151,585,833,859]" pageId="1" pageNumber="144" type="nomenclature">
<paragraph id="8BABC101FFE1506EFF7B8243FDB8EB67" blockId="1.[151,585,833,859]" box="[151,585,833,859]" pageId="1" pageNumber="144">
<heading id="D0E3766DFFE1506EFF7B8243FDB8EB67" bold="true" box="[151,585,833,859]" fontSize="11" level="1" pageId="1" pageNumber="144" reason="1">
<emphasis id="B9601D13FFE1506EFF7B8243FDB8EB67" bold="true" box="[151,585,833,859]" pageId="1" pageNumber="144">
<taxonomicName id="4C14BA82FFE1506EFF7B8243FE12EB67" ID-CoL="8MJLG" box="[151,483,833,859]" class="Insecta" family="Carabidae" genus="Pseudotetracha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="timberensis" status="sp. nov.">
<emphasis id="B9601D13FFE1506EFF7B8243FE12EB67" bold="true" box="[151,483,833,859]" italics="true" pageId="1" pageNumber="144">Pseudotetracha timberensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253A068FFE1506EFE078243FDB8EB67" box="[491,585,833,859]" pageId="1" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30E928AFFE1506EFF7B8285FC6AEC30" pageId="1" pageNumber="144" type="materials_examined">
<paragraph id="8BABC101FFE1506EFF7B8285FC6AEC30" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">
<emphasis id="B9601D13FFE1506EFF7B8285FEBBEB9C" bold="true" box="[151,330,903,928]" pageId="1" pageNumber="144">
<typeStatus id="54AF7FA3FFE1506EFF7B8285FF23EB9C" box="[151,210,903,928]" pageId="1" pageNumber="144">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54AF7FA3FFE1506EFEBB8285FE4EEB9C" box="[343,447,903,928]" pageId="1" pageNumber="144" type="holotype">Holotype</typeStatus>
(♂): “
<collectingCountry id="F3038191FFE1506EFDFE8285FD8EEB9C" box="[530,639,903,928]" name="Australia" pageId="1" pageNumber="144">Australia</collectingCountry>
. Northern Territory,
<quantity id="4CEC6CE4FFE1506EFC99828AFC21EB9C" box="[885,976,903,928]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="144" unit="km" value="120.0">120 km</quantity>
w Timber Creek, Saddle Creek R. A.
<geoCoordinate id="EE20A7C6FFE1506EFF7B82AEFEE0EBF9" box="[151,273,940,965]" direction="south" orientation="latitude" pageId="1" pageNumber="144" precision="9" value="-15.954333">15°57.26'S</geoCoordinate>
<geoCoordinate id="EE20A7C6FFE1506EFEF582AEFE54EBF8" box="[281,421,940,964]" direction="east" orientation="longitude" pageId="1" pageNumber="144" precision="9" value="129.557">129°33.42'E</geoCoordinate>
, lgt. loc. collector (
<figureCitation id="132FDD84FFE1506EFD9382AEFD32EBF8" box="[639,707,940,965]" captionStart="FIGURE 2" captionStartId="2.[151,250,1933,1955]" captionTargetBox="[328,1259,1158,1905]" captionTargetId="figure@2.[321,1265,1149,1912]" captionTargetPageId="2" captionText="FIGURE 2 Pseudotetracha timberensis sp. nov. A—male, habitus; B—aedeagus with expanded endophallus left, upper and right views; C—aedeagus in left lateral view." httpUri="https://zenodo.org/record/238913/files/figure.png" pageId="1" pageNumber="144">Fig. 2</figureCitation>
, cMH, donated to
<collectionCode id="ED0559C4FFE1506EFC7882AEFBF1EBF8" LSID="urn:lsid:biocol.org:col:34986" box="[916,1024,940,964]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34986" name="Museum and Art Gallery of the Northern Territory" pageId="1" pageNumber="144">MAGNT</collectionCode>
).
<typeStatus id="54AF7FA3FFE1506EFBFE82AFFB75EBF9" box="[1042,1156,941,965]" pageId="1" pageNumber="144" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D120A88FFE1506EFB6282AEFB43EBF8" box="[1166,1202,940,964]" pageId="1" pageNumber="144" type="generic">1♂</specimenCount>
, 3♀, same data (
<figureCitation id="132FDD84FFE1506EFA8382AEFF56EBD4" captionStart="FIGURE 3" captionStartId="3.[151,250,1189,1211]" captionTargetBox="[341,1242,213,1148]" captionTargetId="figure@3.[327,1259,193,1167]" captionTargetPageId="3" captionText="FIGURE 3 Pseudotetracha timberensis sp. nov. A—female, habitus; B—proximal antennomeres, detail; C—elytral apex, detail." httpUri="https://zenodo.org/record/238914/files/figure.png" pageId="1" pageNumber="144">Fig. 3</figureCitation>
, cMH, cRS, cSJ);
<specimenCount id="9D120A88FFE1506EFE9782D2FE6EEBD4" box="[379,415,976,1000]" pageId="1" pageNumber="144" type="generic">1♂</specimenCount>
, 2♀, Western
<collectingCountry id="F3038191FFE1506EFDAE82D2FD5AEBD4" box="[578,683,976,1000]" name="Australia" pageId="1" pageNumber="144">Australia</collectingCountry>
, Kimberley Reg.Victoria Hwy, ~
<quantity id="4CEC6CE4FFE1506EFBCC82D2FB9EEBD4" box="[1056,1135,976,1000]" metricMagnitude="5" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="144" unit="km" value="102.0">102km</quantity>
N Halls Creek
<geoCoordinate id="EE20A7C6FFE1506EFACD82D2FA6AEBD4" box="[1313,1435,976,1000]" direction="south" orientation="latitude" pageId="1" pageNumber="144" precision="9" value="-17.426332">17°25.58'S</geoCoordinate>
<geoCoordinate id="EE20A7C6FFE1506EFF7B82F6FED0EC31" box="[151,289,1012,1037]" direction="east" orientation="longitude" pageId="1" pageNumber="144" precision="9" value="127.98617">127°59.17'E</geoCoordinate>
Spring Creek
<date id="FFAAE7C1FFE1506EFE2882F6FDD0EC30" box="[452,545,1012,1036]" pageId="1" pageNumber="144" value="2014-01">I—2014</date>
lgt. J. Horák, S. Jákl (cMH, cSJ).
</paragraph>
</subSubSection>
<subSubSection id="C30E928AFFE1506EFF2B8515FDD0EF51" pageId="1" pageNumber="144" type="description">
<paragraph id="8BABC101FFE1506EFF2B8515FD26EC68" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">
<emphasis id="B9601D13FFE1506EFF2B8515FEA8EC0C" bold="true" box="[199,345,1047,1072]" pageId="1" pageNumber="144">Description.</emphasis>
Length
<quantity id="4CEC6CE4FFE1506EFE548515FDAEEC13" box="[440,607,1047,1072]" metricMagnitude="-2" metricUnit="m" metricValue="1.635" metricValueMax="1.76" metricValueMin="1.51" pageId="1" pageNumber="144" unit="mm" value="16.35" valueMax="17.6" valueMin="15.1">15.117.6 mm</quantity>
, width
<quantity id="4CEC6CE4FFE1506EFD588515FCCEEC13" box="[692,831,1047,1072]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.8" metricValueMin="4.9" pageId="1" pageNumber="144" unit="mm" value="5.85" valueMax="6.8" valueMin="4.9">4.96.8 mm</quantity>
. Proportions: pronotum 1.07x wider than long, 0.93x wider than head, elytra 1,45x wider than pronotum.
</paragraph>
<paragraph id="8BABC101FFE1506EFF2B855DFCEDED3B" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">Coloration. Unicolorous; body black or dark brown with green and often also slight purple metallic sheen; antennae, palps and mandibles yellowish, proximal four antennomeres and mandibles with distal parts dark; labrum dark brown, in some specimens with yellowish base and lateral margins. Legs yellowish, with distal onefifth to one-fourth of femora dark. Underside with sterna reddish brown, or in two instances with metallic green sheen laterally, except 6 and lateral edges of 5 testaceous.</paragraph>
<paragraph id="8BABC101FFE1506EFF2B8416FBF6ED49" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">Head: Large, with widely spaced bulging eyes. Labrum with four very small apical teeth, more developed in female, and four submarginal setae; mandibles with smooth teeth; male with second tooth longer than third, female with third tooth longer than second, both sexes with a small fourth basal tooth.</paragraph>
<paragraph id="8BABC101FFE1506EFF2B847DFAB2EE14" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">Thorax with portion cut off by lateral carina (reddish brown with or without green sheen); pronotum moderately transverse, 1.07x wider than long, trapezoidal; maximum width at anterior third; posterior dorsal sulcus very well developed, lateral carina present throughout length.Elytra ovoid, humeri distinct but rounded, basal third punctate, punctation in irregular rows except third row with foveae parallel to suture, more coarsely and deeply impressed, and extending to apex; in some specimens also first row reaches apical part of elytra. Wingless.</paragraph>
<paragraph id="8BABC101FFE1506EFF2B8737FD0AEE71" blockId="1.[151,1437,903,2009]" box="[199,763,1588,1613]" pageId="1" pageNumber="144">Venter. Sternum VI of male with apex emarginate.</paragraph>
<paragraph id="8BABC101FFE1506EFF2B8755FB21EEE1" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">
Aedeagus similar to that in
<taxonomicName id="4C14BA82FFE1506EFDEE875BFC93EE4C" authority="Sloane, 1906" authorityName="Sloane" authorityYear="1906" box="[514,866,1623,1648]" class="Insecta" family="Carabidae" genus="Pseudotetracha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B9601D13FFE1506EFDEE875BFD45EE4C" box="[514,692,1623,1648]" italics="true" pageId="1" pageNumber="144">P. marginicollis</emphasis>
(
<bibRefCitation id="EF85BCF0FFE1506EFD2F8755FCA8EE4C" author="Sloane" box="[707,857,1623,1648]" pageId="1" pageNumber="144" refString="Sloane, T. G. (1906) Revision of the Cicindelidae of Australia. Proceedings of the Linnean Society of New South Wales, 31, 309 - 360." type="journal article" year="1906">Sloane, 1906</bibRefCitation>
)
</taxonomicName>
but with different outer curvature, especially near base where it widens in funnel-like fashion and creates a fold, whereas in
<taxonomicName id="4C14BA82FFE1506EFC3B877FFB79EEA9" box="[983,1160,1660,1685]" class="Insecta" family="Carabidae" genus="Pseudotetracha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B9601D13FFE1506EFC3B877FFB79EEA9" box="[983,1160,1660,1685]" italics="true" pageId="1" pageNumber="144">P. marginicollis</emphasis>
</taxonomicName>
widening is gradual and curvature in lateral view is uniterrupted, without fold (
<figureCitation id="132FDD84FFE1506EFCD2879DFC7AEE84" box="[830,907,1695,1720]" captionStart="FIGURE 1" captionStartId="2.[151,250,1046,1068]" captionTargetBox="[302,1282,201,1014]" captionTargetId="figure@2.[294,1292,195,1025]" captionTargetPageId="2" captionText="FIGURE 1 Pseudotetracha marginicollis (Sloane, 1906). A—male, habitus; B—aedeagus with expanded endophallus left, upper and right views; C—aedeagus in left lateral view." httpUri="https://zenodo.org/record/238912/files/figure.png" pageId="1" pageNumber="144">Figs 1</figureCitation>
c, 2c); apex is less downturned than in
<emphasis id="B9601D13FFE1506EFA6B87A3FEC2EEE1" italics="true" pageId="1" pageNumber="144">
<taxonomicName id="4C14BA82FFE1506EFA6B87A3FEDEEEE1" class="Insecta" family="Carabidae" genus="Pseudotetracha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="marginicollis">P. marginicollis</taxonomicName>
.
</emphasis>
. More pronounced differences are apparent in shape of endophallus (
<figureCitation id="132FDD84FFE1506EFBD787C6FB74EEE1" box="[1083,1157,1732,1757]" captionStart="FIGURE 1" captionStartId="2.[151,250,1046,1068]" captionTargetBox="[302,1282,201,1014]" captionTargetId="figure@2.[294,1292,195,1025]" captionTargetPageId="2" captionText="FIGURE 1 Pseudotetracha marginicollis (Sloane, 1906). A—male, habitus; B—aedeagus with expanded endophallus left, upper and right views; C—aedeagus in left lateral view." httpUri="https://zenodo.org/record/238912/files/figure.png" pageId="1" pageNumber="144">Figs. 1</figureCitation>
b, 2b).
</paragraph>
<paragraph id="8BABC101FFE1506EFF2B87E5FDD0EF51" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">
<emphasis id="B9601D13FFE1506EFF2B87E5FE3FEF3C" bold="true" box="[199,462,1767,1792]" pageId="1" pageNumber="144">Differential diagnosis.</emphasis>
<taxonomicName id="4C14BA82FFE1506EFE3A87EBFD83EF3C" box="[470,626,1767,1792]" class="Insecta" family="Carabidae" genus="Pseudotetracha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="timberensis" status="sp. nov.">
<emphasis id="B9601D13FFE1506EFE3A87EBFD83EF3C" box="[470,626,1767,1792]" italics="true" pageId="1" pageNumber="144">P. timberensis</emphasis>
</taxonomicName>
<emphasis id="B9601D13FFE1506EFD9687E5FD24EF3C" bold="true" box="[634,725,1767,1792]" pageId="1" pageNumber="144">
<taxonomicNameLabel id="A253A068FFE1506EFD9687E5FD24EF3C" box="[634,725,1767,1792]" pageId="1" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C14BA82FFE1506EFCB887EBFB45EF3C" authority="Sloane, 1906" authorityName="Sloane" authorityYear="1906" box="[852,1204,1767,1792]" class="Insecta" family="Carabidae" genus="Pseudotetracha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B9601D13FFE1506EFCB887EBFBF5EF3C" box="[852,1028,1767,1792]" italics="true" pageId="1" pageNumber="144">P. marginicollis</emphasis>
(
<bibRefCitation id="EF85BCF0FFE1506EFBF887E5FB5CEF3C" author="Sloane" box="[1044,1197,1767,1792]" pageId="1" pageNumber="144" refString="Sloane, T. G. (1906) Revision of the Cicindelidae of Australia. Proceedings of the Linnean Society of New South Wales, 31, 309 - 360." type="journal article" year="1906">Sloane, 1906</bibRefCitation>
)
</taxonomicName>
, differing from it by elytral sculpture with punctures in third row coarser, more deeply impressed and more extended to apex (
<figureCitation id="132FDD84FFE1506EFAD0860EFA8EEF18" box="[1340,1407,1804,1829]" captionStart="FIGURE 3" captionStartId="3.[151,250,1189,1211]" captionTargetBox="[341,1242,213,1148]" captionTargetId="figure@3.[327,1259,193,1167]" captionTargetPageId="3" captionText="FIGURE 3 Pseudotetracha timberensis sp. nov. A—female, habitus; B—proximal antennomeres, detail; C—elytral apex, detail." httpUri="https://zenodo.org/record/238914/files/figure.png" pageId="1" pageNumber="144">Fig. 3</figureCitation>
c); lighter overall coloration; by only distal darkening of femora and first four antennomeres (
<figureCitation id="132FDD84FFE1506EFB49862DFAF2EF74" box="[1189,1283,1839,1864]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1046,1068]" captionStartId-1="2.[151,250,1933,1955]" captionStartId-2="3.[151,250,1189,1211]" captionTargetBox-0="[302,1282,201,1014]" captionTargetBox-1="[328,1259,1158,1905]" captionTargetBox-2="[341,1242,213,1148]" captionTargetId-0="figure@2.[294,1292,195,1025]" captionTargetId-1="figure@2.[321,1265,1149,1912]" captionTargetId-2="figure@3.[327,1259,193,1167]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1 Pseudotetracha marginicollis (Sloane, 1906). A—male, habitus; B—aedeagus with expanded endophallus left, upper and right views; C—aedeagus in left lateral view." captionText-1="FIGURE 2 Pseudotetracha timberensis sp. nov. A—male, habitus; B—aedeagus with expanded endophallus left, upper and right views; C—aedeagus in left lateral view." captionText-2="FIGURE 3 Pseudotetracha timberensis sp. nov. A—female, habitus; B—proximal antennomeres, detail; C—elytral apex, detail." httpUri-0="https://zenodo.org/record/238912/files/figure.png" httpUri-1="https://zenodo.org/record/238913/files/figure.png" httpUri-2="https://zenodo.org/record/238914/files/figure.png" pageId="1" pageNumber="144">Figs 1-3</figureCitation>
a, 3b; and by shape of endophallus (
<figureCitation id="132FDD84FFE1506EFE7E8656FE27EF51" box="[402,470,1876,1901]" captionStart="FIGURE 1" captionStartId="2.[151,250,1046,1068]" captionTargetBox="[302,1282,201,1014]" captionTargetId="figure@2.[294,1292,195,1025]" captionTargetPageId="2" captionText="FIGURE 1 Pseudotetracha marginicollis (Sloane, 1906). A—male, habitus; B—aedeagus with expanded endophallus left, upper and right views; C—aedeagus in left lateral view." httpUri="https://zenodo.org/record/238912/files/figure.png" pageId="1" pageNumber="144">Figs 1</figureCitation>
b, 2b).
</paragraph>
</subSubSection>
<subSubSection id="C30E928AFFE1506EFF2B8675FED4EF89" pageId="1" pageNumber="144" type="etymology">
<paragraph id="8BABC101FFE1506EFF2B8675FED4EF89" blockId="1.[151,1437,903,2009]" pageId="1" pageNumber="144">
<emphasis id="B9601D13FFE1506EFF2B8675FEBFEFAC" bold="true" box="[199,334,1911,1936]" pageId="1" pageNumber="144">Etymology.</emphasis>
Named after Timber Creek, a tributary of Victoria River, the first known area of occurrence of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C30E928AFFE1506EFF2B86C2FBA2EFE5" box="[199,1107,1984,2009]" pageId="1" pageNumber="144" type="distribution">
<paragraph id="8BABC101FFE1506EFF2B86C2FBA2EFE5" blockId="1.[151,1437,903,2009]" box="[199,1107,1984,2009]" pageId="1" pageNumber="144">
<emphasis id="B9601D13FFE1506EFF2B86C2FE91EFE5" bold="true" box="[199,352,1984,2009]" pageId="1" pageNumber="144">Distribution.</emphasis>
Southwestern Northern Territory, northeastern Westerrn
<collectingCountry id="F3038191FFE1506EFC0886C2FBBFEFE5" box="[996,1102,1984,2009]" name="Australia" pageId="1" pageNumber="144">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>