treatments-xml/data/F7/46/87/F74687C8FFBEFFB6E69AFC36FE7289E7.xml
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<mods:title id="6CFD197A4B55999B170DE7D618FA39E0">On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
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<treatment id="F74687C8FFBEFFB6E69AFC36FE7289E7" ID-DOI="http://doi.org/10.5281/zenodo.3804478" ID-GBIF-Taxon="163732527" ID-Zenodo-Dep="3804478" LSID="urn:lsid:plazi:treatment:F74687C8FFBEFFB6E69AFC36FE7289E7" httpUri="http://treatment.plazi.org/id/F74687C8FFBEFFB6E69AFC36FE7289E7" lastPageId="6" lastPageNumber="195" pageId="4" pageNumber="193">
<subSubSection id="37F56555FFBEFFB4E69AFC36FD5C8E3B" box="[178,691,1004,1027]" pageId="4" pageNumber="193" type="nomenclature">
<paragraph id="7F5036DEFFBEFFB4E69AFC36FD5C8E3B" blockId="4.[178,691,1004,1027]" box="[178,691,1004,1027]" pageId="4" pageNumber="193">
<heading id="241881B2FFBEFFB4E69AFC36FD5C8E3B" box="[178,691,1004,1027]" fontSize="8" level="2" pageId="4" pageNumber="193" reason="2">
<emphasis id="4D9BEACCFFBEFFB4E69AFC36FD9A8E3B" bold="true" box="[178,629,1004,1027]" pageId="4" pageNumber="193">
<taxonomicName id="B8EF4D5DFFBEFFB4E69AFC36FE078E39" ID-CoL="4R338" authorityName="Eppelsheim" authorityYear="1888" box="[178,488,1004,1027]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="cohaesus" subGenus="Raphirus">
<emphasis id="4D9BEACCFFBEFFB4E69AFC36FEE68E39" bold="true" box="[178,265,1005,1027]" italics="true" pageId="4" pageNumber="193">Quedius</emphasis>
(
<emphasis id="4D9BEACCFFBEFFB4E730FC36FE968E39" bold="true" box="[280,377,1005,1027]" italics="true" pageId="4" pageNumber="193">Raphirus</emphasis>
)
<emphasis id="4D9BEACCFFBEFFB4E7A0FC36FE078E39" bold="true" box="[392,488,1005,1027]" italics="true" pageId="4" pageNumber="193">cohaesus</emphasis>
</taxonomicName>
<bibRefCitation id="1B7E4B2FFFBEFFB4E7C6FC37FD9A8E3B" author="EPPELSHEIM E" box="[494,629,1004,1025]" pageId="4" pageNumber="193" pagination="49 - 67" refId="ref6006" refString="EPPELSHEIM E. (1888): Neue Staphylinen Central-Asiens. Deutsche Entomolo gische Zeitschrift 32 (1): 49 - 67." type="journal article" year="1888">EPPELSHEIM</bibRefCitation>
</emphasis>
,
<emphasis id="4D9BEACCFFBEFFB4E4A8FC37FD5C8E3B" bold="true" box="[640,691,1004,1025]" pageId="4" pageNumber="193">1888</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="37F56555FFBEFFB4E69AFBC2FBD28C7B" pageId="4" pageNumber="193" type="reference_group">
<paragraph id="7F5036DEFFBEFFB4E69AFBC2FDD28E14" blockId="4.[178,686,1047,1095]" box="[178,573,1047,1070]" pageId="4" pageNumber="193">
<treatmentCitationGroup id="5FFF11F0FFBEFFB4E69AFBC2FDD28E14" box="[178,573,1047,1070]" pageId="4" pageNumber="193">
<taxonomicName id="B8EF4D5DFFBEFFB4E69AFBC2FEB98E17" ID-CoL="4R338" authorityName="Eppelsheim" authorityYear="1888" box="[178,342,1049,1069]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBEFFB4E69AFBC2FEB98E17" box="[178,342,1049,1069]" italics="true" pageId="4" pageNumber="193">Quedius cohaesus</emphasis>
</taxonomicName>
<treatmentCitation id="FE4E10CFFFBEFFB4E774FBC3FDC58E17" author="EPPELSHEIM E" box="[348,554,1047,1069]" page="60" pageId="4" pageNumber="193" year="1888">
<bibRefCitation id="1B7E4B2FFFBEFFB4E774FBC3FDC58E17" author="EPPELSHEIM E" box="[348,554,1047,1069]" pageId="4" pageNumber="193" pagination="49 - 67" refId="ref6006" refString="EPPELSHEIM E. (1888): Neue Staphylinen Central-Asiens. Deutsche Entomolo gische Zeitschrift 32 (1): 49 - 67." type="journal article" year="1888">
EPPELSHEIM, 1888:
<specimenCount id="69E9FD57FFBEFFB4E43BFBCCFDC58E17" box="[531,554,1047,1069]" pageId="4" pageNumber="193" type="generic">60</specimenCount>
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f.
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</paragraph>
<paragraph id="7F5036DEFFBEFFB4E69AFBE8FD418E7D" blockId="4.[178,686,1047,1095]" box="[178,686,1074,1095]" pageId="4" pageNumber="193">
<treatmentCitationGroup id="5FFF11F0FFBEFFB4E69AFBE8FD418E7D" box="[178,686,1074,1095]" pageId="4" pageNumber="193">
<taxonomicName id="B8EF4D5DFFBEFFB4E69AFBE8FE778E7D" ID-CoL="8VS2M" authority="REITTER, 1909" box="[178,408,1075,1095]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis id="4D9BEACCFFBEFFB4E69AFBE8FE778E7D" box="[178,408,1075,1095]" italics="true" pageId="4" pageNumber="193">Quedius pseudonigriceps</emphasis>
</taxonomicName>
<treatmentCitation id="FE4E10CFFFBEFFB4E7B5FBE9FDBA8E7D" author="REITTER E" box="[413,597,1074,1095]" page="113" pageId="4" pageNumber="193" year="1909">
<bibRefCitation id="1B7E4B2FFFBEFFB4E7B5FBE9FDBA8E7D" author="REITTER E" box="[413,597,1074,1095]" pageId="4" pageNumber="193" pagination="1 - 392" refId="ref6249" refString="REITTER E. (1909): Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. 2. 2 K. G. Lutz, Stuttgart: 1 - 392." type="book chapter" year="1909">REITTER, 1909: 113</bibRefCitation>
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;
<emphasis id="4D9BEACCFFBEFFB4E477FBE9FD418E7D" bold="true" box="[607,686,1074,1095]" pageId="4" pageNumber="193">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="7F5036DEFFBEFFB4E69AFB8DFBD28C7B" blockId="4.[178,1086,1110,1602]" pageId="4" pageNumber="193">
C o m m e n t: The original description of
<taxonomicName id="B8EF4D5DFFBEFFB4E451FB8CFD1A8E56" ID-CoL="4R338" authorityName="Eppelsheim" authorityYear="1888" box="[633,757,1111,1132]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBEFFB4E451FB8CFD1A8E56" box="[633,757,1111,1132]" italics="true" pageId="4" pageNumber="193">Q. cohaesus</emphasis>
</taxonomicName>
is based on three
<typeStatus id="A054887CFFBEFFB4E584FB8DFBEA8E51" box="[940,1029,1110,1131]" pageId="4" pageNumber="193" type="syntype">syntypes</typeStatus>
from ̎Turcmenien̎ (
<bibRefCitation id="1B7E4B2FFFBEFFB4E77EFBAFFDF78EB0" author="EPPELSHEIM E" box="[342,536,1140,1162]" pageId="4" pageNumber="193" pagination="49 - 67" refId="ref6006" refString="EPPELSHEIM E. (1888): Neue Staphylinen Central-Asiens. Deutsche Entomolo gische Zeitschrift 32 (1): 49 - 67." type="journal article" year="1888">EPPELSHEIM 1888</bibRefCitation>
), of which
<bibRefCitation id="1B7E4B2FFFBEFFB4E489FBAFFC7A8EB0" author="SOLODOVNIKOV A" box="[673,917,1140,1162]" pageId="4" pageNumber="193" pagination="221 - 243" refId="ref6455" refString="SOLODOVNIKOV A. (2004): Taxonomy and faunistics of some West Palearctic Quedius STEPHENS subgenus Raphirus STEPHENS (Coleoptera: Staphylinidae: Staphylininae). 2 Koleopterologische Rundschau 74: 221 - 243." type="journal article" year="2004">SOLODOVNIKOV (2004)</bibRefCitation>
designated the
<typeStatus id="A054887CFFBEFFB4E69AFB4AFEFD8E9D" box="[178,274,1169,1191]" pageId="4" pageNumber="193" type="lectotype">lectotype</typeStatus>
, that of
<emphasis id="4D9BEACCFFBEFFB4E74BFB49FDC78E9D" box="[355,552,1170,1191]" italics="true" pageId="4" pageNumber="193">Q. pseudonigriceps</emphasis>
on an unspecified number of
<typeStatus id="A054887CFFBEFFB4E57FFB4AFC5F8E9C" box="[855,944,1169,1190]" pageId="4" pageNumber="193" type="syntype">syntypes</typeStatus>
from ̎
<collectingCountry id="07F8764EFFBEFFB4E5DAFB4BFF108EFE" name="Bosnia and Herzegovina" pageId="4" pageNumber="193">Herzegowina</collectingCountry>
,
<collectingCountry id="07F8764EFFBEFFB4E724FB75FEBD8EFE" box="[268,338,1198,1220]" name="Turkey" pageId="4" pageNumber="193">Türkei</collectingCountry>
, Kleinasien̎ (
<bibRefCitation id="1B7E4B2FFFBEFFB4E7CFFB75FD918EFE" author="REITTER E" box="[487,638,1198,1220]" pageId="4" pageNumber="193" pagination="1 - 392" refId="ref6249" refString="REITTER E. (1909): Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. 2. 2 K. G. Lutz, Stuttgart: 1 - 392." type="book chapter" year="1909">REITTER 1909</bibRefCitation>
).
<taxonomicName id="B8EF4D5DFFBEFFB4E4BBFB74FCA28EFE" ID-CoL="4R338" authorityName="Eppelsheim" authorityYear="1888" box="[659,845,1199,1220]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBEFFB4E4BBFB74FCA28EFE" box="[659,845,1199,1220]" italics="true" pageId="4" pageNumber="193">Quedius cohaesus</emphasis>
</taxonomicName>
has three junior synonyms,
<emphasis id="4D9BEACCFFBEFFB4E72EFB16FE338EDB" box="[262,476,1228,1250]" italics="true" pageId="4" pageNumber="193">Q. pseudonigriceps</emphasis>
five (SCHÜLKE &amp; SMETANA 2015, SALNITSKA &amp; SOLODOVNIKOV 2018). According to the latest edition of the Palaearctic catalogue, the distribution of
<taxonomicName id="B8EF4D5DFFBEFFB4E762FAD3FE298F26" ID-CoL="4R338" authorityName="Eppelsheim" authorityYear="1888" box="[330,454,1287,1309]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBEFFB4E762FAD3FE298F26" box="[330,454,1287,1309]" italics="true" pageId="4" pageNumber="193">Q. cohaesus</emphasis>
</taxonomicName>
ranges from
<collectingCountry id="07F8764EFFBEFFB4E467FADDFD498F26" box="[591,678,1286,1308]" name="Bulgaria" pageId="4" pageNumber="193">Bulgaria</collectingCountry>
across
<collectingCountry id="07F8764EFFBEFFB4E4DBFADDFCD38F26" box="[755,828,1286,1308]" name="Turkey" pageId="4" pageNumber="193">Turkey</collectingCountry>
and the Caucasus region to
<collectingCountry id="07F8764EFFBEFFB4E6E7FAFFFF178F03" box="[207,248,1316,1337]" name="Iran" pageId="4" pageNumber="193">Iran</collectingCountry>
and
<collectingCountry id="07F8764EFFBEFFB4E707FAF8FE518F03" box="[303,446,1315,1337]" name="Turkmenistan" pageId="4" pageNumber="193">Turkmenistan</collectingCountry>
, whereas SALNITSKA &amp; SOLODOVNIKOV (2018) state that the distribution of this species includes
<collectingCountry id="07F8764EFFBEFFB4E411FA9AFD8B8F6C" box="[569,612,1345,1366]" name="Iran" pageId="4" pageNumber="193">Iran</collectingCountry>
,
<collectingCountry id="07F8764EFFBEFFB4E45CFA9AFCEC8F6D" box="[628,771,1345,1367]" name="Turkmenistan" pageId="4" pageNumber="193">Turkmenistan</collectingCountry>
,
<collectingCountry id="07F8764EFFBEFFB4E53BFA9AFC948F6D" box="[787,891,1345,1367]" name="Tajikistan" pageId="4" pageNumber="193">Tajikistan</collectingCountry>
, and
<collectingCountry id="07F8764EFFBEFFB4E593FA9AFBD58F6D" box="[955,1082,1345,1367]" name="Afghanistan" pageId="4" pageNumber="193">Afghanistan</collectingCountry>
. There is no reference to the records from other regions in the latter study, probably because it focuses on Middle Asia. The distributions given for
<emphasis id="4D9BEACCFFBEFFB4E564FAA6FBFB8FAB" box="[844,1044,1404,1426]" italics="true" pageId="4" pageNumber="193">Q. pseudonigriceps</emphasis>
are Southeast Europe northwards to
<collectingCountry id="07F8764EFFBEFFB4E439FA42FDB18F95" box="[529,606,1433,1455]" name="Austria" pageId="4" pageNumber="193">Austria</collectingCountry>
,
<collectingCountry id="07F8764EFFBEFFB4E445FA42FD568F95" box="[621,697,1433,1455]" name="Turkey" pageId="4" pageNumber="193">Turkey</collectingCountry>
, and the Caucasus region (
<collectingCountry id="07F8764EFFBEFFB4E5C9FA42FBD98F95" box="[993,1078,1433,1455]" name="Georgia" pageId="4" pageNumber="193">Georgia</collectingCountry>
) (SCHÜLKE &amp; SMETANA 2015), and South Europe, West Asia eastwards to Middle Asia (
<collectingCountry id="07F8764EFFBEFFB4E691FA0FFEDD8FD3" box="[185,306,1492,1513]" name="Kyrgyzstan" pageId="4" pageNumber="193">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="07F8764EFFBEFFB4E763FA08FE2C8FD3" box="[331,451,1491,1513]" name="Kazakhstan" pageId="4" pageNumber="193">Kazakhstan</collectingCountry>
) (SALNITSKA &amp; SOLODOVNIKOV 2018). SALNITSKA &amp; SOLODOVNIKOV (2018) regard
<taxonomicName id="B8EF4D5DFFBEFFB4E7DCFA29FD9C8C3D" ID-CoL="4R338" authorityName="Eppelsheim" authorityYear="1888" box="[500,627,1522,1543]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBEFFB4E7DCFA29FD9C8C3D" box="[500,627,1522,1543]" italics="true" pageId="4" pageNumber="193">Q. cohaesus</emphasis>
</taxonomicName>
and
<emphasis id="4D9BEACCFFBEFFB4E482FA29FC9D8C3D" box="[682,882,1522,1543]" italics="true" pageId="4" pageNumber="193">Q. pseudonigriceps</emphasis>
as distinct species, provide redescriptions and illustrations of the aedeagi based on material from Middle Asia, and state that
<taxonomicName id="B8EF4D5DFFBEFFB4E7B9F9F6FDFD8C78" ID-CoL="4R338" authorityName="Eppelsheim" authorityYear="1888" box="[401,530,1581,1602]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBEFFB4E7B9F9F6FDFD8C78" box="[401,530,1581,1602]" italics="true" pageId="4" pageNumber="193">Q. cohaesus</emphasis>
</taxonomicName>
is most similar to
<emphasis id="4D9BEACCFFBEFFB4E4C5F9F6FC578C78" box="[749,952,1581,1602]" italics="true" pageId="4" pageNumber="193">Q. pseudonigriceps</emphasis>
, but ̎easily
</paragraph>
</subSubSection>
<subSubSection id="37F56555FFBFFFB6E69AFF00FE7289E7" lastPageId="6" lastPageNumber="195" pageId="5" pageNumber="194" type="description">
<paragraph id="7F5036DEFFBFFFB5E69AFF00FC698B34" blockId="5.[178,1085,219,270]" pageId="5" pageNumber="194">distinguished by the presence of an apical seam of palisade fringe VII and normally developed elytra, as well as by the characters of the aedeagus̎ (p. 143).</paragraph>
<paragraph id="7F5036DEFFBFFFB5E69AFEC6FEE98BB1" blockId="5.[178,1085,285,395]" pageId="5" pageNumber="194">
Although I have not studied material from Middle Asia personally, the argumentation and conclusions laid out by SALNITSKA &amp; SOLODOVNIKOV (2018) are difficult to follow and
<emphasis id="4D9BEACCFFBFFFB5E6C8FE81FE468B54" box="[224,425,345,367]" italics="true" pageId="5" pageNumber="194">Q. pseudonigriceps</emphasis>
is considered a junior synonym of
<taxonomicName id="B8EF4D5DFFBFFFB5E501FE81FC478B54" authorityName="Eppelsheim" authorityYear="1888" box="[809,936,345,367]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBFFFB5E501FE81FC478B54" box="[809,936,345,367]" italics="true" pageId="5" pageNumber="194">Q. cohaesus</emphasis>
</taxonomicName>
for two main reasons:
</paragraph>
<paragraph id="7F5036DEFFBFFFB5E69AFE40FF1588EC" blockId="5.[178,1085,411,726]" pageId="5" pageNumber="194">
1. Based on personal observations and abundant material, the species previously identified as
<emphasis id="4D9BEACCFFBFFFB5E71FFE62FDD58BF4" box="[311,570,441,462]" italics="true" pageId="5" pageNumber="194">
<taxonomicName id="B8EF4D5DFFBFFFB5E71FFE62FE648BF4" box="[311,395,441,462]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
pseudonigriceps
</emphasis>
is one of the most widespread and most common representatives of the genus in the Balkans,
<collectingCountry id="07F8764EFFBFFFB5E456FE0EFD258BD1" box="[638,714,469,491]" name="Turkey" pageId="5" pageNumber="194">Turkey</collectingCountry>
, and the Caucasus region (
<collectingCountry id="07F8764EFFBFFFB5E5CBFE0EFBD78BD1" box="[995,1080,469,491]" name="Georgia" pageId="5" pageNumber="194">Georgia</collectingCountry>
,
<collectingCountry id="07F8764EFFBFFFB5E69AFE28FEE08833" box="[178,271,499,521]" name="Armenia" pageId="5" pageNumber="194">Armenia</collectingCountry>
). In this region, the species always has normally developed elytra and a palisade fringe at the posterior margin of tergite VII. This would mean that, according to SALNITSKA &amp; SOLODOVNIKOV (2018), there is an abrupt character displacement in the range of the species somewhere in North
<collectingCountry id="07F8764EFFBFFFB5E448FD90FD63885A" box="[608,652,587,608]" name="Iran" pageId="5" pageNumber="194">Iran</collectingCountry>
. True, there are several examples of wing di- or polymorphic
<taxonomicName id="B8EF4D5DFFBFFFB5E7A9FDB2FE3A8844" box="[385,469,617,638]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="genus">
<emphasis id="4D9BEACCFFBFFFB5E7A9FDB2FE3A8844" box="[385,469,617,638]" italics="true" pageId="5" pageNumber="194">Quedius</emphasis>
</taxonomicName>
species, sometimes even with clinal variation. It does not seem plausible, however, that there should be another, highly similar macropterous species (
<taxonomicName id="B8EF4D5DFFBFFFB5E724FD7FFE648883" authorityName="Eppelsheim" authorityYear="1888" box="[268,395,676,697]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBFFFB5E724FD7FFE648883" box="[268,395,676,697]" italics="true" pageId="5" pageNumber="194">Q. cohaesus</emphasis>
</taxonomicName>
) replacing the macropterous morph of
<emphasis id="4D9BEACCFFBFFFB5E502FD7FFC1D8883" box="[810,1010,676,697]" italics="true" pageId="5" pageNumber="194">Q. pseudonigriceps</emphasis>
in this region.
</paragraph>
<paragraph id="7F5036DEFFBFFFB5E69AFD3DFBED894B" blockId="5.[178,1085,741,881]" pageId="5" pageNumber="194">
2. The aedeagus of
<taxonomicName id="B8EF4D5DFFBFFFB5E752FD3CFE2188C6" box="[378,462,743,764]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="genus">
<emphasis id="4D9BEACCFFBFFFB5E752FD3CFE2188C6" box="[378,462,743,764]" italics="true" pageId="5" pageNumber="194">Quedius</emphasis>
</taxonomicName>
species is subject to - sometimes considerable - interspecific variation. This particularly applies to widespread species (
<bibRefCitation id="1B7E4B2FFFBFFFB5E538FCD8FC728923" author="ASSING V" box="[784,925,771,793]" pageId="5" pageNumber="194" pagination="129 - 182" refId="ref5867" refString="ASSING V. (2018): On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae). Linzer Biologische Beitrage 50 (1): 129 - 182." type="journal article" year="2018">ASSING 2018</bibRefCitation>
). SALNITSKA &amp; SOLODOVNIKOV (2018) may state that there are differences between
<taxonomicName id="B8EF4D5DFFBFFFB5E55BFCF9FC1F890C" authorityName="Eppelsheim" authorityYear="1888" box="[883,1008,801,823]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBFFFB5E55BFCF9FC1F890C" box="[883,1008,801,823]" italics="true" pageId="5" pageNumber="194">Q. cohaesus</emphasis>
</taxonomicName>
and
<emphasis id="4D9BEACCFFBFFFB5E20CFCF9FEB7896E" italics="true" pageId="5" pageNumber="194">Q. pseudonigriceps</emphasis>
in aedeagal characters, but no such discrete differences are visible in the illustrations they provide. Instead, they illustrate a continuum of aedeagal variation.
</paragraph>
<paragraph id="7F5036DEFFBFFFB5E69AFC5AFE3189EB" blockId="5.[178,1085,896,977]" pageId="5" pageNumber="194">
In consequence, the available evidence suggests that a)
<taxonomicName id="B8EF4D5DFFBFFFB5E52EFC59FC6889AC" authorityName="Eppelsheim" authorityYear="1888" box="[774,903,897,919]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="cohaesus">
<emphasis id="4D9BEACCFFBFFFB5E52EFC59FC6889AC" box="[774,903,897,919]" italics="true" pageId="5" pageNumber="194">Q. cohaesus</emphasis>
</taxonomicName>
is distributed on both sides of the Caspian Sea and that b) this species is wing-dimorphic in Middle Asia and monomorphic elsewhere.
</paragraph>
<paragraph id="7F5036DEFFBFFFB5E69AFC27FF028E11" blockId="5.[178,1064,1019,1067]" pageId="5" pageNumber="194">
<emphasis id="4D9BEACCFFBFFFB5E69AFC27FD878E2A" bold="true" box="[178,616,1019,1042]" pageId="5" pageNumber="194">
<taxonomicName id="B8EF4D5DFFBFFFB5E69AFC27FE158E2B" authorityName="Gridelli" authorityYear="1924" box="[178,506,1019,1042]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="invreae" subGenus="Microsaurus">
<emphasis id="4D9BEACCFFBFFFB5E69AFC27FEE68E28" bold="true" box="[178,265,1020,1042]" italics="true" pageId="5" pageNumber="194">Quedius</emphasis>
(
<emphasis id="4D9BEACCFFBFFFB5E730FC27FE728E2B" bold="true" box="[280,413,1020,1041]" italics="true" pageId="5" pageNumber="194">Microsaurus</emphasis>
)
<emphasis id="4D9BEACCFFBFFFB5E784FC27FE158E2B" bold="true" box="[428,506,1020,1041]" italics="true" pageId="5" pageNumber="194">invreae</emphasis>
</taxonomicName>
GRIDELLI
</emphasis>
,
<emphasis id="4D9BEACCFFBFFFB5E45BFC20FBCD8E2A" bold="true" box="[627,1058,1019,1042]" pageId="5" pageNumber="194">
1924 and
<taxonomicName id="B8EF4D5DFFBFFFB5E4F4FC27FC478E28" box="[732,936,1019,1042]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Microsaurus">
<emphasis id="4D9BEACCFFBFFFB5E4F4FC27FD1B8E2B" bold="true" box="[732,756,1020,1041]" italics="true" pageId="5" pageNumber="194">Q.</emphasis>
(
<emphasis id="4D9BEACCFFBFFFB5E52BFC26FCCF8E28" bold="true" box="[771,800,1021,1042]" italics="true" pageId="5" pageNumber="194">M.</emphasis>
)
<emphasis id="4D9BEACCFFBFFFB5E507FC27FC478E28" bold="true" box="[815,936,1020,1042]" italics="true" pageId="5" pageNumber="194">puncticollis</emphasis>
</taxonomicName>
(THOMSON
</emphasis>
,
<emphasis id="4D9BEACCFFBFFFB5E69AFBCEFF028E11" bold="true" box="[178,237,1045,1067]" pageId="5" pageNumber="194">1867)</emphasis>
</paragraph>
<paragraph id="7F5036DEFFBFFFB5E69AFB9DFD4A8F7F" blockId="5.[178,1085,1092,1351]" pageId="5" pageNumber="194">
C o m m e n t: According to the latest edition of the Palaearctic catalogue (SCHÜLKE &amp; SMETANA 2015), the distribution of
<taxonomicName id="B8EF4D5DFFBFFFB5E419FBBFFD4F8E40" authorityName="Gridelli" authorityYear="1924" box="[561,672,1124,1146]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBFFFB5E419FBBFFD4F8E40" box="[561,672,1124,1146]" italics="true" pageId="5" pageNumber="194">Q. invreae</emphasis>
</taxonomicName>
ranges from
<collectingCountry id="07F8764EFFBFFFB5E519FBB8FC978E42" box="[817,888,1123,1144]" name="France" pageId="5" pageNumber="194">France</collectingCountry>
,
<collectingCountry id="07F8764EFFBFFFB5E5AEFBB8FBE18E43" box="[902,1038,1123,1145]" name="United Kingdom" pageId="5" pageNumber="194">Great Britain</collectingCountry>
and Scandinavia across Central Europe southeastwards to the Russian South European territory and that of
<taxonomicName id="B8EF4D5DFFBFFFB5E74AFB44FE148E8E" box="[354,507,1183,1204]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBFFFB5E74AFB44FE148E8E" box="[354,507,1183,1204]" italics="true" pageId="5" pageNumber="194">Q. puncticollis</emphasis>
</taxonomicName>
from
<collectingCountry id="07F8764EFFBFFFB5E468FB45FD688E89" box="[576,647,1182,1203]" name="France" pageId="5" pageNumber="194">France</collectingCountry>
, the British Isles (including
<collectingCountry id="07F8764EFFBFFFB5E590FB46FBEB8E89" box="[952,1028,1181,1203]" name="Ireland" pageId="5" pageNumber="194">Ireland</collectingCountry>
), and Scandinavia across Central Europe southwards to
<collectingCountry id="07F8764EFFBFFFB5E4EAFB60FD1F8EEB" box="[706,752,1211,1233]" name="Italy" pageId="5" pageNumber="194">Italy</collectingCountry>
and eastwards to the Caucasus region and West Siberia. Recently,
<taxonomicName id="B8EF4D5DFFBFFFB5E40FFB01FD508ED4" box="[551,703,1241,1263]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBFFFB5E40FFB01FD508ED4" box="[551,703,1241,1263]" italics="true" pageId="5" pageNumber="194">Q. puncticollis</emphasis>
</taxonomicName>
was recorded also from
<collectingCountry id="07F8764EFFBFFFB5E5EFFB03FBE08ED7" box="[967,1039,1240,1261]" name="Greece" pageId="5" pageNumber="194">Greece</collectingCountry>
and Middle Asia (
<bibRefCitation id="1B7E4B2FFFBFFFB5E762FB2DFE388F31" author="ASSING V" box="[330,471,1270,1291]" pageId="5" pageNumber="194" pagination="207 - 228" refId="ref5811" refString="ASSING V. (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer B iologische Beitrage 49 (1): 207 - 228." type="journal article" year="2017">ASSING 2017</bibRefCitation>
, SALNITSKA &amp; SOLODOVNIKOV 2018). In his key to the Central European
<taxonomicName id="B8EF4D5DFFBFFFB5E746FAC8FE228F13" authorityName="Kraatz" authorityYear="1857" box="[366,461,1299,1321]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="subTribe" subTribe="Quediina">Quediina</taxonomicName>
,
<bibRefCitation id="1B7E4B2FFFBFFFB5E7F1FAC8FD278F13" author="SOLODOVNIKOV A" box="[473,712,1299,1321]" pageId="5" pageNumber="194" pagination="451 - 484" refId="ref6491" refString="SOLODOVNIKOV A. (2012): Subtribus Quediina KRAATZ, 1857; pp. 451 - 484. 2 In: A SSING V. &amp; M. SCHULKE (eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4, Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg: I-XII, 1 - 560." type="book chapter" year="2012">SOLODOVNIKOV (2012)</bibRefCitation>
doubted the taxonomic status of
<taxonomicName id="B8EF4D5DFFBFFFB5E20CFACFFF118F7D" authorityName="Gridelli" authorityYear="1924" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBFFFB5E20CFACFFF118F7D" italics="true" pageId="5" pageNumber="194">Q. invreae</emphasis>
</taxonomicName>
and suspected synonymy of these names.
</paragraph>
<paragraph id="7F5036DEFFBFFFB6E69AFA8EFE9C886A" blockId="5.[178,1085,1365,1593]" lastBlockId="6.[178,1085,219,594]" lastPageId="6" lastPageNumber="195" pageId="5" pageNumber="194">
Inspired by records of specimens preliminarily identified as
<taxonomicName id="B8EF4D5DFFBFFFB5E504FA8CFC758F56" authorityName="Gridelli" authorityYear="1924" box="[812,922,1367,1388]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBFFFB5E504FA8CFC758F56" box="[812,922,1367,1388]" italics="true" pageId="5" pageNumber="194">Q. invreae</emphasis>
</taxonomicName>
from
<collectingCountry id="07F8764EFFBFFFB5E5F6FA8EFBD68F51" box="[990,1081,1365,1387]" name="Armenia" pageId="5" pageNumber="194">Armenia</collectingCountry>
, material of
<taxonomicName id="B8EF4D5DFFBFFFB5E705FAAFFE2A8FB3" box="[301,453,1396,1417]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBFFFB5E705FAAFFE2A8FB3" box="[301,453,1396,1417]" italics="true" pageId="5" pageNumber="194">Q. puncticollis</emphasis>
</taxonomicName>
and
<taxonomicName id="B8EF4D5DFFBFFFB5E7D5FAAFFD838FB0" authorityName="Gridelli" authorityYear="1924" box="[509,620,1396,1418]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBFFFB5E7D5FAAFFD838FB0" box="[509,620,1396,1418]" italics="true" pageId="5" pageNumber="194">Q. invreae</emphasis>
</taxonomicName>
in the authoŕs collection was revised. This study revealed that both names undoubtedly represent distinct taxa and that both species are easily distinguished. The aedeagi of both species may be rather similar, but the paramere of
<taxonomicName id="B8EF4D5DFFBFFFB5E713FA17FE448FD8" authorityName="Gridelli" authorityYear="1924" box="[315,427,1484,1506]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBFFFB5E713FA17FE448FD8" box="[315,427,1484,1506]" italics="true" pageId="5" pageNumber="194">Q. invreae</emphasis>
</taxonomicName>
is broader and usually weakly concave apically, more rarely truncate or weakly convex, whereas in
<taxonomicName id="B8EF4D5DFFBFFFB5E47DFA32FD018FC4" box="[597,750,1513,1534]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBFFFB5E47DFA32FD018FC4" box="[597,750,1513,1534]" italics="true" pageId="5" pageNumber="194">Q. puncticollis</emphasis>
</taxonomicName>
it is more slender and apically distinctly convex. The best characters for the separation of both species, however, are the colour, shape, and especially the chaetotaxy of sternite VIII, especially in the male, as already stated by
<bibRefCitation id="1B7E4B2FFFBCFFB6E742FF00FE128ACB" author="LOHSE G" box="[362,509,219,241]" pageId="6" pageNumber="195" pagination="1 - 264" refId="ref6165" refString="LOHSE G. A. (1964): Staphylinidae I (Micropeplinae bis Tachyporinae). 2 In: F REUDE H., HARDE K. W. &amp; G. A. LOHSE (eds.), Die Kafer Mitteleuropas. Band 4. Goecke &amp; Evers, Krefeld: 1 - 264." type="book chapter" year="1964">LOHSE (1964)</bibRefCitation>
. In
<taxonomicName id="B8EF4D5DFFBCFFB6E40DFF07FD7D8AC8" authorityName="Gridelli" authorityYear="1924" box="[549,658,220,242]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBCFFB6E40DFF07FD7D8AC8" box="[549,658,220,242]" italics="true" pageId="6" pageNumber="195">Q. invreae</emphasis>
</taxonomicName>
, sternite VIII is darker, often completely black or at least black in anterior three-fourths with the posterior fourth sometimes slightly paler, whereas in
<taxonomicName id="B8EF4D5DFFBCFFB6E79FFECCFDA38B16" box="[439,588,279,300]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E79FFECCFDA38B16" box="[439,588,279,300]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
tergite VIII is paler with at least the anterior and posterior portions pale-brownish and the median portion usually blackish-brown. More importantly, however, the posterior margin of the male sternite VIII is broadly concave and furnished with extremely long black setae (especially laterally), the longest of them longer than antennomere I in
<taxonomicName id="B8EF4D5DFFBCFFB6E7D1FE57FD858B98" authorityName="Gridelli" authorityYear="1924" box="[505,618,396,418]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBCFFB6E7D1FE57FD858B98" box="[505,618,396,418]" italics="true" pageId="6" pageNumber="195">Q. invreae</emphasis>
</taxonomicName>
. In
<taxonomicName id="B8EF4D5DFFBCFFB6E4B5FE57FCD88B9B" box="[669,823,396,417]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E4B5FE57FCD88B9B" box="[669,823,396,417]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
, on the other hand, the posterior margin is shallowly concave only in the middle and furnished with short brown setae, the longest of them barely half as long as antennomere I. The posterior margin of the female sternite VIII is furnished with relatively short setae in both species, but these setae are black in
<taxonomicName id="B8EF4D5DFFBCFFB6E756FDD9FE00882D" authorityName="Gridelli" authorityYear="1924" box="[382,495,514,535]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBCFFB6E756FDD9FE00882D" box="[382,495,514,535]" italics="true" pageId="6" pageNumber="195">Q. invreae</emphasis>
</taxonomicName>
and brown in
<taxonomicName id="B8EF4D5DFFBCFFB6E4B0FDD9FCDC882C" box="[664,819,513,535]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E4B0FDD9FCDC882C" box="[664,819,513,535]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
. The illustrations of the aedeagus of
<taxonomicName id="B8EF4D5DFFBCFFB6E71EFDC4FE21880E" box="[310,462,543,564]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E71EFDC4FE21880E" box="[310,462,543,564]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="1B7E4B2FFFBCFFB6E475FDC5FCA68809" author="SOLODOVNIKOV A" box="[605,841,542,563]" pageId="6" pageNumber="195" pagination="451 - 484" refId="ref6491" refString="SOLODOVNIKOV A. (2012): Subtribus Quediina KRAATZ, 1857; pp. 451 - 484. 2 In: A SSING V. &amp; M. SCHULKE (eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4, Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg: I-XII, 1 - 560." type="book chapter" year="2012">SOLODOVNIKOV (2012</bibRefCitation>
: figures 191t-v) clearly refer to
<taxonomicName id="B8EF4D5DFFBCFFB6E72AFDE7FE828868" authorityName="Gridelli" authorityYear="1924" box="[258,365,572,594]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBCFFB6E72AFDE7FE828868" box="[258,365,572,594]" italics="true" pageId="6" pageNumber="195">Q. invreae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7F5036DEFFBCFFB6E69AFDBBFE7289E7" blockId="6.[178,1085,608,989]" pageId="6" pageNumber="195">
According to
<bibRefCitation id="1B7E4B2FFFBCFFB6E76CFDBAFE38884C" author="LOHSE G" box="[324,471,608,630]" pageId="6" pageNumber="195" pagination="1 - 264" refId="ref6165" refString="LOHSE G. A. (1964): Staphylinidae I (Micropeplinae bis Tachyporinae). 2 In: F REUDE H., HARDE K. W. &amp; G. A. LOHSE (eds.), Die Kafer Mitteleuropas. Band 4. Goecke &amp; Evers, Krefeld: 1 - 264." type="book chapter" year="1964">LOHSE (1964)</bibRefCitation>
,
<taxonomicName id="B8EF4D5DFFBCFFB6E7CDFDB9FD92884C" box="[485,637,609,631]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E7CDFDB9FD92884C" box="[485,637,609,631]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
is the most common representative of the genus in mole nests. This may be true of the environs of
<collectingRegion id="BD2BF83CFFBCFFB6E514FDA5FC7188AE" box="[828,926,638,660]" country="Germany" name="Hamburg" pageId="6" pageNumber="195">Hamburg</collectingRegion>
, G.A. Lohsés hometown, but certainly not of regions farther south. Based on personal observations (studies of mole nests, extensive pitfall trap studies),
<taxonomicName id="B8EF4D5DFFBCFFB6E4F3FD60FC9E88EA" box="[731,881,699,720]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E4F3FD60FC9E88EA" box="[731,881,699,720]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
is rare, much more so than
<taxonomicName id="B8EF4D5DFFBCFFB6E724FD02FE9288D5" authorityName="Gridelli" authorityYear="1924" box="[268,381,729,751]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBCFFB6E724FD02FE9288D5" box="[268,381,729,751]" italics="true" pageId="6" pageNumber="195">Q. invreae</emphasis>
</taxonomicName>
, and the southernmost record examined is from South
<collectingCountry id="07F8764EFFBCFFB6E5F6FD03FBD388D7" box="[990,1084,728,749]" name="Germany" pageId="6" pageNumber="195">Germany</collectingCountry>
(München). While most of the few specimens of
<taxonomicName id="B8EF4D5DFFBCFFB6E49AFD2CFCA68936" box="[690,841,759,780]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E49AFD2CFCA68936" box="[690,841,759,780]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
in cAss were extracted from mole nests (some also collected with pitfall traps), material of
<taxonomicName id="B8EF4D5DFFBCFFB6E54FFCCEFBE18910" authorityName="Gridelli" authorityYear="1924" box="[871,1038,789,810]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBCFFB6E54FFCCEFBE18910" box="[871,1038,789,810]" italics="true" pageId="6" pageNumber="195">Quedius invreae</emphasis>
</taxonomicName>
was exclusively collected with pitfall and flight interception traps (especially in warmer habitats; never from mole nests). A re-examination of a specimen from
<collectingCountry id="07F8764EFFBCFFB6E58FFC94FC1E895E" box="[935,1009,847,868]" name="Greece" pageId="6" pageNumber="195">Greece</collectingCountry>
, which the record of
<taxonomicName id="B8EF4D5DFFBCFFB6E779FCB4FE0189B9" box="[337,494,878,900]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E779FCB4FE0189B9" box="[337,494,878,900]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
in
<bibRefCitation id="1B7E4B2FFFBCFFB6E437FCB6FD5289B9" author="ASSING V" box="[543,701,877,899]" pageId="6" pageNumber="195" pagination="207 - 228" refId="ref5811" refString="ASSING V. (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer B iologische Beitrage 49 (1): 207 - 228." type="journal article" year="2017">ASSING (2017)</bibRefCitation>
is based on, revealed that it was misidentified and in fact belongs to
<taxonomicName id="B8EF4D5DFFBCFFB6E436FC56FD668998" authorityName="Gridelli" authorityYear="1924" box="[542,649,909,930]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="invreae">
<emphasis id="4D9BEACCFFBCFFB6E436FC56FD668998" box="[542,649,909,930]" italics="true" pageId="6" pageNumber="195">Q. invreae</emphasis>
</taxonomicName>
(first record from
<collectingCountry id="07F8764EFFBCFFB6E56EFC50FC7D899B" box="[838,914,907,929]" name="Greece" pageId="6" pageNumber="195">Greece</collectingCountry>
). The same most likely applies to records of
<taxonomicName id="B8EF4D5DFFBCFFB6E42EFC71FD478985" box="[518,680,938,959]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="4D9BEACCFFBCFFB6E42EFC71FD478985" box="[518,680,938,959]" italics="true" pageId="6" pageNumber="195">Q. puncticollis</emphasis>
</taxonomicName>
from Middle Asia (SALNITSKA &amp; SOLODOVNIKOV 2018).
</paragraph>
</subSubSection>
</treatment>
</document>