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<mods:title id="0B45C25EDD2860D8D6D493A92B4730DF">The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa</mods:title>
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<mods:namePart id="60A2AC59812E1F4AF7F0FCA3B1D97FF1">Daniel, Gimo M.</mods:namePart>
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<mods:namePart id="5EF46ABE630199D3AB56CEAE97860A51">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="C9D4F978604F3B26879DF6F467009E91">Sole, Catherine L.</mods:namePart>
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Identification key to
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<emphasis id="B937D606157BA14611D48D48FDE94144" bold="true" box="[411,561,565,592]" italics="true" pageId="9" pageNumber="403">Stiptopodius</emphasis>
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species modified from
<bibRefCitation id="EFD277E5157BA146131E8D4BFBD94144" author="Branco" box="[849,1025,566,592]" firstAuthor="Branco" pageId="9" pageNumber="403" pagination="87 - 91" refId="ref7180" refString="Branco, T. (2009) Notes on &quot; Stiptopodius &quot; Harold, 1871: number of tarsal segments of S. longipedis Branco, 1991, and description of a new species from Zambia (Coleoptera, Scarabaeidae). Boletin Sociedad Entomologica Aragonesa, 44, 87 - 91." type="journal article" year="2009">Branco (2009)</bibRefCitation>
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1 Sides of pronotum (lateral view) weakly sinuate before postero-lateral angle (
<taxonomicName id="4C437197157BA14613848D06FBFA419B" ID-CoL="52RTY" authorityName="Harold" authorityYear="1871" box="[971,1058,634,656]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="B937D606157BA14613848D06FBFA419B" box="[971,1058,634,656]" italics="true" pageId="9" pageNumber="403">S. doriae</emphasis>
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species group).................. 2
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Sides of pronotum (lateral view) strongly sinuate before postero-lateral angle (
<taxonomicName id="4C437197157BA146139B8DE5FB9441B9" ID-CoL="6ZVQR" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[980,1100,664,685]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D606157BA146139B8DE5FB9441B9" box="[980,1100,664,685]" italics="true" pageId="9" pageNumber="403">S. singularis</emphasis>
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species group).............. 4
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2 Body elongate, parallel-sided. Pronotum quadrate, much shorter than elytra. Head with two oblique tubercles on the vertex, often obsolete. Length
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...........................................................................
<taxonomicName id="4C437197157BA14614148D8DFA444011" ID-CoL="52RTY" authority="Harold, 1871" authorityName="Harold" authorityYear="1871" box="[1115,1436,752,774]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="doriae">
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<bibRefCitation id="EFD277E5157BA14615518D8DFA444011" author="Harold" box="[1310,1436,752,774]" firstAuthor="Harold" pageId="9" pageNumber="403" pagination="1 - 28" refId="ref7328" refString="Harold, E. von (1871) Verzeichniss der von Dr Beccari in Bogos gesammelten Coprophagen Lamellicornien. Coleopterologische Hefte, 8, 1 - 28. https: // doi. org / 10.1002 / mmnd. 47918710229" type="journal article" year="1871">Harold, 1871</bibRefCitation>
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<keyLead id="30B2A921157BA14610D88C70FA44402B" pageId="9" pageNumber="403"> Body short and wide. Pronotum with sides arcuate, only slightly shorter than elytra Head with two transversely-aligned weak tubercles on vertex.................................................................................... 3</keyLead>
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3 Pronotal disc brown with metallic sheen. Meso- and metatarsi with five tarsomeres and apical pair of claws, meso- and metatarsomeres I to IV triangular and wider than long. (male; female unknown). Length 4.0 mm.
<collectingCountry id="F3544A84157BA146143B8C18FB28406F" box="[1140,1264,869,891]" name="South Africa" pageId="9" pageNumber="403">South Africa</collectingCountry>
(
<collectingRegion id="4987C4F6157BA14614B18C18FA41406F" box="[1278,1433,869,891]" country="South Africa" name="KwaZulu-Natal" pageId="9" pageNumber="403">KwaZulu-Natal</collectingRegion>
, Richards Bay)...........................................
<taxonomicName id="4C437197157BA14613368CFFFAC7408C" ID-CoL="6ZVQK" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[889,1311,898,920]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="peringueyi" status="sp. nov.">
<emphasis id="B937D606157BA14613368CFFFB86408C" box="[889,1118,898,920]" italics="true" pageId="9" pageNumber="403">Stiptopodius peringueyi</emphasis>
Daniel &amp; Deschodt
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,
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Pronotal disc dull. Meso- and metatarsi clawless with three tarsomeres, meso and metatarsomeres I and II much longer than wide, tarsomere III small and not bilobate. Length
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,
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....
<taxonomicName id="4C437197157BA146147C8CC0FA4440C6" ID-CoL="6ZW2G" authority="Branco, 1991" authorityName="Branco" authorityYear="1991" box="[1075,1436,957,978]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="longipedis">
<emphasis id="B937D606157BA146147C8CC0FACD40C6" box="[1075,1301,957,978]" italics="true" pageId="9" pageNumber="403">Stiptopodius longipedis</emphasis>
<bibRefCitation id="EFD277E5157BA14615548CC0FA4440C6" author="Branco" box="[1307,1436,957,978]" firstAuthor="Branco" pageId="9" pageNumber="403" pagination="265 - 285" refId="ref7093" refString="Branco, T. (1991) Revision des genres du &quot; groupe &quot; Stiptopodius: Les genres Stiptopodius Harold et Stiptotarsus Branco (Coleoptera: Scarabaeidae). Annales de la Societe Entomologique de France, Nouvelle Serie, 27, 265 - 285." type="journal article" year="1991">Branco, 1991</bibRefCitation>
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<keyLead id="30B2A921157BA14610D88CA7FA4440E4" box="[151,1436,986,1008]" pageId="9" pageNumber="403">4 Head with two clearly separated tubercles on vertex. Clypeo-frontal carina present................................. 5</keyLead>
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<keyLead id="30B2A921157BA14610D88C85FA44473F" pageId="9" pageNumber="403"> Head with carina on vertex, entire or slightly interrupted in the middle, but never divided into two separated small tubercles. Clypeo-frontal carina absent............................................................................ 6</keyLead>
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5 Cephalic tubercles approximately equidistant from each other and the inner edge of eyes; space between tubercles flat or very slightly depressed. Length
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<collectingCountry id="F3544A84157BA146122B8B2DFD704772" box="[612,680,1104,1126]" name="Eritrea" pageId="9" pageNumber="403">Eritrea</collectingCountry>
,
<collectingCountry id="F3544A84157BA14612FE8B2DFCDD4772" box="[689,773,1104,1126]" name="Ethiopia" pageId="9" pageNumber="403">Ethiopia</collectingCountry>
,
<collectingCountry id="F3544A84157BA14613418B2DFC874772" box="[782,863,1104,1126]" name="Somalia" pageId="9" pageNumber="403">Somalia</collectingCountry>
,
<collectingCountry id="F3544A84157BA14613268B2DFC724771" box="[873,938,1104,1125]" name="Kenya" pageId="9" pageNumber="403">Kenya</collectingCountry>
,
<collectingCountry id="F3544A84157BA14613FC8B2DFBD14772" box="[947,1033,1104,1126]" name="Tanzania" pageId="9" pageNumber="403">Tanzania</collectingCountry>
.....................................................................................................
<taxonomicName id="4C437197157BA14614688B10FA444796" ID-CoL="6ZW2L" authority="Boucomont, 1923" authorityName="Boucomont" authorityYear="1923" box="[1063,1436,1133,1155]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="patrizii">
<emphasis id="B937D606157BA14614688B10FB324797" box="[1063,1258,1133,1155]" italics="true" pageId="9" pageNumber="403">Stiptopodius patrizii</emphasis>
Boucomont, 1923
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D88BF6FA4447A9" blockId="9.[151,1437,1074,1214]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88BF6FA4447A9" pageId="9" pageNumber="403">
Cephalic tubercles much closer to inner edge of eyes than to each other; space between tubercles strongly depressed (male; female unknown). Length
<quantity id="4CBBA7F1157BA14611AF8BD5FDF447A9" box="[480,556,1192,1213]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="9" pageNumber="403" unit="mm" value="7.3">7.3 mm</quantity>
.
<collectingCountry id="F3544A84157BA14612798BD5FDAD47A9" box="[566,629,1192,1213]" name="Kenya" pageId="9" pageNumber="403">Kenya</collectingCountry>
......................................
<taxonomicName id="4C437197157BA146141F8BD5FA4447A9" ID-CoL="6ZW2H" authority="Branco, 1991" authorityName="Branco" authorityYear="1991" box="[1104,1436,1192,1213]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="krausei">
<emphasis id="B937D606157BA146141F8BD5FACD47A9" box="[1104,1301,1192,1213]" italics="true" pageId="9" pageNumber="403">Stiptopodius krausei</emphasis>
<bibRefCitation id="EFD277E5157BA14615548BD5FA4447A9" author="Branco" box="[1307,1436,1192,1213]" firstAuthor="Branco" pageId="9" pageNumber="403" pagination="265 - 285" refId="ref7093" refString="Branco, T. (1991) Revision des genres du &quot; groupe &quot; Stiptopodius: Les genres Stiptopodius Harold et Stiptotarsus Branco (Coleoptera: Scarabaeidae). Annales de la Societe Entomologique de France, Nouvelle Serie, 27, 265 - 285." type="journal article" year="1991">Branco, 1991</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14610D88BB8FA4447E3" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D88BB8FA4447CE" blockId="9.[151,1436,1221,1272]" box="[151,1436,1221,1243]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88BB8FA4447CE" box="[151,1436,1221,1243]" pageId="9" pageNumber="403">6 Clypeal surface not granular, punctation varies from small, simple punctures to large and ocellate..................... 7</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D88B9EFA4447E3" blockId="9.[151,1436,1221,1272]" box="[151,1436,1250,1272]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88B9EFA4447E3" box="[151,1436,1250,1272]" pageId="9" pageNumber="403"> Clypeal surface at least partly granular and punctate........................................................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14610D88A7DFA444679" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D88A7DFA444626" blockId="9.[151,1436,1280,1390]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88A7DFA444626" pageId="9" pageNumber="403">7 Second elytral interstria with punctation aligned alongside the striae, middle of interstria apunctate. Head and pronotum of metallic colour or with a metallic sheen................................................................... 8</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D88A46FA444679" blockId="9.[151,1436,1280,1390]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88A46FA444679" pageId="9" pageNumber="403"> Second elytral interstria with punctation irregularly distributed over entire surface. Head and pronotum black or brown, without metallic sheen...................................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14610D88A08FA4446D2" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D88A08FA44469E" blockId="9.[151,1437,1397,1478]" box="[151,1436,1397,1419]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88A08FA44469E" box="[151,1436,1397,1419]" pageId="9" pageNumber="403">8 Disc of pronotum glabrous at basal half, densely punctate. Frontal carina most often obsolete........................ 9</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D88AEEFA4446D2" blockId="9.[151,1437,1397,1478]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88AEEFA4446D2" pageId="9" pageNumber="403">
Disc of pronotum pilose at basal half, with sparsely set double punctation, the larger punctures setiferous. Frontal carina most often strong. Length
<quantity id="4CBBA7F1157BA14611E48ACDFDF946D1" box="[427,545,1456,1478]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.6" metricValueMin="5.3" pageId="9" pageNumber="403" unit="mm" value="5.949999999999999" valueMax="6.6" valueMin="5.3">5.36.6 mm</quantity>
.
<collectingCountry id="F3544A84157BA14612648ACDFDB446D1" box="[555,620,1456,1477]" name="Kenya" pageId="9" pageNumber="403">Kenya</collectingCountry>
,
<collectingCountry id="F3544A84157BA146123A8ACDFD1646D2" box="[629,718,1456,1478]" name="Tanzania" pageId="9" pageNumber="403">Tanzania</collectingCountry>
,
<collectingCountry id="F3544A84157BA14612988ACDFCF846D2" box="[727,800,1456,1478]" name="Malawi" pageId="9" pageNumber="403">Malawi</collectingCountry>
...................
<taxonomicName id="4C437197157BA14614548ACDFA4446D2" ID-CoL="6ZVQM" authority="(Boucomont, 1923)" baseAuthorityName="Boucomont" baseAuthorityYear="1923" box="[1051,1436,1456,1478]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B937D606157BA14614548ACDFB0246D1" box="[1051,1242,1456,1477]" italics="true" pageId="9" pageNumber="403">Stiptopodius nitidus</emphasis>
(Boucomont, 1923)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14610D88AB0FA454562" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D88AB0FA44450A" blockId="9.[151,1437,1485,1654]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88AB0FA44450A" pageId="9" pageNumber="403">
9 Pronotum with uniform, dense and medium-sized punctation. Elytra sparsely setose posteriorly. Mesotibiae not sinuate externally. Parameres short, with blunt tips. Length 4.56.0 mm.
<collectingCountry id="F3544A84157BA14613538A97FCAD4514" box="[796,885,1514,1536]" name="Tanzania" pageId="9" pageNumber="403">Tanzania</collectingCountry>
,
<collectingCountry id="F3544A84157BA14613318A97FC124514" box="[894,970,1514,1536]" name="Malawi" pageId="9" pageNumber="403">Malawi</collectingCountry>
,
<collectingCountry id="F3544A84157BA146139D8A97FB8B4514" box="[978,1107,1514,1536]" name="Mozambique" pageId="9" pageNumber="403">Mozambique</collectingCountry>
,
<collectingCountry id="F3544A84157BA14614138A97FB704514" box="[1116,1192,1514,1536]" name="Zambia" pageId="9" pageNumber="403">Zambia</collectingCountry>
,
<collectingCountry id="F3544A84157BA14614FE8A97FAC24514" box="[1201,1306,1514,1536]" name="Zimbabwe" pageId="9" pageNumber="403">Zimbabwe</collectingCountry>
,
<collectingCountry id="F3544A84157BA146156C8A97FA444514" box="[1315,1436,1514,1536]" name="South Africa" pageId="9" pageNumber="403">South Africa</collectingCountry>
(
<collectingRegion id="4987C4F6157BA14610A78975FE9C450A" box="[232,324,1544,1566]" country="South Africa" name="Limpopo" pageId="9" pageNumber="403">Limpopo</collectingRegion>
,
<collectingRegion id="4987C4F6157BA14611018975FE474509" box="[334,415,1544,1565]" country="South Africa" name="Gauteng" pageId="9" pageNumber="403">Gauteng</collectingRegion>
and Natal),
<collectingCountry id="F3544A84157BA146125B8975FD83450A" box="[532,603,1544,1566]" name="Angola" pageId="9" pageNumber="403">Angola</collectingCountry>
...................................
<taxonomicName id="4C437197157BA14614438975FA44450A" ID-CoL="6ZVQR" authority="(Peringuey, 1901)" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[1036,1436,1544,1566]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D606157BA14614438975FB324509" box="[1036,1258,1544,1565]" italics="true" pageId="9" pageNumber="403">Stiptopodius singularis</emphasis>
(
<bibRefCitation id="EFD277E5157BA14614B88975FA4D450A" author="Peringuey" box="[1271,1429,1544,1566]" firstAuthor="Peringuey" pageId="9" pageNumber="403" pagination="1 - 920" refId="ref7395" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12, 1 - 920. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D88958FA454562" blockId="9.[151,1437,1485,1654]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88958FA454562" pageId="9" pageNumber="403">
Pronotum with punctures of mixed size from fine to large. Elytra fully setose. Mesotibiae sinuate externally. Parameres with tips sharp and projecting forward (male; female unknown). Length 7.6. mm.
<collectingCountry id="F3544A84157BA146138D893FFBD1454C" box="[962,1033,1602,1624]" name="Angola" pageId="9" pageNumber="403">Angola</collectingCountry>
.......................................................................................
<taxonomicName id="4C437197157BA1461335891DFAC14562" ID-CoL="6ZW2N" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[890,1305,1632,1654]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D606157BA1461335891DFB804561" box="[890,1112,1632,1653]" italics="true" pageId="9" pageNumber="403">Stiptopodius muellerae</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D157BA146156C891DFA414562" box="[1315,1433,1632,1654]" pageId="9" pageNumber="403" rank="species">new species</taxonomicNameLabel>
,
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14610D88900FA4445FF" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D88900FA4445BB" blockId="9.[151,1437,1661,1771]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88900FA4445BB" pageId="9" pageNumber="403">10 Disc of pronotum glabrous, with uniform punctation, punctures of fine to medium-size. Elytral interstriae either pilose or glabrous............................................................................................. 11</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D889C5FA4445FF" blockId="9.[151,1437,1661,1771]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D889C5FA4445FF" pageId="9" pageNumber="403">
Disc of pronotum pilose, with double punctation, the larger punctures fairly strong, ocellate and setigerous. Elytra always entirely pilose. Length
<quantity id="4CBBA7F1157BA14611E589A8FDF845FE" box="[426,544,1749,1771]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.3" metricValueMin="6.5" pageId="9" pageNumber="403" unit="mm" value="6.9" valueMax="7.3" valueMin="6.5">6.57.3 mm</quantity>
.
<collectingCountry id="F3544A84157BA146126689A8FDB945FF" box="[553,609,1749,1771]" name="Niger" pageId="9" pageNumber="403">Niger</collectingCountry>
,
<collectingCountry id="F3544A84157BA146122489A8FD6D45FF" box="[619,693,1749,1771]" name="Nigeria" pageId="9" pageNumber="403">Nigeria</collectingCountry>
,
<collectingCountry id="F3544A84157BA14612F089A8FCFB45FE" box="[703,803,1749,1770]" name="Cameroon" pageId="9" pageNumber="403">Cameroon</collectingCountry>
..................
<taxonomicName id="4C437197157BA146144689A8FA4445FF" ID-CoL="52RTZ" authority="(Boucomont, 1923)" baseAuthorityName="Boucomont" baseAuthorityYear="1923" box="[1033,1436,1749,1771]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="gaillardi">
<emphasis id="B937D606157BA146144689A8FB0245FE" box="[1033,1242,1749,1770]" italics="true" pageId="9" pageNumber="403">Stiptopodius gaillardi</emphasis>
(Boucomont, 1923)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14610D8898FFA44446A" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D8898FFA444431" blockId="9.[151,1436,1778,1918]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D8898FFA444431" pageId="9" pageNumber="403">
11 Frontal carina strong. Upper surface of the body entirely glabrous, except sometimes on sides of pronotum and on the eighth elytral interstria. Length 6.0
<quantity id="4CBBA7F1157BA14611B4886DFD9E4431" box="[507,582,1808,1830]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="9" pageNumber="403" unit="mm" value="7.2">7.2 mm</quantity>
.
<collectingCountry id="F3544A84157BA146121F886DFD544432" box="[592,652,1808,1830]" name="Sudan" pageId="9" pageNumber="403">Sudan</collectingCountry>
.....................................
<taxonomicName id="4C437197157BA1461417886DFA444431" ID-CoL="6ZVQJ" authority="Branco, 1992" authorityName="Branco" authorityYear="1992" box="[1112,1436,1808,1829]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="B937D606157BA1461417886DFACD4431" box="[1112,1301,1808,1829]" italics="true" pageId="9" pageNumber="403">Stiptopodius glaber</emphasis>
<bibRefCitation id="EFD277E5157BA1461554886DFA444431" author="Branco" box="[1307,1436,1808,1829]" firstAuthor="Branco" pageId="9" pageNumber="403" pagination="101 - 108" refId="ref7139" refString="Branco, T. (1992) Revision des genres du &quot; groupe &quot; Stiptopodius: Donnes complementaires (Coleoptera, Scarabaeidae). Annales de la Societe Entomologique de France, Nouvelle Serie, 28, 101 - 108." type="journal article" year="1992">Branco, 1992</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D88850FA44446A" blockId="9.[151,1436,1778,1918]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D88850FA44446A" pageId="9" pageNumber="403">
Frontal carina obsolete. Upper surface of the body either pilose or glabrous. Pronotum always pilose on the sides; the eighth elytral interstriae varies from pilose or glabrous. Length
<quantity id="4CBBA7F1157BA146134E8837FCAF4474" box="[769,887,1866,1888]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.5" metricValueMin="4.8" pageId="9" pageNumber="403" unit="mm" value="5.65" valueMax="6.5" valueMin="4.8">4.86.5 mm</quantity>
.
<collectingCountry id="F3544A84157BA14613CF8837FC124474" box="[896,970,1866,1888]" name="Nigeria" pageId="9" pageNumber="403">Nigeria</collectingCountry>
,
<collectingCountry id="F3544A84157BA146139B8837FBC84474" box="[980,1040,1866,1888]" name="Benin" pageId="9" pageNumber="403">Benin</collectingCountry>
,
<collectingCountry id="F3544A84157BA14614558837FB454474" box="[1050,1181,1866,1888]" name="Burkina Faso" pageId="9" pageNumber="403">Burkina Faso</collectingCountry>
,
<collectingCountry id="F3544A84157BA14614E98837FB3F4474" box="[1190,1255,1866,1888]" name="Ghana" pageId="9" pageNumber="403">Ghana</collectingCountry>
,
<collectingCountry id="F3544A84157BA14614BE8837FAE54474" box="[1265,1341,1866,1888]" name="Senegal" pageId="9" pageNumber="403">Senegal</collectingCountry>
..........................................................................
<taxonomicName id="4C437197157BA14614588815FA44446A" ID-CoL="6ZW2M" authority="(Boucomont, 1923)" baseAuthorityName="Boucomont" baseAuthorityYear="1923" box="[1047,1436,1896,1918]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="nodieri">
<emphasis id="B937D606157BA14614588815FB024469" box="[1047,1242,1896,1917]" italics="true" pageId="9" pageNumber="403">Stiptopodius nodieri</emphasis>
(Boucomont, 1923)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14610D888F8FA4444C1" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D888F8FA44448E" blockId="9.[151,1436,1925,2005]" box="[151,1436,1925,1947]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D888F8FA44448E" box="[151,1436,1925,1947]" pageId="9" pageNumber="403">12 Clypeal surface only partly granular; genae and vertex not granulose. Integument of pronotum not rugose.............. 13</keyLead>
</paragraph>
<paragraph id="8BFC0A14157BA14610D888DEFA4444C1" blockId="9.[151,1436,1925,2005]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D888DEFA4444C1" pageId="9" pageNumber="403">
Head entirely granular. Integument of pronotum strongly rugose laterally. Length 4.95.0 mm.
<collectingCountry id="F3544A84157BA146143D88DFFB6344AC" box="[1138,1211,1954,1976]" name="Zambia" pageId="9" pageNumber="403">Zambia</collectingCountry>
.......................................................................................
<taxonomicName id="4C437197157BA146146388BDFA4444C1" ID-CoL="6ZVQH" authority="Branco, 2009" authorityName="Branco" authorityYear="2009" box="[1068,1436,1984,2005]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B937D606157BA146146388BDFACD44C1" box="[1068,1301,1984,2005]" italics="true" pageId="9" pageNumber="403">Stiptopodius granulosus</emphasis>
<bibRefCitation id="EFD277E5157BA146155488BDFA4444C1" author="Branco" box="[1307,1436,1984,2005]" firstAuthor="Branco" pageId="9" pageNumber="403" pagination="87 - 91" refId="ref7180" refString="Branco, T. (2009) Notes on &quot; Stiptopodius &quot; Harold, 1871: number of tarsal segments of S. longipedis Branco, 1991, and description of a new species from Zambia (Coleoptera, Scarabaeidae). Boletin Sociedad Entomologica Aragonesa, 44, 87 - 91." type="journal article" year="2009">Branco, 2009</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B712B1157BA14510D888A0FA4443D9" lastPageId="10" lastPageNumber="404" pageId="9" pageNumber="403">
<paragraph id="8BFC0A14157BA14610D888A0FA4444E6" blockId="9.[151,1436,2013,2035]" box="[151,1436,2013,2035]" pageId="9" pageNumber="403">
<keyLead id="30B2A921157BA14610D888A0FA4444E6" box="[151,1436,2013,2035]" pageId="9" pageNumber="403">
13 Head and pronotum of bronze colour. Length
<quantity id="4CBBA7F1157BA14612D588A0FCC844E6" box="[666,784,2013,2035]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" metricValueMax="5.9" metricValueMin="4.5" pageId="9" pageNumber="403" unit="mm" value="5.2" valueMax="5.9" valueMin="4.5">4.55.9 mm</quantity>
.
<collectingCountry id="F3544A84157BA146135688A0FCB044E7" box="[793,872,2013,2035]" name="Somalia" pageId="9" pageNumber="403">Somalia</collectingCountry>
...............
<taxonomicName id="4C437197157BA146146288A0FA4444E6" ID-CoL="6ZW2J" authority="Muller, 1942" authorityName="Muller" authorityYear="1942" box="[1069,1436,2013,2035]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="species" species="glabricollis">
<emphasis id="B937D606157BA146146288A0FAC244E6" box="[1069,1306,2013,2034]" italics="true" pageId="9" pageNumber="403">Stiptopodius glabricollis</emphasis>
Müller, 1942
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC0A141578A14510D88FE7FA4443D9" blockId="10.[151,1436,153,205]" pageId="10" pageNumber="404">
<keyLead id="30B2A9211578A14510D88FE7FA4443D9" pageId="10" pageNumber="404">
Head and pronotum of black colour (female; male unknown). Length
<quantity id="4CBBA7F11578A14513CF8FE7FC1443BB" box="[896,972,154,175]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="404" unit="mm" value="4.3">4.3 mm</quantity>
.
<collectingCountry id="F3544A841578A145139A8FE7FBEC43BB" box="[981,1076,154,175]" name="Botswana" pageId="10" pageNumber="404">Botswana</collectingCountry>
(Savuti)................................................................................
<taxonomicName id="4C4371971578A14513E98FCAFAC743D9" ID-CoL="6ZW2K" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[934,1311,183,205]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="404" phylum="Arthropoda" rank="species" species="savuti" status="sp. nov.">
<emphasis id="B937D6061578A14513E98FCAFB8543D8" box="[934,1117,183,204]" italics="true" pageId="10" pageNumber="404">Stiptopodius savuti</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D1578A14515668FCAFA4443D9" box="[1321,1436,183,205]" pageId="10" pageNumber="404" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>