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<mods:title id="3B587D93FEB229DCDE526E98F9B512FD">Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)</mods:title>
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<treatment id="03C79010FFC5D724FF7079A822FFFC6A" ID-DOI="http://doi.org/10.5281/zenodo.7621927" ID-GBIF-Taxon="209295633" ID-Zenodo-Dep="7621927" LSID="urn:lsid:plazi:treatment:03C79010FFC5D724FF7079A822FFFC6A" httpUri="http://treatment.plazi.org/id/03C79010FFC5D724FF7079A822FFFC6A" lastPageId="59" lastPageNumber="60" pageId="51" pageNumber="52">
<subSubSection id="C374728DFFC5D72CFF7079A82182FD56" box="[151,644,675,702]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8BD12106FFC5D72CFF7079A82182FD56" blockId="51.[151,644,675,737]" box="[151,644,675,702]" pageId="51" pageNumber="52">
<heading id="D099966AFFC5D72CFF7079A82182FD56" bold="true" box="[151,644,675,702]" fontSize="11" level="2" pageId="51" pageNumber="52" reason="4">
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<taxonomicName id="4C6E5A85FFC5D72CFF7079A82103FD56" ID-CoL="5XWVY" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[151,517,675,702]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="51" pageNumber="52" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFC5D72CFF7079A82238FD56" bold="true" box="[151,318,675,702]" italics="true" pageId="51" pageNumber="52">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFC5D72CFEA179A82103FD56" author="M'Intosh, W. C." box="[326,517,675,701]" pageId="51" pageNumber="52" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
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restricted
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<subSubSection id="C374728DFFC5D72CFF7079CC2237FD0A" box="[151,305,711,737]" pageId="51" pageNumber="52" type="description">
<paragraph id="8BD12106FFC5D72CFF7079CC2237FD0A" blockId="51.[151,644,675,737]" box="[151,305,711,737]" pageId="51" pageNumber="52">
<figureCitation id="13553D83FFC5D72CFF7079CC23D8FD0A" box="[151,222,711,737]" captionStart="FIGURE 4" captionStartId="10.[152,255,1827,1851]" captionTargetBox="[154,1433,635,1802]" captionTargetId="figure-246@10.[151,1436,633,1804]" captionTargetPageId="10" captionText="FIGURE 4. Chloeia fusca M’Intosh, 1885, non-type specimen (UF 4368). A. Ventral view. B. Ventral view after body wall removed. C. Same, after Methyl Green staining. D. Same, lower half of digestive tube removed. (AI, anterior intestine; MB, midventral band (nerve chord); MR, muscular ridge; Oe, oesophagus; PI, posterior intestine; R, rectum; St, stomach). All scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7621812" httpUri="https://zenodo.org/record/7621812/files/figure.png" pageId="51" pageNumber="52">Figs 4</figureCitation>
,
<figureCitation id="13553D83FFC5D72CFF0D79CC2237FD0A" box="[234,305,711,737]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStart-2="FIGURE 24" captionStart-3="FIGURE 25" captionStart-4="FIGURE 26" captionStartId-0="53.[152,255,1645,1669]" captionStartId-1="54.[152,255,1889,1913]" captionStartId-2="55.[152,255,1202,1226]" captionStartId-3="56.[152,255,1727,1751]" captionStartId-4="57.[152,255,1755,1779]" captionTargetBox-0="[155,1436,183,1621]" captionTargetBox-1="[152,1435,564,1864]" captionTargetBox-2="[154,1436,183,1175]" captionTargetBox-3="[154,1433,417,1703]" captionTargetBox-4="[178,1411,1140,1731]" captionTargetId-0="figure-18@53.[151,1436,181,1621]" captionTargetId-1="figure-182@54.[151,1436,561,1866]" captionTargetId-2="figure-18@55.[151,1436,181,1178]" captionTargetId-3="figure-123@56.[151,1436,417,1704]" captionTargetId-4="figure-439@57.[176,1411,1137,1732]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionTargetPageId-2="55" captionTargetPageId-3="56" captionTargetPageId-4="57" captionText-0="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." captionText-1="FIGURE 23. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1).A. Largest epitoke (35x6 mm), anterior region, dorsal view. B. Same, posterior region, dorsal view. C. Smallest epitoke (16x4 mm), anterior region, dorsal view. D. Same, posterior region, dorsal view. E. Medium-sized epitoke (22x4 mm), anterior region, dorsal view. F. Same, posterior region, dorsal view. Scale bars: A, B, 1.4 mm; C, D, 0.9 mm; E, F, 1.3 mm." captionText-2="FIGURE 24. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1). A. Largest epitoke, anterior end, right lateral view. B. Chaetiger 3, notochaetae. C. Same, neurochaetal tips. D. Chaetiger 14, harpoon notochaetae and capillary furcates. E. Same, furcates and capillary furcates (inset: capillary furcates furca and tips). F. Smallest epitoke, anterior end, right lateral view. G. Medium-sized epitoke, anterior end, right lateral view. Scale bars: A, 1.2 mm; B, 80 μm; C, 30 μm; D, 35 μm; E, 90 μm; F, 0.6 mm; G, 1 mm." captionText-3="FIGURE 25. Chloeia fusca M’Intosh, 1885, non-type specimens (UF 4368) showing variation in reddish pigmentation and branchial contraction. A) Anterior region, dorsal view of specimen with visible dorsal oblique bands continued to the anterior notopodial surface, and bipinnate branchiae contracted. B) Same, right lateral view. C) Another specimen with oblique bands covered by fully extended bipinnate branchiae. D) Same, right lateral view, with less intense pigmentation along notopidal anterior surface. E) Another specimen with a few reddish chaetae, anterior region dorsal view, bipinnate branchiae covering oblique bands. F) Same, right lateral view showing pigmentation along a few anterior chaetigers on notopodial anterior surface. Scale bars: A, 1.1 mm; B, 1.4 mm; C, 1.8 mm; D, 1.3 mm; E, 0.9 mm; F, 1.5 mm." captionText-4="FIGURE 26. Chloeia fusca M’Intosh, 1885, freshly collected epitokes (CAS 218217). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 10-11, dorsal view. D. Posterior region, slightly oblique dorsal view. Scale bars: A, 1.4 mm; B, 0.7 mm; C, 0.8 mm; D, 1.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7621875" figureDoi-1="http://doi.org/10.5281/zenodo.7621879" figureDoi-2="http://doi.org/10.5281/zenodo.7621883" figureDoi-3="http://doi.org/10.5281/zenodo.7621887" figureDoi-4="http://doi.org/10.5281/zenodo.7621893" httpUri-0="https://zenodo.org/record/7621875/files/figure.png" httpUri-1="https://zenodo.org/record/7621879/files/figure.png" httpUri-2="https://zenodo.org/record/7621883/files/figure.png" httpUri-3="https://zenodo.org/record/7621887/files/figure.png" httpUri-4="https://zenodo.org/record/7621893/files/figure.png" pageId="51" pageNumber="52">22–26</figureCitation>
</paragraph>
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<treatmentCitation id="0ACF0717FFC5D72CFF70780D211AFCF5" author="M'Intosh, W. C." box="[151,540,774,798]" page="15" pageId="51" pageNumber="52" year="1885">
<taxonomicName id="4C6E5A85FFC5D72CFF70780D211AFCF5" ID-CoL="5XWVY" authority="M'Intosh, 1885: 14 - 15" authorityName="M'Intosh" authorityPageNumber="14 - 15" authorityYear="1885" box="[151,540,774,798]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="51" pageNumber="52" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFC5D72CFF70780D2222FCF5" box="[151,292,774,798]" italics="true" pageId="51" pageNumber="52">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFC5D72CFECB780D211AFCF5" author="M'Intosh, W. C." box="[300,540,774,798]" pageId="51" pageNumber="52" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885: 14–15</bibRefCitation>
</taxonomicName>
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, Pl. 2,
<figureCitation id="13553D83FFC5D72CFD82780D21A0FCF5" box="[613,678,774,798]" captionStart="FIGURE 1" captionStartId="7.[152,255,1176,1200]" captionTargetBox="[179,1410,184,1150]" captionTargetId="figure-18@7.[176,1410,181,1152]" captionTargetPageId="7" captionText="FIGURE 1. Living specimens of four Philippine species of Chloeia Savigny in Lamarck, 1818. A. C. amphora Horst, 1910, dorsal view. B. C. bemisae sp. n., pink variant, dorsal view. C. Same, reddish and brownish variants, dorsal view. D. C. flava (Pallas, 1766), oblique right dorsal view. E. C. violacea Horst, 1910, oblique left dorsal view (scales not available; photos: Philippe Poppe; © Philippe &amp; Guido Poppe – www.poppe-images.com)." figureDoi="http://doi.org/10.5281/zenodo.7621800" httpUri="https://zenodo.org/record/7621800/files/figure.png" pageId="51" pageNumber="52">Figs 1</figureCitation>
,
<figureCitation id="13553D83FFC5D72CFD54780D21C7FCF5" box="[691,705,774,798]" captionStart="FIGURE 2" captionStartId="8.[152,255,1518,1542]" captionTargetBox="[155,1434,185,1492]" captionTargetId="figure-28@8.[151,1436,181,1495]" captionTargetPageId="8" captionText="FIGURE 2. Chloeia viridis Schmarda, 1861, non-type specimens (BMNH 1931.6.22.16-18). A. Black form, anterior region, dorsal view. B. Same, chaetigers 10-12, dorsal view (branchiae removed from chaetiger 11). C. Same, posterior region, dorsal view. D. Red form, anterior region, dorsal view. E. Same, chaetigers 12-14, dorsal view (branchiae removed from chaetiger 13). F. Same, posterior region, dorsal view. Scale bars: A, F, 0.9 mm; B, E, 0.7 mm; C, 1 mm; D, 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621804" httpUri="https://zenodo.org/record/7621804/files/figure.png" pageId="51" pageNumber="52">2</figureCitation>
, Pl 1A,
<figureCitation id="13553D83FFC5D72CFCFF780D2060FCF5" box="[792,870,774,798]" captionStart="FIGURE 14" captionStartId="31.[152,255,1789,1813]" captionTargetBox="[152,1435,778,1763]" captionTargetId="figure-274@31.[151,1436,776,1765]" captionTargetPageId="31" captionText="FIGURE 14. Chloeia bistriata Grube, 1868, neotype (SMF 19403). A. Dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Same, posterior region, dorsal view. D. Paraneotype (SMF 30268), anterior end, right lateral view. E. Same, chaetiger 3, notochaetae. F. Same, neurochaetal tips. G. Chaetiger 12, notochaetae (inset: section of denticulate tine). H. Same, neurochaetae. Scale bars: A, 2.5 mm; B, 0.6 mm; C, 0.9 mm; D, 0.5 mm; E, 80 μm; F, 40 μm; G, 100 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621850" httpUri="https://zenodo.org/record/7621850/files/figure.png" pageId="51" pageNumber="52">Figs 14</figureCitation>
,
<figureCitation id="13553D83FFC5D72CFC94780D2088FCF5" box="[883,910,774,798]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="51" pageNumber="52">15</figureCitation>
, Pl. 2A, Figs 1,2;
<treatmentCitation id="0ACF0717FFC5D72CFBAB780C2609FCF5" author="Horst, R." box="[1100,1295,774,798]" page="22" pageId="51" pageNumber="52" year="1912">
<bibRefCitation id="EFFF5CF7FFC5D72CFBAB780C2609FCF5" author="Horst, R." box="[1100,1295,774,798]" pageId="51" pageNumber="52" refId="ref108337" refString="Horst, R. (1912) Polychaeta errantia of the Siboga Expedition. Part 1. Amphinomidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 a, 1 - 43, 10 pls." type="journal volume" year="1912">Horst 1912: 22–24</bibRefCitation>
</treatmentCitation>
, Pl. 7,
<figureCitation id="13553D83FFC5D72CFABF780D2691FCF5" box="[1368,1431,774,798]" captionStart="FIGURE 7" captionStartId="20.[152,255,1856,1880]" captionTargetBox="[151,1435,851,1832]" captionTargetId="figure-294@20.[151,1435,849,1832]" captionTargetPageId="20" captionText="FIGURE 7. Chloeia amoureuxi sp. n., holotype (MMSUCO Amp 2). A. Oblique dorsal view. B. Anterior end, left lateral view. C. Median segments, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 16, harpoon notochaetae. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 0.7 mm; B, 0.05 mm; C, 1.5 mm; D, 160 μm; E, 110 μm; F, 180 μm; G, 30 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621824" httpUri="https://zenodo.org/record/7621824/files/figure.png" pageId="51" pageNumber="52">Fig. 7</figureCitation>
;
<treatmentCitation id="0ACF0717FFC5D72CFF20782E2289FCD6" author="Hartman, O." box="[199,399,805,829]" page="131" pageId="51" pageNumber="52" year="1959">
<bibRefCitation id="EFFF5CF7FFC5D72CFF20782E2289FCD6" author="Hartman, O." box="[199,399,805,829]" pageId="51" pageNumber="52" pagination="1 - 353" refId="ref107655" refString="Hartman, O. (1959) Catalogue of the polychaetous annelids of the World. Part 1. Allan Hancock Foundation Publications, Occasional Paper, 23, 1 - 353." type="journal article" year="1959">Hartman 1959: 131</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD12106FFC5D72CFF70784F2665FCB0" blockId="51.[151,1436,774,860]" box="[151,1379,835,860]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB7E0628FFC5D72CFF70784F2665FCB0" box="[151,1379,835,860]" pageId="51" pageNumber="52">
<treatmentCitation id="0ACF0717FFC5D72CFF70784F2108FCB0" author="Treadwell, A. L." box="[151,526,835,860]" page="1164" pageId="51" pageNumber="52" year="1906">
<taxonomicName id="4C6E5A85FFC5D72CFF70784F2108FCB0" ID-CoL="69WW9" authority=": Treadwell 1906: 1164" authorityName=": Treadwell 1906:" authorityPageNumber="1164" authorityYear="1906" box="[151,526,835,860]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="51" pageNumber="52" phylum="Annelida" rank="species" species="flava">
<emphasis id="B91AFD14FFC5D72CFF70784F2227FCB7" box="[151,289,836,860]" italics="true" pageId="51" pageNumber="52">Chloeia flava</emphasis>
:
<bibRefCitation id="EFFF5CF7FFC5D72CFECA784F2108FCB0" author="Treadwell, A. L." box="[301,526,835,860]" pageId="51" pageNumber="52" pagination="1145 - 1181" refId="ref113334" refString="Treadwell, A. L. (1906) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (1903), 3, 1145 - 1181." type="journal article" year="1906">Treadwell 1906: 1164</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ACF0717FFC5D72CFDFE784F2015FCB7" author="Hartman, O." box="[537,787,835,860]" page="178" pageId="51" pageNumber="52" year="1966">
<bibRefCitation id="EFFF5CF7FFC5D72CFDFE784F2015FCB7" author="Hartman, O." box="[537,787,835,860]" pageId="51" pageNumber="52" pagination="163 - 252" refId="ref107718" refString="Hartman, O. (1966) Polychaetous annelids of the Hawaiian Islands. Occasional Papers of Bernice P. Bishop Museum, 23 (11), 163 - 252." type="journal article" year="1966">Hartman 1966: 178–179</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACF0717FFC5D72CFCF9784F278AFCB0" author="Bailey-Brock, J. H. &amp; Hartman, O." box="[798,1164,835,860]" page="245" pageId="51" pageNumber="52" year="1987">
<bibRefCitation id="EFFF5CF7FFC5D72CFCF9784F278AFCB0" author="Bailey-Brock, J. H. &amp; Hartman, O." box="[798,1164,835,860]" pageId="51" pageNumber="52" pagination="216 - 454" refId="ref102950" refString="Bailey-Brock, J. H. &amp; Hartman, O. (1987) Class Polychaeta. In: Devaney, D. M. &amp; Eldredge, L. G. (Eds.), Reef and Shore Fauna of Hawaii. Bishop Museum Special Publications, 64 (2 - 3), pp. 216 - 454" type="journal article" year="1987">Bailey-Brock &amp; Hartman 1987: 245</bibRefCitation>
</treatmentCitation>
(
<emphasis id="B91AFD14FFC5D72CFB7D784F27C6FCB0" box="[1178,1216,836,859]" italics="true" pageId="51" pageNumber="52">non</emphasis>
(Pallas, 1766)).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFC5D72BFF70788E2058F90A" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BD12106FFC5D72CFF70788E272AFBE1" blockId="51.[151,1437,900,2043]" pageId="51" pageNumber="52">
<emphasis id="B91AFD14FFC5D72CFF70788E2239FC75" bold="true" box="[151,319,900,927]" pageId="51" pageNumber="52">Type material</emphasis>
.
<emphasis id="B91AFD14FFC5D72CFEA8788F2010FC74" bold="true" box="[335,790,900,927]" pageId="51" pageNumber="52">
<collectingCountry id="F3796196FFC5D72CFEA8788F22C3FC75" box="[335,453,900,926]" name="Indonesia" pageId="51" pageNumber="52">Indonesia</collectingCountry>
, Banda Sea, Banda Islands.
</emphasis>
<typeStatus id="54D59FA4FFC5D72CFCF8788F2081FC75" box="[799,903,900,926]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED7FB9C3FFC5D72CFC70788E20EBFC74" box="[919,1005,901,927]" pageId="51" pageNumber="52">BMNH</collectionCode>
1885.12.1.9), H.M.S. Challenger, unnumb. station, depth not indicated (in
<bibRefCitation id="EFFF5CF7FFC5D72CFDB078A32000FC28" author="M'Intosh, W. C." box="[599,774,936,963]" pageId="51" pageNumber="52" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh 1885</bibRefCitation>
: XXV; Admiralty Islands,
<collectingRegion id="49AAEFE4FFC5D72CFBD978A22784FC28" box="[1086,1154,937,963]" country="Indonesia" name="Papua Barat" pageId="51" pageNumber="52">Papua</collectingRegion>
New Guinea but this is wrong,
<quantity id="4C968CE3FFC5D72CFF0878C72251FC0C" box="[239,343,972,999]" metricMagnitude="1" metricUnit="m" metricValue="4.5" metricValueMax="5.4" metricValueMin="3.6" pageId="51" pageNumber="52" unit="m" value="45.0" valueMax="54.0" valueMin="36.0">36–54 m</quantity>
; after
<bibRefCitation id="EFFF5CF7FFC5D72CFE4678C721ABFC0C" author="Tizard, T. H. &amp; Moseley, H. N. &amp; Buchanan, J. Y. &amp; Murray, J." box="[417,685,972,999]" pageId="51" pageNumber="52" pagination="511 - 1110" refId="ref113219" refString="Tizard, T. H., Moseley, H. N., Buchanan, J. Y. &amp; Murray, J. (1885) Narrative of the cruise of H. M. S. Challenger with a general accoung of the scientific results of the expedition. Results of the Voyage of H. M. S. Challenger during the years 1873 - 76, Narrative, 1 (2), 511 - 1110." type="journal article" year="1885">
Tizard
<emphasis id="B91AFD14FFC5D72CFE1578C6212BFC0D" box="[498,557,972,998]" italics="true" pageId="51" pageNumber="52">et al.</emphasis>
1885: 569
</bibRefCitation>
),
<date id="FFD007C6FFC5D72CFD2578C6204CFC0C" box="[706,842,973,999]" pageId="51" pageNumber="52" value="1874-10-01">1 Oct. 1874</date>
(during these days, some people remained in Neira Island, whereas the ship was sent to Ceram trying to help another ship in trouble).
</paragraph>
<paragraph id="8BD12106FFC5D72BFF207F1F2058F90A" blockId="51.[151,1437,900,2043]" lastBlockId="52.[151,1437,151,2013]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<emphasis id="B91AFD14FFC5D72CFF207F1F22B5FBC5" bold="true" box="[199,435,1044,1070]" pageId="51" pageNumber="52">Additional material</emphasis>
.
<emphasis id="B91AFD14FFC5D72CFE257F1F2083FBC4" bold="true" box="[450,901,1044,1071]" pageId="51" pageNumber="52">
<collectingCountry id="F3796196FFC5D72CFE257F1F213EFBC5" box="[450,568,1044,1070]" name="Indonesia" pageId="51" pageNumber="52">Indonesia</collectingCountry>
, Banda Sea, Banda Islands.
</emphasis>
Six specimens (
<collectionCode id="ED7FB9C3FFC5D72CFBDA7F1E2795FBC4" box="[1085,1171,1045,1071]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="51" pageNumber="52" type="Museum">RMNH</collectionCode>
1245),
<collectingRegion id="49AAEFE4FFC5D72CFB0D7F1F2642FBC5" box="[1258,1348,1044,1070]" country="Indonesia" name="Maluku Utara" pageId="51" pageNumber="52">Maluku</collectingRegion>
, RV Siboga Exped., Sta. 240 (Banda anchorage),
<quantity id="4C968CE3FFC5D72CFD9B7F3321D2FBB8" box="[636,724,1080,1107]" metricMagnitude="1" metricUnit="m" metricValue="2.7" metricValueMax="4.5" metricValueMin="0.9" pageId="51" pageNumber="52" unit="m" value="27.0" valueMax="45.0" valueMin="9.0">9–45 m</quantity>
, trawl + dredge + reef expl., black sand and coral,
<date id="FFD007C6FFC5D72CFAF07F32220CFB9C" pageId="51" pageNumber="52" value="1899-11-22" valueMax="1899-12-01" valueMin="1899-11-22">22 Nov. – 1 Dec. 1899</date>
(bipinnate branchiae from chaetiger 5; median antenna as long as caruncle (2/3–1/1); mouth in chaetiger 2;
<quantity id="4C968CE3FFC5D72CFF507F8A2237FB70" box="[183,305,1153,1179]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.2" metricValueMin="1.1" pageId="51" pageNumber="52" unit="mm" value="16.5" valueMax="22.0" valueMin="11.0">11–22 mm</quantity>
long, 2.8–4.0 mm wide, 18–25 chaetigers). Three specimens (
<collectionCode id="ED7FB9C3FFC5D72CFBE67F8A2746FB70" box="[1025,1088,1153,1179]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="51" pageNumber="52">ZMA</collectionCode>
VPol 156.1), epitokes,
<collectingRegion id="49AAEFE4FFC5D72CFAB47F8A2231FB55" country="Indonesia" name="Nusa Tenggara Timur" pageId="51" pageNumber="52">Lesser Sunda Islands</collectingRegion>
, RV Siboga Exped., Sta. 37, Paternoster Islands, Sailus Ketjil (Pulau Sailus Kecil), close to reef,
<quantity id="4C968CE3FFC5D72CFA677FAE23A9FB08" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="51" pageNumber="52" unit="m" value="27.0">27 m</quantity>
, dredge, coral and coral sand, Monaco-trap in
<quantity id="4C968CE3FFC5D72CFD527FC221F8FB08" box="[693,766,1225,1251]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="100.0">100 m</quantity>
, close to reef,
<date id="FFD007C6FFC5D72CFC787FC22767FB08" box="[927,1121,1225,1251]" pageId="51" pageNumber="52" value="1899-03-30" valueMax="1899-03-31" valueMin="1899-03-30">30–31 Mar. 1899</date>
. Ten specimens (
<collectionCode id="ED7FB9C3FFC5D72CFAF97FC2265BFB08" box="[1310,1373,1225,1251]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="51" pageNumber="52">ZMA</collectionCode>
VPol 156.5),
<collectingRegion id="49AAEFE4FFC5D72CFF087FE7224FFAED" box="[239,329,1260,1286]" country="Indonesia" name="Maluku Utara" pageId="51" pageNumber="52">Maluku</collectingRegion>
, RV Siboga Exped., Sta. 240 (Banda anchorage),
<quantity id="4C968CE3FFC5D72CFC987FE720D1FAEC" box="[895,983,1260,1287]" metricMagnitude="1" metricUnit="m" metricValue="2.7" metricValueMax="4.5" metricValueMin="0.9" pageId="51" pageNumber="52" unit="m" value="27.0" valueMax="45.0" valueMin="9.0">9–45 m</quantity>
, trawl + dredge + reef expl., black sand and coral,
<date id="FFD007C6FFC5D72CFEEB7E1A2100FAC0" box="[268,518,1297,1323]" pageId="51" pageNumber="52" value="1899-11-22" valueMax="1899-12-01" valueMin="1899-11-22">22 Nov. – 1 Dec. 1899</date>
(bipinnate branchiae from chaetiger 5; median antennae 4/5 as long as caruncle or slightly longer; mouth in chaetiger 2;
<quantity id="4C968CE3FFC5D72CFDA07E3E21B2FAA4" box="[583,692,1333,1359]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="2.1" metricValueMin="0.8" pageId="51" pageNumber="52" unit="mm" value="14.5" valueMax="21.0" valueMin="8.0">8–21 mm</quantity>
long, 2.5–5.0 mm wide, 18–24 chaetigers). One specimen (
<collectionCode id="ED7FB9C3FFC5D72CFAB97E3E269BFAA4" box="[1374,1437,1333,1359]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="51" pageNumber="52">ZMA</collectionCode>
VPol 156.7),
<collectingRegion id="49AAEFE4FFC5D72CFED77E53228CFA99" box="[304,394,1368,1394]" country="Indonesia" name="Maluku Utara" pageId="51" pageNumber="52">Maluku</collectingRegion>
, RV Siboga Exped., Sta. 240 (Banda anchorage),
<quantity id="4C968CE3FFC5D72CFC5A7E532713FA98" box="[957,1045,1368,1395]" metricMagnitude="1" metricUnit="m" metricValue="2.7" metricValueMax="4.5" metricValueMin="0.9" pageId="51" pageNumber="52" unit="m" value="27.0" valueMax="45.0" valueMin="9.0">9–45 m</quantity>
, trawl + dredge + reef expl., black sand and coral,
<date id="FFD007C6FFC5D72CFEA17E762147FA7C" box="[326,577,1405,1431]" pageId="51" pageNumber="52" value="1899-11-22" valueMax="1899-12-01" valueMin="1899-11-22">22 Nov. – 1 Dec. 1899</date>
(bipinnate branchiae from chaetiger 5; median antenna longer than caruncle; mouth in chaetiger 2;
<quantity id="4C968CE3FFC5D72CFE687EAB22D9FA50" box="[399,479,1440,1467]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="mm" value="15.0">15 mm</quantity>
long,
<quantity id="4C968CE3FFC5D72CFDC07EAB217BFA50" box="[551,637,1440,1467]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="51" pageNumber="52" unit="mm" value="3.5">3.5 mm</quantity>
wide, 23 chaetigers).
<emphasis id="B91AFD14FFC5D72CFC907EAB20EAFA51" bold="true" box="[887,1004,1440,1466]" pageId="51" pageNumber="52">
<collectingCountry id="F3796196FFC5D72CFC907EAB20EAFA51" box="[887,1004,1440,1466]" name="Indonesia" pageId="51" pageNumber="52">Indonesia</collectingCountry>
</emphasis>
. Three specimens (
<collectionCode id="ED7FB9C3FFC5D72CFB217EAA2603FA50" box="[1222,1285,1441,1467]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="51" pageNumber="52">ZMA</collectionCode>
VPol 156.2),
<collectingRegion id="49AAEFE4FFC5D72CFF707ECF23F9FA35" box="[151,255,1476,1502]" country="Indonesia" name="Sulawesi Utara" pageId="51" pageNumber="52">Sulawesi</collectingRegion>
, RV Siboga Exped., Sta. 66 (Bank between islands of Bahuluwang and Tambolungan, S of Saleyer),
<quantity id="4C968CE3FFC5D72CFA837ECE23A9F9E8" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="51" pageNumber="52" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8–10 m</quantity>
, dredge, dead coral,
<taxonomicName id="4C6E5A85FFC5D72CFE727EE321B3F9E9" authority=", Lithothamnion" authorityName="Lithothamnion" box="[405,693,1512,1538]" class="Ulvophyceae" family="Halimedaceae" genus="Halimeda" kingdom="Plantae" order="Bryopsidales" pageId="51" pageNumber="52" phylum="Chlorophyta" rank="genus">
<emphasis id="B91AFD14FFC5D72CFE727EE321B3F9E9" box="[405,693,1512,1538]" italics="true" pageId="51" pageNumber="52">Halimeda, Lithothamnion</emphasis>
</taxonomicName>
,
<date id="FFD007C6FFC5D72CFD247EE22063F9E8" box="[707,869,1513,1539]" pageId="51" pageNumber="52" value="1899-05-07" valueMax="1899-05-08" valueMin="1899-05-07">7–8 May 1899</date>
(body dark purple; bipinnate branchiae from chaetiger 5; median antenna as long as caruncle in
<specimenCount id="9D68EA8FFFC5D72CFD727D062037F9CD" box="[661,817,1548,1575]" pageId="51" pageNumber="52" type="generic">one specimen</specimenCount>
, broken in the others; mouth in chaetiger 2;
<quantity id="4C968CE3FFC5D72CFAF07D07269BF9CC" box="[1303,1437,1548,1575]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.5" metricValueMin="6.3" pageId="51" pageNumber="52" unit="mm" value="6.9" valueMax="7.5" valueMin="6.3">6.3–7.5 mm</quantity>
long, 1.5–2.0 mm wide, 17 chaetigers). One specimen (
<collectionCode id="ED7FB9C3FFC5D72CFCF47D3A2054F9A0" box="[787,850,1585,1611]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="51" pageNumber="52">ZMA</collectionCode>
VPol 156.6),
<collectingRegion id="49AAEFE4FFC5D72CFC127D3B2749F9A1" box="[1013,1103,1584,1610]" country="Indonesia" name="Maluku Utara" pageId="51" pageNumber="52">Maluku</collectingRegion>
, RV Siboga Exped., Sta. 251 (
<geoCoordinate id="EE5A47C1FFC5D72CFF787D5F221AF984" box="[159,284,1620,1647]" degrees="05" direction="south" minutes="28.4" orientation="latitude" pageId="51" pageNumber="52" precision="92" value="-5.4733334">05°28.4´S</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC5D72CFECC7D5E22BEF984" box="[299,440,1621,1647]" degrees="132" direction="east" minutes="00.2" orientation="longitude" pageId="51" pageNumber="52" precision="92" value="132.00333">132°00.2´E</geoCoordinate>
),
<quantity id="4C968CE3FFC5D72CFE297D5E211FF984" box="[462,537,1621,1647]" metricMagnitude="2" metricUnit="m" metricValue="2.04" pageId="51" pageNumber="52" unit="m" value="204.0">204 m</quantity>
, trawl, coral + sand,
<date id="FFD007C6FFC5D72CFCE97D5E209DF984" box="[782,923,1621,1647]" pageId="51" pageNumber="52" value="1899-12-08">8 Dec. 1899</date>
(dorsal bands blackish; body pale; bipinnate branchiae from chaetiger 5; median antenna as long as caruncle; mouth in chaetiger 2;
<quantity id="4C968CE3FFC5D72CFB8D7D7227BCF978" box="[1130,1210,1657,1683]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="51" pageNumber="52" unit="mm" value="19.0">19 mm</quantity>
long,
<quantity id="4C968CE3FFC5D72CFAE67D73265EF978" box="[1281,1368,1656,1683]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="51" pageNumber="52" unit="mm" value="4.5">4.5 mm</quantity>
wide, 24 chaetigers).
<emphasis id="B91AFD14FFC5D72CFEA97D9722D2F95D" bold="true" box="[334,468,1692,1718]" pageId="51" pageNumber="52">
<collectingCountry id="F3796196FFC5D72CFEA97D9722D2F95D" box="[334,468,1692,1718]" name="Philippines" pageId="51" pageNumber="52">Philippines</collectingCountry>
</emphasis>
. One specimen (
<collectionCode id="ED7FB9C3FFC5D72CFD787D9621D1F95C" box="[671,727,1693,1719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
187295), Hearst Philippine Biodiversity Expedition, Luzon,
<collectingRegion id="49AAEFE4FFC5D72CFF707DCA2274F931" box="[151,370,1728,1755]" country="Philippines" name="Batangas" pageId="51" pageNumber="52">Batangas Province</collectingRegion>
, Calumpan Peninsula, Mainit Bubbles dive site (
<geoCoordinate id="EE5A47C1FFC5D72CFC2E7DCB273CF930" box="[969,1082,1728,1755]" degrees="13.69" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="555" value="13.69">13.69° N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC5D72CFBAA7DCA27CEF930" box="[1101,1224,1729,1755]" degrees="120.90" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="120.9">120.90° E</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC5D72CFB3C7DC9269EF930" box="[1243,1432,1729,1757]" degrees="13" direction="north" minutes="41" orientation="latitude" pageId="51" pageNumber="52" precision="1" seconds="24.00" value="13.690001">13°41´24.00″ N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC5D72CFF707DED225EF914" box="[151,344,1765,1793]" degrees="120" direction="east" minutes="54" orientation="longitude" pageId="51" pageNumber="52" precision="1" seconds="00.00" value="120.9">120°54´00.00″ E</geoCoordinate>
), night dive,
<quantity id="4C968CE3FFC5D72CFE0B7DEE2145F914" box="[492,579,1765,1791]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="19.0" metricValueMin="0.0" pageId="51" pageNumber="52" unit="m" value="9.5" valueMax="19.0" valueMin="0.0">0–19 m</quantity>
,
<date id="FFD007C6FFC5D72CFDA87DEE21E2F914" box="[591,740,1765,1791]" pageId="51" pageNumber="52" value="2011-04-29">29 Apr. 2011</date>
, M. Weber, coll. (complete, with some medioposterior dissection for molecular studies; pigmentation pattern retained; body
<quantity id="4C968CE3FFC5D72CFCBD7C0220AEF8C8" box="[858,936,1801,1827]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="51" pageNumber="52" unit="mm" value="20.0">20 mm</quantity>
long,
<quantity id="4C968CE3FFC5D72CFC097C022729F8C8" box="[1006,1071,1801,1827]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="51" pageNumber="52" unit="mm" value="4.0">4 mm</quantity>
wide, 23 chaetigers). One specimen (
<collectionCode id="ED7FB9C3FFC5D72CFF3F7C262216F8AC" box="[216,272,1837,1863]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
214530), Verde Island Passage Expedition, Mindoro, Puerto Galera,
<collectingRegion id="49AAEFE4FFC5D72CFBC37C26278DF8AC" box="[1060,1163,1837,1863]" country="Philippines" name="Batangas" pageId="51" pageNumber="52">Batangas</collectingRegion>
Channel, School Beach (
<geoCoordinate id="EE5A47C1FFC5D72CFF787C5B220DF880" box="[159,267,1872,1899]" degrees="13.52" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="555" value="13.52">13.52° N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC5D72CFEFE7C5B2289F880" box="[281,399,1872,1899]" degrees="120.96" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="120.96">120.96° E</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC5D72CFE7A7C592153F880" box="[413,597,1873,1901]" degrees="13" direction="north" minutes="31" orientation="latitude" pageId="51" pageNumber="52" precision="1" seconds="12.00" value="13.5199995">13°31´12.00″ N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC5D72CFD847C592020F880" box="[611,806,1873,1901]" degrees="120" direction="east" minutes="57" orientation="longitude" pageId="51" pageNumber="52" precision="1" seconds="36.00" value="120.96">120°57´36.00″ E</geoCoordinate>
),
<quantity id="4C968CE3FFC5D72CFCDA7C5A2090F880" box="[829,918,1873,1899]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="3.1" metricValueMin="0.7" pageId="51" pageNumber="52" unit="m" value="19.0" valueMax="31.0" valueMin="7.0">7–31 m</quantity>
,
<date id="FFD007C6FFC5D72CFC437C5A2736F880" box="[932,1072,1872,1899]" pageId="51" pageNumber="52" value="2015-04-08">8 Apr. 2015</date>
, C. Piotrowski, coll. (complete, chaetae bright yellowish, anterior eyes 2× larger than posterior ones; median antenna slightly shorter than caruncle; body
<quantity id="4C968CE3FFC5D72CFF3E7C92222CF858" box="[217,298,1945,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="51" pageNumber="52" unit="mm" value="24.0">24 mm</quantity>
long,
<quantity id="4C968CE3FFC5D72CFE927C9222BFF858" box="[373,441,1945,1971]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="51" pageNumber="52" unit="mm" value="4.0">4 mm</quantity>
wide, 26 chaetigers). One specimen (
<collectionCode id="ED7FB9C3FFC5D72CFC8A7C9220A3F858" box="[877,933,1945,1971]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
214561), Luzon Island,
<collectingRegion id="49AAEFE4FFC5D72CFB257C92269EF859" box="[1218,1432,1944,1971]" country="Philippines" name="Batangas" pageId="51" pageNumber="52">Batangas Province</collectingRegion>
, Verde Island, Blackfish Corner dive site (
<geoCoordinate id="EE5A47C1FFC5D72CFD8C7CB721D0F83C" box="[619,726,1980,2007]" degrees="13.57" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="555" value="13.57">13.57° N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC5D72CFD057CB72050F83C" box="[738,854,1980,2007]" degrees="121.05" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="121.05">121.05° E</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC5D72CFC847CB62712F83C" box="[867,1044,1981,2007]" degrees="13" direction="north" minutes="34" orientation="latitude" pageId="51" pageNumber="52" precision="1" seconds="12.00" value="13.57">13°34'12.00&quot; N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC5D72CFBC77CB627DBF83C" box="[1056,1245,1981,2007]" degrees="121" direction="east" minutes="03" orientation="longitude" pageId="51" pageNumber="52" precision="1" seconds="00.00" value="121.05">121°03'00.00&quot; E</geoCoordinate>
),
<quantity id="4C968CE3FFC5D72CFB157CB6264CF83C" box="[1266,1354,1981,2007]" metricMagnitude="1" metricUnit="m" metricValue="2.05" metricValueMax="3.3" metricValueMin="0.8" pageId="51" pageNumber="52" unit="m" value="20.5" valueMax="33.0" valueMin="8.0">8–33 m</quantity>
,
<date id="FFD007C6FFC5D72CFAB07CB623D4F810" pageId="51" pageNumber="52" value="2015-04-09">9 Apr. 2015</date>
, P.J. Aristorenas, coll. (complete, bent ventrally, dorsal bands discontinuous after extension after ventral bent, chaetae colorless, anterior eyes 6× larger than posterior ones; median antenna as long as caruncle; body
<quantity id="4C968CE3FFC2D72BFAF77B9C2658FF5A" box="[1296,1374,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="52" pageNumber="53" unit="mm" value="12.0">12 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFF707BB023D1FF3E" box="[151,215,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="53" unit="mm" value="3.0">3 mm</quantity>
wide, 24 chaetigers). Four specimens (
<collectionCode id="ED7FB9C3FFC2D72BFD6F7BB021C6FF3E" box="[648,704,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="52" pageNumber="53" type="Museum">CAS</collectionCode>
218217), Visayas, Negros Island, Zamboanguita, off Bask Barangay (
<geoCoordinate id="EE5A47C1FFC2D72BFF297BD4223EFF12" box="[206,312,223,249]" degrees="09" direction="north" minutes="10" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="9.166667">09°10´N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFEA57BD422B0FF12" box="[322,438,223,249]" degrees="123" direction="east" minutes="21" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="123.35">123°21´E</geoCoordinate>
),
<quantity id="4C968CE3FFC2D72BFE2E7BD42119FF12" box="[457,543,223,249]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.4" metricValueMin="0.7" pageId="52" pageNumber="53" unit="m" value="10.5" valueMax="14.0" valueMin="7.0">7–14 m</quantity>
, sand,
<date id="FFD007C6FFC2D72BFD8D7BD421E8FF12" box="[618,750,223,249]" pageId="52" pageNumber="53" value="2016-04-03">3 Apr. 2016</date>
, C. Piotrowski, coll. (complete; anterior eyes 3–4× larger than posterior ones, in darker areas in larger specimens; median antenna longer than caruncle;
<quantity id="4C968CE3FFC2D72BFB667A0827FCFEF6" box="[1153,1274,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.8" metricValueMin="2.7" pageId="52" pageNumber="53" unit="mm" value="37.5" valueMax="48.0" valueMin="27.0">27–48 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFAA77A08269BFEF6" box="[1344,1437,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="52" pageNumber="53" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
wide, 25–28 chaetigers). Seven specimens (
<collectionCode id="ED7FB9C3FFC2D72BFD777A2C21B3FEAA" box="[656,693,295,321]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="52" pageNumber="53" type="Museum">UF</collectionCode>
4368),
<collectingRegion id="49AAEFE4FFC2D72BFCF47A2C274DFEAA" box="[787,1099,295,321]" country="Philippines" name="Mindoro Oriental" pageId="52" pageNumber="53">Oriental Mindoro Province</collectingRegion>
, Mindoro, Puerto Galera, off Sabang Beach (
<geoCoordinate id="EE5A47C1FFC2D72BFEA07A4122B2FE8E" box="[327,436,330,357]" degrees="13.52207" direction="north" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="13.52207">13.52207</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFE277A41213DFE8E" box="[448,571,330,357]" degrees="120.97522" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="120.97522">120.97522</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC2D72BFDA07A4721FBFE8E" box="[583,765,331,359]" degrees="13" direction="north" minutes="31" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="15.42" value="13.520949">13°31´15.42″ N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFCEE7A4720DEFE8E" box="[777,984,331,359]" degrees="121" direction="east" minutes="58" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="30.792" value="121.97522">121°58´30.792″ E</geoCoordinate>
), gentle sand slope with reef heads,
<quantity id="4C968CE3FFC2D72BFA957A4023A9FE62" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="52" pageNumber="53" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6 m</quantity>
,
<date id="FFD007C6FFC2D72BFF5C7A642249FE62" box="[187,335,366,393]" pageId="52" pageNumber="53" value="2015-04-13">13 Apr. 2015</date>
, G. Paulay, coll. (all with pharynx partially exposed; anterior eyes 5–6× larger than posterior ones; longitudinal bands reddish, of similar width throughout dorsum; some specimens with dorsal oblique reddish bands continued towards parapodial anterior surface, usually covered by bipinnate branchiae; chaetae reddish to pale yellowish, sometimes with a wide median golden band; body
<quantity id="4C968CE3FFC2D72BFCD27AD020A9FE1E" box="[821,943,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="52" pageNumber="53" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">16–19 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFC1F7AD12750FE1E" box="[1016,1110,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="52" pageNumber="53" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
wide, 22–23 chaetigers;
<specimenCount id="9D68EA8FFFC2D72BFA937AD02207FDF2" pageId="52" pageNumber="53" type="generic">one specimen</specimenCount>
dissected for anatomy). Two specimens (
<collectionCode id="ED7FB9C3FFC2D72BFD367AF421F0FDF2" box="[721,758,511,537]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="52" pageNumber="53" type="Museum">UF</collectionCode>
4392),
<collectingRegion id="49AAEFE4FFC2D72BFCAA7AF42779FDF2" box="[845,1151,511,537]" country="Philippines" name="Mindoro Oriental" pageId="52" pageNumber="53">Oriental Mindoro Province</collectingRegion>
, Mindoro, Puerto Galera, Sabang Beach (
<geoCoordinate id="EE5A47C1FFC2D72BFEA4792922B6FDD6" box="[323,432,546,573]" degrees="13.52095" direction="north" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="13.52095">13.52095</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFE5C79282130FDD6" box="[443,566,547,573]" degrees="120.97439" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="120.97439">120.97439</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC2D72BFDD8792F21F5FDD6" box="[575,755,547,575]" degrees="13" direction="north" minutes="31" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="15.42" value="13.520949">13°31´15.42″ N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFD1A792F20CCFDD6" box="[765,970,547,575]" degrees="121" direction="east" minutes="58" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="27.804" value="121.97439">121°58´27.804″ E</geoCoordinate>
), sand, seagrass flat,
<quantity id="4C968CE3FFC2D72BFB54792827FDFDD6" box="[1203,1275,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="52" pageNumber="53" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
,
<date id="FFD007C6FFC2D72BFAE17928269EFDD6" box="[1286,1432,546,573]" pageId="52" pageNumber="53" value="2015-04-23">23 Apr. 2015</date>
, G. Paulay, coll. (shorter without posterior end, showing branching gonads each branch slightly wider than corresponding blood vessel; larger complete, bent ventrally; anterior eyes 3–4× larger than posterior ones; black anterior prostomial area split in two halves; median antenna without tips, shorter than caruncle;
<quantity id="4C968CE3FFC2D72BFB9C798427F0FD42" box="[1147,1270,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="4.3" metricValueMin="3.1" pageId="52" pageNumber="53" unit="mm" value="37.0" valueMax="43.0" valueMin="31.0">31–43 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFAD97984269BFD42" box="[1342,1437,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="52" pageNumber="53" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
wide, 18–27 chaetigers). One specimen (
<collectionCode id="ED7FB9C3FFC2D72BFD8479B8218EFD26" box="[611,648,691,717]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="52" pageNumber="53" type="Museum">UF</collectionCode>
4394),
<collectingRegion id="49AAEFE4FFC2D72BFD0779B82715FD26" box="[736,1043,691,717]" country="Philippines" name="Mindoro Oriental" pageId="52" pageNumber="53">Oriental Mindoro Province</collectingRegion>
, Mindoro, Puerto Galera, off Point West of Bayanan Beach (
<geoCoordinate id="EE5A47C1FFC2D72BFE5779DD2108FD1A" box="[432,526,726,753]" degrees="13.5118" direction="north" orientation="latitude" pageId="52" pageNumber="53" precision="5" value="13.5118">13.5118</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFDFE79DC2180FD1A" box="[537,646,727,753]" degrees="120.9088" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="5" value="120.9088">120.9088</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC2D72BFD7779DC2047FD1A" box="[656,833,727,753]" degrees="13" direction="north" minutes="30" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="42.48" value="13.5118">13°30'42.48&quot; N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFCAB79DD2701FD1A" box="[844,1031,726,753]" degrees="120" direction="east" minutes="54" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="31.68" value="120.9088">120°54'31.68&quot; E</geoCoordinate>
),
<quantity id="4C968CE3FFC2D72BFBFC79DC2786FD1A" box="[1051,1152,727,753]" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="52" pageNumber="53" unit="m" value="11.5" valueMax="13.0" valueMin="10.0">10–13 m</quantity>
, sand slope with few corals,
<date id="FFD007C6FFC2D72BFF2179F02259FCFE" box="[198,351,762,789]" pageId="52" pageNumber="53" value="2015-04-28">28 Apr. 2015</date>
, G. Paulay, coll. (atoke; thin transverse black intersegmental bands in addition to other typical banding; anterior eyes 3–4× larger than posterior ones; anterior prostomial area barely split; median antenna slightly longer than caruncle;
<quantity id="4C968CE3FFC2D72BFE40784822F0FCB6" box="[423,502,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="52" pageNumber="53" unit="mm" value="29.0">29 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFDDA78482178FCB6" box="[573,638,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="mm" value="6.0">6 mm</quantity>
wide, 26 chaetigers). Two specimens (
<collectionCode id="ED7FB9C3FFC2D72BFBC878482768FCB6" box="[1071,1134,835,861]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="52" pageNumber="53">ZMA</collectionCode>
VPol 156.3),
<collectingRegion id="49AAEFE4FFC2D72BFAEC78482638FCB6" box="[1291,1342,835,861]" country="Philippines" name="Sulu" pageId="52" pageNumber="53">Sulu</collectingRegion>
Islands, anchorage off North Ubian, RV Siboga Exped., Sta. 99 (
<geoCoordinate id="EE5A47C1FFC2D72BFCC2786D20A0FC6A" box="[805,934,870,897]" degrees="06" direction="north" minutes="07.5" orientation="latitude" pageId="52" pageNumber="53" precision="92" value="6.125">06°07.5’ N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFC53786C272CFC6A" box="[948,1066,871,897]" degrees="120" direction="east" minutes="26" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="120.433334">120°26’ E</geoCoordinate>
),
<quantity id="4C968CE3FFC2D72BFBA5786C27AFFC6A" box="[1090,1193,871,897]" metricMagnitude="1" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="52" pageNumber="53" unit="m" value="19.5" valueMax="23.0" valueMin="16.0">16–23 m</quantity>
, dredge and tow-net, lithothamnion bottom,
<date id="FFD007C6FFC2D72BFE7D78802153FC4E" box="[410,597,907,933]" pageId="52" pageNumber="53" value="1899-06-28" valueMax="1899-06-30" valueMin="1899-06-28">28–30 Jun. 1899</date>
(bipinnate branchiae from chaetiger 5; median antenna longer than caruncle; mouth in chaetiger 2;
<quantity id="4C968CE3FFC2D72BFE5E78A42125FC22" box="[441,547,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.1" metricValueMin="0.9" pageId="52" pageNumber="53" unit="mm" value="15.0" valueMax="21.0" valueMin="9.0">9–21 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFD8F78A421C2FC22" box="[616,708,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="52" pageNumber="53" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
wide, 17–24 chaetigers). One specimen (
<collectionCode id="ED7FB9C3FFC2D72BFB6878A427C8FC22" box="[1167,1230,943,969]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="52" pageNumber="53">ZMA</collectionCode>
VPol 156.4),
<collectingRegion id="49AAEFE4FFC2D72BFA8E78A4269AFC22" box="[1385,1436,943,969]" country="Philippines" name="Sulu" pageId="52" pageNumber="53">Sulu</collectingRegion>
Islands, RV Siboga Exped., Sta. 104 (
<collectingRegion id="49AAEFE4FFC2D72BFDA778D82173FC06" box="[576,629,979,1005]" country="Philippines" name="Sulu" pageId="52" pageNumber="53">Sulu</collectingRegion>
Harbour),
<quantity id="4C968CE3FFC2D72BFD1578D82028FC06" box="[754,814,979,1005]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="52" pageNumber="53" unit="m" value="14.0">14 m</quantity>
, dredge, sand,
<date id="FFD007C6FFC2D72BFC3F78D82777FC06" box="[984,1137,979,1005]" pageId="52" pageNumber="53" value="1899-07-02" valueMax="1899-07-03" valueMin="1899-07-02">2–3 Jul. 1899</date>
(pale purple, dorsal bands almost completely faded off; bipinnate branchiae from chaetiger 5; median antenna longer than caruncle; mouth in chaetiger 2; body
<quantity id="4C968CE3FFC2D72BFE807F1122BEFBDE" box="[359,440,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="52" pageNumber="53" unit="mm" value="15.0">15 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFDE57F102143FBDE" box="[514,581,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="53" unit="mm" value="3.0">3 mm</quantity>
wide, 21 chaetigers).
<emphasis id="B91AFD14FFC2D72BFCA37F102729FBDF" bold="true" box="[836,1071,1050,1077]" pageId="52" pageNumber="53">
<collectingCountry id="F3796196FFC2D72BFCA37F102729FBDF" box="[836,1071,1050,1077]" name="Papua New Guinea" pageId="52" pageNumber="53">Papua New Guinea</collectingCountry>
</emphasis>
. One specimen (
<collectionCode id="ED7FB9C3FFC2D72BFB167F102637FBDE" box="[1265,1329,1051,1077]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="52" pageNumber="53">NTM</collectionCode>
W5050), Horseshoe Reef,
<quantity id="4C968CE3FFC2D72BFEB17F342297FBB2" box="[342,401,1087,1113]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="52" pageNumber="53" unit="m" value="10.0">10 m</quantity>
,
<date id="FFD007C6FFC2D72BFE7C7F342115FBB2" box="[411,531,1087,1113]" pageId="52" pageNumber="53" value="1980-08">Aug. 1980</date>
, N. Coleman, coll. (colorless; anterior prostomial area blackish; eyes black, anterior ones 3–4× larger than posterior ones; body
<quantity id="4C968CE3FFC2D72BFD2F7F692029FB96" box="[712,815,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="52" pageNumber="53" unit="mm" value="17.5">17.5 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFC9A7F6820C7FB96" box="[893,961,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="52" pageNumber="53" unit="mm" value="4.0">4 mm</quantity>
wide, 22 chaetigers).
<emphasis id="B91AFD14FFC2D72BFB217F692665FB96" bold="true" box="[1222,1379,1122,1149]" pageId="52" pageNumber="53">Line Islands.</emphasis>
One specimen (
<collectionCode id="ED7FB9C3FFC2D72BFEF77F8C2233FB4A" box="[272,309,1159,1185]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="52" pageNumber="53" type="Museum">UF</collectionCode>
576), Kiritimati, just N of main reef passage, West side of atoll, outer reef slope (
<geoCoordinate id="EE5A47C1FFC2D72BFB217F8C2611FB4A" box="[1222,1303,1159,1185]" degrees="1.9841" direction="north" orientation="latitude" pageId="52" pageNumber="53" precision="5" value="1.9841">1.9841</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFAC57F8D269EFB4A" box="[1314,1432,1158,1185]" degrees="157.4819" direction="west" orientation="longitude" pageId="52" pageNumber="53" precision="5" value="-157.4819">-157.4819</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC2D72BFF707FA7224BFB2E" box="[151,333,1195,1223]" degrees="01" direction="north" minutes="59" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="02.76" value="1.9841">01°59´02.76″ N</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFEBF7FA72125FB2E" box="[344,547,1195,1223]" degrees="157" direction="west" minutes="28" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="54.84" value="-157.48189">157°28´54.84″ W</geoCoordinate>
), outer reef slope,
<quantity id="4C968CE3FFC2D72BFD137FA0205CFB2E" box="[756,858,1195,1221]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="52" pageNumber="53" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10–12 m</quantity>
, from
<taxonomicName id="4C6E5A85FFC2D72BFC437FA02714FB2E" box="[932,1042,1195,1221]" class="Ulvophyceae" family="Halimedaceae" genus="Halimeda" kingdom="Plantae" order="Bryopsidales" pageId="52" pageNumber="53" phylum="Chlorophyta" rank="genus">
<emphasis id="B91AFD14FFC2D72BFC437FA02714FB2E" box="[932,1042,1195,1221]" italics="true" pageId="52" pageNumber="53">Halimeda</emphasis>
</taxonomicName>
sample,
<date id="FFD007C6FFC2D72BFB907FA12603FB2E" box="[1143,1285,1194,1221]" pageId="52" pageNumber="53" value="2005-08-05">5 Aug. 2005</date>
, G. Paulay &amp; N. Knowlton, coll. (without posterior end; median antenna twisted, about as long as caruncle; anterior eyes 3–4× larger than posterior ones;
<quantity id="4C968CE3FFC2D72BFE257FF82117FAE6" box="[450,529,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="52" pageNumber="53" unit="mm" value="13.0">13 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFDB17FF82191FAE6" box="[598,663,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="52" pageNumber="53" unit="mm" value="4.0">4 mm</quantity>
wide, 16 chaetigers).
<emphasis id="B91AFD14FFC2D72BFC6A7FF8274AFAE6" bold="true" box="[909,1100,1267,1293]" pageId="52" pageNumber="53">Loyalty Islands.</emphasis>
Two specimens (
<collectionCode id="ED7FB9C3FFC2D72BFAE97FF82663FAE6" box="[1294,1381,1267,1293]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="52" pageNumber="53">MNHN</collectionCode>
Musorstom P1231), Sta. 1231 (
<geoCoordinate id="EE5A47C1FFC2D72BFE2B7E1C2140FADA" box="[460,582,1303,1329]" degrees="20" direction="south" minutes="31.2" orientation="latitude" pageId="52" pageNumber="53" precision="92" value="-20.52">20°31.2´S</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFDB67E1C21DDFADA" box="[593,731,1303,1329]" degrees="166" direction="east" minutes="22.9" orientation="longitude" pageId="52" pageNumber="53" precision="92" value="166.38167">166°22.9´E</geoCoordinate>
),
<quantity id="4C968CE3FFC2D72BFD087E1C202FFADA" box="[751,809,1303,1329]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="52" pageNumber="53" unit="m" value="23.0">23 m</quantity>
,
<date id="FFD007C6FFC2D72BFCD37E1C20BEFADA" box="[820,952,1303,1329]" pageId="52" pageNumber="53" value="1992-09-03">3 Sep. 1992</date>
(slightly damaged, depressed; dorsal bands fading off; anterior eyes 6–8× larger than posterior ones; body
<quantity id="4C968CE3FFC2D72BFC977E3020EDFABE" box="[880,1003,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="52" pageNumber="53" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
long, 3.5–4.0 mm wide, 22–24 chaetigers).
<emphasis id="B91AFD14FFC2D72BFF0A7E5422C7FA93" bold="true" box="[237,449,1374,1401]" pageId="52" pageNumber="53">
<collectingCountry id="F3796196FFC2D72BFF0A7E5422BBFA93" box="[237,445,1374,1401]" name="French Polynesia" pageId="52" pageNumber="53">French Polynesia</collectingCountry>
.
</emphasis>
One specimen (
<collectionCode id="ED7FB9C3FFC2D72BFD917E54219DFA92" box="[630,667,1375,1401]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="52" pageNumber="53" type="Museum">UF</collectionCode>
2921),
<collectingRegion id="49AAEFE4FFC2D72BFD157E542099FA92" box="[754,927,1375,1401]" inferredBySuffix="true" pageId="52" pageNumber="53">Society Islands</collectingRegion>
, Moorea Island, Moorea, off Motu Ahi, north of Afareitu Pass (
<geoCoordinate id="EE5A47C1FFC2D72BFEB17E8922CBFA76" box="[342,461,1410,1437]" degrees="17.55258" direction="south" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="-17.55258">-17.55258</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFE307E88215CFA76" box="[471,602,1411,1437]" degrees="149.77347" direction="west" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="-149.77347">-149.77347</geoCoordinate>
;
<geoCoordinate id="EE5A47C1FFC2D72BFD827E8F2027FA76" box="[613,801,1411,1439]" degrees="17" direction="south" minutes="33" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="09.288" value="-17.55258">17°33´09.288″ S</geoCoordinate>
,
<geoCoordinate id="EE5A47C1FFC2D72BFCCA7E8F2702FA76" box="[813,1028,1411,1439]" degrees="149" direction="west" minutes="46" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="24.492" value="-149.77347">149°46´24.492″ W</geoCoordinate>
), deep reef slope, large rubble,
<quantity id="4C968CE3FFC2D72BFABA7E892691FA76" box="[1373,1431,1410,1437]" metricMagnitude="1" metricUnit="m" metricValue="5.7" pageId="52" pageNumber="53" unit="m" value="57.0">57 m</quantity>
,
<date id="FFD007C6FFC2D72BFF707EAC222EFA2A" box="[151,296,1447,1473]" pageId="52" pageNumber="53" value="2012-01-27">27 Jan. 2012</date>
, R. Whitton, coll. (anterior fragment; anterior eyes 3× larger than posterior ones; two dorsal longitudinal bands;
<quantity id="4C968CE3FFC2D72BFEF77EC12263FA0E" box="[272,357,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="52" pageNumber="53" unit="mm" value="3.5">3.5 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFE4C7EC02106FA0E" box="[427,512,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="52" pageNumber="53" unit="mm" value="1.2">1.2 mm</quantity>
wide, 9 chaetigers).
<emphasis id="B91AFD14FFC2D72BFD007EC1204BFA0F" bold="true" box="[743,845,1482,1508]" pageId="52" pageNumber="53">Hawai’i.</emphasis>
One specimen (
<collectionCode id="ED7FB9C3FFC2D72BFBE67EC02728FA0E" box="[1025,1070,1483,1509]" country="United Kingdom" name="Bristol Museum" pageId="52" pageNumber="53">BM</collectionCode>
R3441), KOK Pisces V, Expedition HURL-09, Sta. P5-716,
<date id="FFD007C6FFC2D72BFE307EE4216EF9E2" box="[471,616,1519,1545]" pageId="52" pageNumber="53" value="2009-02-22">22 Feb. 2009</date>
(no further data; complete, juvenile, dorsal pigmentation with two irregular longitudinal lines, paler than in more recent specimens; dorsal cirri purple, bipinnate branchial tips purple; chaetae colorless; Prostomium with anterior brownish area as a half-moon; eyes blackish, anterior eyes about 6× larger than posterior ones; median antenna slightly longer than caruncle;
<quantity id="4C968CE3FFC2D72BFC3E7D50271DF99E" box="[985,1051,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="mm" value="6.0">6 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFB827D5027AEF99E" box="[1125,1192,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="52" pageNumber="53" unit="mm" value="2.0">2 mm</quantity>
wide, 17 chaetigers). One specimen (
<collectionCode id="ED7FB9C3FFC2D72BFEA37D74226FF972" box="[324,361,1663,1689]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="52" pageNumber="53" type="Museum">UF</collectionCode>
418), Maui Island, Buzzs Wharf, caught swimming,
<date id="FFD007C6FFC2D72BFC507D752748F972" box="[951,1102,1662,1689]" pageId="52" pageNumber="53" value="2005-05-25">25 May 2005</date>
, C. Pittman &amp; D. Kheler, coll. (epitoke, complete, chaetae yellowish; anterior eyes 7–8× larger than posterior ones; median antenna without tip, longer than caruncle;
<quantity id="4C968CE3FFC2D72BFE6C7DCD22DCF90A" box="[395,474,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="52" pageNumber="53" unit="mm" value="55.0">55 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFDC77DCC2168F90A" box="[544,622,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="52" pageNumber="53" unit="mm" value="11.0">11 mm</quantity>
wide, 31 chaetigers).
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFC2D72BFF207DE12188F8A6" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8BD12106FFC2D72BFF207DE12188F8A6" blockId="52.[151,1437,151,2013]" pageId="52" pageNumber="53">
<emphasis id="B91AFD14FFC2D72BFF207DE1223CF8EF" bold="true" box="[199,314,1770,1796]" pageId="52" pageNumber="53">Diagnosis</emphasis>
.
<taxonomicName id="4C6E5A85FFC2D72BFEAE7DE022A4F8EE" authorityName="Savigny" authorityYear="1818" box="[329,418,1771,1797]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="52" pageNumber="53" phylum="Annelida" rank="genus">
<emphasis id="B91AFD14FFC2D72BFEAE7DE022A4F8EE" box="[329,418,1771,1797]" italics="true" pageId="52" pageNumber="53">Chloeia</emphasis>
</taxonomicName>
with bipinnate branchiae from chaetiger 5, progressively smaller posteriorly; dorsum with two longitudinal bands; caruncle pale; lips darker than adjacent ventral areas; notochaetae include furcates and furcate harpoon-chaetae; neurochaetae furcates.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFC2D725FF207C5D2143FCB6" lastPageId="58" lastPageNumber="59" pageId="52" pageNumber="53" type="description">
<paragraph id="8BD12106FFC2D72BFF207C5D204FF836" blockId="52.[151,1437,151,2013]" pageId="52" pageNumber="53">
<emphasis id="B91AFD14FFC2D72BFF207C5D2257F89B" bold="true" box="[199,337,1878,1904]" pageId="52" pageNumber="53">Description</emphasis>
.
<typeStatus id="54D59FA4FFC2D72BFEB87C5C22C1F89A" box="[351,455,1879,1905]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
(BMNH 1885.12.1.9), almost without chaetae, with a midventral dissection along chaetiger 4–19, cut into two portions (
<figureCitation id="13553D83FFC2D72BFDF67C70219AF87E" box="[529,668,1915,1941]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="52" pageNumber="53">Fig. 22A, D</figureCitation>
): anterior region
<quantity id="4C968CE3FFC2D72BFCBA7C712098F87E" box="[861,926,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="52" pageNumber="53" unit="mm" value="5.0">5 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFC027C702720F87E" box="[997,1062,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="52" pageNumber="53" unit="mm" value="4.0">4 mm</quantity>
wide, 8 chaetigers, and a median to posterior region
<quantity id="4C968CE3FFC2D72BFE8A7C9522D7F852" box="[365,465,1950,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="52" pageNumber="53" unit="mm" value="10.5">10.5 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFDF07C94215EF852" box="[535,600,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="52" pageNumber="53" unit="mm" value="4.0">4 mm</quantity>
wide, 14 chaetigers; body narrow fusiform,
<quantity id="4C968CE3FFC2D72BFBAD7C9527A8F852" box="[1098,1198,1950,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="52" pageNumber="53" unit="mm" value="15.5">15.5 mm</quantity>
long,
<quantity id="4C968CE3FFC2D72BFB137C942633F852" box="[1268,1333,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="52" pageNumber="53" unit="mm" value="4.0">4 mm</quantity>
wide, 22 chaetigers (
<quantity id="4C968CE3FFC2D72BFEFF7CC8226EF836" box="[280,360,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="52" pageNumber="53" unit="mm" value="18.0">18 mm</quantity>
long, 23 segments in original description).
</paragraph>
<caption id="DF11718EFFC3D72AFF7F7D662076F8FE" ID-DOI="http://doi.org/10.5281/zenodo.7621875" ID-Zenodo-Dep="7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="53" pageNumber="54" startId="53.[152,255,1645,1669]" targetBox="[155,1436,183,1621]" targetPageId="53">
<paragraph id="8BD12106FFC3D72AFF7F7D662076F8FE" blockId="53.[151,1437,1645,1813]" pageId="53" pageNumber="54">
<emphasis id="B91AFD14FFC3D72AFF7F7D662218F96E" bold="true" box="[152,286,1645,1669]" pageId="53" pageNumber="54">FIGURE 22</emphasis>
.
<taxonomicName id="4C6E5A85FFC3D72AFECD7D66215AF96E" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[298,604,1645,1669]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="53" pageNumber="54" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFC3D72AFECD7D6622B3F96E" box="[298,437,1645,1669]" italics="true" pageId="53" pageNumber="54">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFC3D72AFE5C7D66215AF96E" author="M'Intosh, W. C." box="[443,604,1645,1669]" pageId="53" pageNumber="54" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
. A. Holotype (BMNH 1885.12.1.9). A. Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm.
</paragraph>
</caption>
<paragraph id="8BD12106FFC3D72AFF207C4822E7F84E" blockId="53.[151,1437,1859,2030]" pageId="53" pageNumber="54">
<typeStatus id="54D59FA4FFC3D72AFF207C482229F8B6" box="[199,303,1859,1885]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
brownish; anterior prostomial margin blackish; middorsal bands brownish, continued throughout body, slightly expanded medially (
<figureCitation id="13553D83FFC3D72AFDFE7C6C2186F86A" box="[537,640,1895,1921]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="53" pageNumber="54">Fig. 22C</figureCitation>
). Dorsal cirri with cirrophores pale, cirrostyles dark purple. Venter pale with a midventral paler band.
</paragraph>
<paragraph id="8BD12106FFC3D729FF207CA422A0FF3E" blockId="53.[151,1437,1859,2030]" lastBlockId="54.[151,1437,151,537]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
Prostomium anteriorly entire. Eyes blackish, separate, anterior eyes 3–4× larger than posterior ones (
<figureCitation id="13553D83FFC3D72AFACE7CA42696F822" box="[1321,1424,1967,1993]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="53" pageNumber="54">Fig. 22B</figureCitation>
). Median antenna pale, inserted at anterior caruncular margin, slightly longer than caruncle, 2× longer than lateral antennae. Lateral antennae blackish, bases separate from each other, 2× longer than palps. Mouth ventral on chaetiger 2. Pharynx not exposed.
</paragraph>
<paragraph id="8BD12106FFC0D729FF207BD42781FEF6" blockId="54.[151,1437,151,537]" pageId="54" pageNumber="55">Caruncle pale, sigmoid, trilobed, tapered, reaching chaetiger 5. Median ridge plicate, with about 18 vertical folds, partially concealing lateral lobes. Lateral lobes narrow, with about 28 vertical folds.</paragraph>
<paragraph id="8BD12106FFC0D729FF207A2C20F3FE62" blockId="54.[151,1437,151,537]" pageId="54" pageNumber="55">Bipinnate branchiae from chaetiger 5, continued throughout body, partially eroded, parallel along body; progressively larger to chaetiger 9, about half as long as successive segments, decreasing in size posteriorly (becoming as long as successive segments); median segments with 5–6 lateral branches.</paragraph>
<paragraph id="8BD12106FFC0D729FF207A98271DFDF2" blockId="54.[151,1437,151,537]" pageId="54" pageNumber="55">Parapodia biramous, notopodia with purple cirriform branchiae along chaetigers 1–4, 2/3 as long as dorsal cirri. Dorsal cirri 3–4× longer than bipinnate branchiae along median chaetigers, 5–6× longer in posterior chaetigers. Second ventral cirri with cirrophores 2× longer and wider, and cirrostyle 3× longer than adjacent ones, directed dorsally. Other ventral cirri directed ventrolaterally, as long as 1.5× subsequent segments.</paragraph>
<caption id="DF11718EFFC0D729FF7F7C6A21C5F80E" ID-DOI="http://doi.org/10.5281/zenodo.7621879" ID-Zenodo-Dep="7621879" httpUri="https://zenodo.org/record/7621879/files/figure.png" pageId="54" pageNumber="55" startId="54.[152,255,1889,1913]" targetBox="[152,1435,564,1864]" targetPageId="54">
<paragraph id="8BD12106FFC0D729FF7F7C6A21C5F80E" blockId="54.[151,1437,1889,2021]" pageId="54" pageNumber="55">
<emphasis id="B91AFD14FFC0D729FF7F7C6A221BF892" bold="true" box="[152,285,1889,1913]" pageId="54" pageNumber="55">FIGURE 23</emphasis>
.
<taxonomicName id="4C6E5A85FFC0D729FECE7C6A215EF892" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[297,600,1889,1913]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="54" pageNumber="55" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFC0D729FECE7C6A22B2F892" box="[297,436,1889,1913]" italics="true" pageId="54" pageNumber="55">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFC0D729FE5E7C6A215EF892" author="M'Intosh, W. C." box="[441,600,1889,1913]" pageId="54" pageNumber="55" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
, epitokes (ZMA VPol 156.1). A. Largest epitoke (35x6 mm), anterior region, dorsal view. B. Same, posterior region, dorsal view. C. Smallest epitoke (16x4 mm), anterior region, dorsal view. D. Same, posterior region, dorsal view. E. Medium-sized epitoke (22x4 mm), anterior region, dorsal view. F. Same, posterior region, dorsal view. Scale bars: A, B, 1.4 mm; C, D, 0.9 mm; E, F, 1.3 mm.
</paragraph>
</caption>
<caption id="DF11718EFFC1D728FF7F7FB9222BFAB1" ID-DOI="http://doi.org/10.5281/zenodo.7621883" ID-Zenodo-Dep="7621883" httpUri="https://zenodo.org/record/7621883/files/figure.png" pageId="55" pageNumber="56" startId="55.[152,255,1202,1226]" targetBox="[154,1436,183,1175]" targetPageId="55">
<paragraph id="8BD12106FFC1D728FF7F7FB9222BFAB1" blockId="55.[151,1437,1202,1370]" pageId="55" pageNumber="56">
<emphasis id="B91AFD14FFC1D728FF7F7FB92219FB21" bold="true" box="[152,287,1202,1226]" pageId="55" pageNumber="56">FIGURE 24</emphasis>
.
<taxonomicName id="4C6E5A85FFC1D728FECA7FB92164FB21" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[301,610,1202,1226]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="55" pageNumber="56" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFC1D728FECA7FB922BFFB21" box="[301,441,1202,1226]" italics="true" pageId="55" pageNumber="56">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFC1D728FE267FB92164FB21" author="M'Intosh, W. C." box="[449,610,1202,1226]" pageId="55" pageNumber="56" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
, epitokes (ZMA VPol 156.1). A. Largest epitoke, anterior end, right lateral view. B. Chaetiger 3, notochaetae. C. Same, neurochaetal tips. D. Chaetiger 14, harpoon notochaetae and capillary furcates. E. Same, furcates and capillary furcates (inset: capillary furcates furca and tips). F. Smallest epitoke, anterior end, right lateral view. G. Medium-sized epitoke, anterior end, right lateral view. Scale bars: A, 1.2 mm; B, 80 μm; C, 30 μm; D, 35 μm; E, 90 μm; F, 0.6 mm; G, 1 mm.
</paragraph>
</caption>
<paragraph id="8BD12106FFC1D728FF207E832198F991" blockId="55.[151,1437,1416,2019]" pageId="55" pageNumber="56">
<materialsCitation id="3B062B5BFFC1D728FF207E83219FF991" ID-GBIF-Occurrence="4029882311" collectionCode="RMNH" pageId="55" pageNumber="56" specimenCode="RMNH 1245" specimenCount="1" typeStatus="holotype">
Chaetae most broken, a few remaining (not removed for avoiding further damage). Complete chaetae (after largest topotype
<specimenCode id="DBC8897DFFC1D728FEA87EA622E4FA2C" box="[335,482,1452,1479]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="55" pageNumber="56" type="Museum">RMNH 1245</specimenCode>
) with distal hoods. Anterior notochaetae furcates (
<figureCitation id="13553D83FFC1D728FBE87EA72775FA2D" box="[1039,1139,1452,1478]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="55" pageNumber="56">Fig. 22E</figureCitation>
), major tines 3—4× longer than minor ones; median chaetigers with harpoon-chaetae and furcates of
<specimenCount id="9D68EA8FFFC1D728FC007EDA2750FA00" box="[999,1110,1489,1515]" pageId="55" pageNumber="56" type="generic" typeStatus="types">two types</specimenCount>
(
<figureCitation id="13553D83FFC1D728FB807EDB27D4FA01" box="[1127,1234,1488,1514]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="55" pageNumber="56">Fig. 22G</figureCitation>
), thinnest golden, with major tines 5–14× longer than minor ones, and wider with major tines 4× longer than minor ones. Harpoonchaetae with short, almost blunt denticles and smooth tines (denticulate tines 3–6× longer than smooth ones). Neurochaetae all furcates, major tines 4–6× longer than minor ones, anterior chaetigers with tines longer (
<figureCitation id="13553D83FFC1D728FB107D37265CF9BD" box="[1271,1370,1596,1622]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="55" pageNumber="56">Fig. 22F</figureCitation>
), than those present in median chaetigers (
<figureCitation id="13553D83FFC1D728FDCF7D6B2197F991" box="[552,657,1632,1658]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="55" pageNumber="56">Fig. 22H</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD12106FFC1D728FF207D8F274AF975" blockId="55.[151,1437,1416,2019]" box="[199,1100,1668,1695]" pageId="55" pageNumber="56">
Posterior region tapered (
<figureCitation id="13553D83FFC1D728FE037D8F214BF975" box="[484,589,1668,1694]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="55" pageNumber="56">Fig. 22D</figureCitation>
); pygidium with anus terminal, anal cirri lost.
</paragraph>
<paragraph id="8BD12106FFC1D728FF207DA32229F929" blockId="55.[151,1437,1416,2019]" box="[199,303,1704,1730]" pageId="55" pageNumber="56">
<emphasis id="B91AFD14FFC1D728FF207DA32229F929" bold="true" box="[199,303,1704,1730]" pageId="55" pageNumber="56">Epitokes</emphasis>
</paragraph>
<paragraph id="8BD12106FFC1D728FF207DC72747F8E1" blockId="55.[151,1437,1416,2019]" pageId="55" pageNumber="56">
Epitokes (ZMA VPol 156.1), complete, posterior end bent ventrally or laterally (
<figureCitation id="13553D83FFC1D728FBB47DC727A1F90D" box="[1107,1191,1740,1766]" captionStart="FIGURE 23" captionStartId="54.[152,255,1889,1913]" captionTargetBox="[152,1435,564,1864]" captionTargetId="figure-182@54.[151,1436,561,1866]" captionTargetPageId="54" captionText="FIGURE 23. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1).A. Largest epitoke (35x6 mm), anterior region, dorsal view. B. Same, posterior region, dorsal view. C. Smallest epitoke (16x4 mm), anterior region, dorsal view. D. Same, posterior region, dorsal view. E. Medium-sized epitoke (22x4 mm), anterior region, dorsal view. F. Same, posterior region, dorsal view. Scale bars: A, B, 1.4 mm; C, D, 0.9 mm; E, F, 1.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7621879" httpUri="https://zenodo.org/record/7621879/files/figure.png" pageId="55" pageNumber="56">Fig. 23</figureCitation>
), lateral antennae lost in largest epitoke; body fusiform,
<quantity id="4C968CE3FFC1D728FDF37DFB218BF8E0" box="[532,653,1776,1803]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="3.5" metricValueMin="1.6" pageId="55" pageNumber="56" unit="mm" value="25.5" valueMax="35.0" valueMin="16.0">16–35 mm</quantity>
long,
<quantity id="4C968CE3FFC1D728FD347DFB2036F8E0" box="[723,816,1776,1803]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="55" pageNumber="56" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
wide, 23–26 chaetigers.
</paragraph>
<paragraph id="8BD12106FFC1D728FF207C1E23D3F871" blockId="55.[151,1437,1416,2019]" pageId="55" pageNumber="56">
Largest epitoke (ZMA VPol 156.1) pale; anterior prostomial area grayish (
<figureCitation id="13553D83FFC1D728FBFF7C1F278DF8C5" box="[1048,1163,1812,1838]" captionStart="FIGURE 22" captionStartId="53.[152,255,1645,1669]" captionTargetBox="[155,1436,183,1621]" captionTargetId="figure-18@53.[151,1436,181,1621]" captionTargetPageId="53" captionText="FIGURE 22. Chloeia fusca M’Intosh, 1885.A. Holotype (BMNH 1885.12.1.9).A.Anterior fragment, dorsal view. B. Anterior end, dorsal view. C. Posterior fragment, anterior chaetigers. D. Posterior fragment, dorsal view. E. Largest topotype specimen (RMNH 1245), chaetiger 3, notochaetae (inset: close-up of notochaetal tip). F. Same, neurochaetal tips. G. Chaetiger 13, notochetae (inset: harpoon notochaetal tip). H. Same, neurochaetae (insets: shorter and longer tines). Scale bars: A, 0.7 mm; B, 0.3 mm; C, 0.4 mm; D, 1.6 mm; E, 60 μm; F, 50 μm; G, 90 μm; H, 110 μm." figureDoi="http://doi.org/10.5281/zenodo.7621875" httpUri="https://zenodo.org/record/7621875/files/figure.png" pageId="55" pageNumber="56">Figs. 22A</figureCitation>
,
<figureCitation id="13553D83FFC1D728FB7E7C1F27CBF8C5" box="[1177,1229,1812,1838]" captionStart="FIGURE 23" captionStartId="54.[152,255,1889,1913]" captionTargetBox="[152,1435,564,1864]" captionTargetId="figure-182@54.[151,1436,561,1866]" captionTargetPageId="54" captionText="FIGURE 23. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1).A. Largest epitoke (35x6 mm), anterior region, dorsal view. B. Same, posterior region, dorsal view. C. Smallest epitoke (16x4 mm), anterior region, dorsal view. D. Same, posterior region, dorsal view. E. Medium-sized epitoke (22x4 mm), anterior region, dorsal view. F. Same, posterior region, dorsal view. Scale bars: A, B, 1.4 mm; C, D, 0.9 mm; E, F, 1.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7621879" httpUri="https://zenodo.org/record/7621879/files/figure.png" pageId="55" pageNumber="56">23A</figureCitation>
); middorsal bands purple (
<figureCitation id="13553D83FFC1D728FF097C332273F8B9" box="[238,373,1848,1874]" captionStart="FIGURE 23" captionStartId="54.[152,255,1889,1913]" captionTargetBox="[152,1435,564,1864]" captionTargetId="figure-182@54.[151,1436,561,1866]" captionTargetPageId="54" captionText="FIGURE 23. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1).A. Largest epitoke (35x6 mm), anterior region, dorsal view. B. Same, posterior region, dorsal view. C. Smallest epitoke (16x4 mm), anterior region, dorsal view. D. Same, posterior region, dorsal view. E. Medium-sized epitoke (22x4 mm), anterior region, dorsal view. F. Same, posterior region, dorsal view. Scale bars: A, B, 1.4 mm; C, D, 0.9 mm; E, F, 1.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7621879" httpUri="https://zenodo.org/record/7621879/files/figure.png" pageId="55" pageNumber="56">Fig. 23A, B</figureCitation>
); accessory lateral bands barely pigmented, continued along internotopodial space. Dorsal cirri blackish. Bipinnate branchiae with stems pale, lateral branches purple. Venter pale, midventral band shiny, slightly paler.
</paragraph>
<paragraph id="8BD12106FFC1D727FF207CAF2715FF3E" blockId="55.[151,1437,1416,2019]" lastBlockId="56.[151,1436,151,393]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">Prostomium anteriorly entire. Eyes blackish, coalescent, anterior eyes 6× larger than posterior ones. Median antenna pale, inserted at anterior caruncular margin, half as long as caruncle, size relationship to lateral antennae unknown. Lateral antennae with blackish ceratophores, ceratostyles lost, bases separate from each other, size relationship to palps unknown. Mouth ventral on chaetiger 2. Pharynx not exposed.</paragraph>
<paragraph id="8BD12106FFCED727FF207BD42781FEF6" blockId="56.[151,1436,151,393]" pageId="56" pageNumber="57">Caruncle pale, straight, trilobed, tapered, reaching chaetiger 4. Median ridge plicate, with about 36 vertical folds, partially concealing lateral lobes. Lateral lobes narrow, with about 34 vertical folds.</paragraph>
<paragraph id="8BD12106FFCED727FF207A2C2079FE62" blockId="56.[151,1436,151,393]" pageId="56" pageNumber="57">Bipinnate branchiae from chaetiger 5, continued throughout body, divergent along body; progressively larger to chaetiger 9–10, about half as long as successive segments, decreasing in size posteriorly (becoming as long as successive segments); median segments with 5–6 lateral branches.</paragraph>
<caption id="DF11718EFFCED727FF7F7DB42090F844" ID-DOI="http://doi.org/10.5281/zenodo.7621887" ID-Zenodo-Dep="7621887" httpUri="https://zenodo.org/record/7621887/files/figure.png" pageId="56" pageNumber="57" startId="56.[152,255,1727,1751]" targetBox="[154,1433,417,1703]" targetPageId="56">
<paragraph id="8BD12106FFCED727FF7F7DB42090F844" blockId="56.[151,1437,1727,1967]" pageId="56" pageNumber="57">
<emphasis id="B91AFD14FFCED727FF7F7DB42226F93C" bold="true" box="[152,288,1727,1751]" pageId="56" pageNumber="57">FIGURE 25</emphasis>
.
<taxonomicName id="4C6E5A85FFCED727FEC87DB42161F93C" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[303,615,1727,1751]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="56" pageNumber="57" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCED727FEC87DB422BAF93C" box="[303,444,1727,1751]" italics="true" pageId="56" pageNumber="57">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFCED727FE227DB42161F93C" author="M'Intosh, W. C." box="[453,615,1727,1751]" pageId="56" pageNumber="57" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
, non-type specimens (UF 4368) showing variation in reddish pigmentation and branchial contraction. A) Anterior region, dorsal view of specimen with visible dorsal oblique bands continued to the anterior notopodial surface, and bipinnate branchiae contracted. B) Same, right lateral view. C) Another specimen with oblique bands covered by fully extended bipinnate branchiae. D) Same, right lateral view, with less intense pigmentation along notopidal anterior surface. E) Another specimen with a few reddish chaetae, anterior region dorsal view, bipinnate branchiae covering oblique bands. F) Same, right lateral view showing pigmentation along a few anterior chaetigers on notopodial anterior surface. Scale bars: A, 1.1 mm; B, 1.4 mm; C, 1.8 mm; D, 1.3 mm; E, 0.9 mm; F, 1.5 mm.
</paragraph>
</caption>
<paragraph id="8BD12106FFCFD726FF207B9C22F3FEAA" blockId="57.[151,1437,151,1113]" pageId="57" pageNumber="58">Parapodia biramous, notopodia with purple cirriform branchiae along chaetigers 1–4, half as long as dorsal cirri in chaetiger 1, progressively shorter becoming 1/3–1/4 as long in chaetiger 4. Dorsal cirri 2× longer than bipinnate branchiae along median and posterior chaetigers. Second ventral cirri with cirrophores 2× longer and wider, and cirrostyle 2× longer than those present in chaetiger 3, directed dorsally. Other ventral cirri directed ventrolaterally, as long as subsequent segment.</paragraph>
<paragraph id="8BD12106FFCFD726FF207A4020F6FDF2" blockId="57.[151,1437,151,1113]" pageId="57" pageNumber="58">
Chaetae most complete with distal hoods. Anterior notochaetae furcates (
<figureCitation id="13553D83FFCFD726FC157A402750FE8E" box="[1010,1110,331,357]" captionStart="FIGURE 23" captionStartId="54.[152,255,1889,1913]" captionTargetBox="[152,1435,564,1864]" captionTargetId="figure-182@54.[151,1436,561,1866]" captionTargetPageId="54" captionText="FIGURE 23. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1).A. Largest epitoke (35x6 mm), anterior region, dorsal view. B. Same, posterior region, dorsal view. C. Smallest epitoke (16x4 mm), anterior region, dorsal view. D. Same, posterior region, dorsal view. E. Medium-sized epitoke (22x4 mm), anterior region, dorsal view. F. Same, posterior region, dorsal view. Scale bars: A, B, 1.4 mm; C, D, 0.9 mm; E, F, 1.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7621879" httpUri="https://zenodo.org/record/7621879/files/figure.png" pageId="57" pageNumber="58">Fig. 23B</figureCitation>
), major tines about 4× longer than minor ones; median chaetigers with harpoon-chaetae and furcate capillaries with major tines about 12× longer than minor ones; harpoon notochaetae with short, almost blunt denticles and shorter tine, denticulate tines 3–6× longer than smooth ones (
<figureCitation id="13553D83FFCFD726FE507ABC2119FE3A" box="[439,543,439,465]" captionStart="FIGURE 23" captionStartId="54.[152,255,1889,1913]" captionTargetBox="[152,1435,564,1864]" captionTargetId="figure-182@54.[151,1436,561,1866]" captionTargetPageId="54" captionText="FIGURE 23. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1).A. Largest epitoke (35x6 mm), anterior region, dorsal view. B. Same, posterior region, dorsal view. C. Smallest epitoke (16x4 mm), anterior region, dorsal view. D. Same, posterior region, dorsal view. E. Medium-sized epitoke (22x4 mm), anterior region, dorsal view. F. Same, posterior region, dorsal view. Scale bars: A, B, 1.4 mm; C, D, 0.9 mm; E, F, 1.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7621879" httpUri="https://zenodo.org/record/7621879/files/figure.png" pageId="57" pageNumber="58">Fig. 23D</figureCitation>
). Neurochaetae all furcates; anterior chaetigers with major tines about 4× longer than minor ones (
<figureCitation id="13553D83FFCFD726FEB87AD022CEFE1E" box="[351,456,475,501]" captionStart="FIGURE 24" captionStartId="55.[152,255,1202,1226]" captionTargetBox="[154,1436,183,1175]" captionTargetId="figure-18@55.[151,1436,181,1178]" captionTargetPageId="55" captionText="FIGURE 24. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1). A. Largest epitoke, anterior end, right lateral view. B. Chaetiger 3, notochaetae. C. Same, neurochaetal tips. D. Chaetiger 14, harpoon notochaetae and capillary furcates. E. Same, furcates and capillary furcates (inset: capillary furcates furca and tips). F. Smallest epitoke, anterior end, right lateral view. G. Medium-sized epitoke, anterior end, right lateral view. Scale bars: A, 1.2 mm; B, 80 μm; C, 30 μm; D, 35 μm; E, 90 μm; F, 0.6 mm; G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7621883" httpUri="https://zenodo.org/record/7621883/files/figure.png" pageId="57" pageNumber="58">Fig. 24C</figureCitation>
); median chaetigers with furcates with major tines 5–12× longer than minor ones, and capillary furcates with major tines 7–15× longer than minor ones (
<figureCitation id="13553D83FFCFD726FC997AF420E2FDF2" box="[894,996,511,537]" captionStart="FIGURE 24" captionStartId="55.[152,255,1202,1226]" captionTargetBox="[154,1436,183,1175]" captionTargetId="figure-18@55.[151,1436,181,1178]" captionTargetPageId="55" captionText="FIGURE 24. Chloeia fusca M’Intosh, 1885, epitokes (ZMA VPol 156.1). A. Largest epitoke, anterior end, right lateral view. B. Chaetiger 3, notochaetae. C. Same, neurochaetal tips. D. Chaetiger 14, harpoon notochaetae and capillary furcates. E. Same, furcates and capillary furcates (inset: capillary furcates furca and tips). F. Smallest epitoke, anterior end, right lateral view. G. Medium-sized epitoke, anterior end, right lateral view. Scale bars: A, 1.2 mm; B, 80 μm; C, 30 μm; D, 35 μm; E, 90 μm; F, 0.6 mm; G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7621883" httpUri="https://zenodo.org/record/7621883/files/figure.png" pageId="57" pageNumber="58">Fig. 24E</figureCitation>
).
</paragraph>
<paragraph id="8BD12106FFCFD726FF2079282644FDD6" blockId="57.[151,1437,151,1113]" box="[199,1346,546,573]" pageId="57" pageNumber="58">Posterior region tapered; pygidium with anus terminal, anal cirri pale, digitate, 4–5× longer than wide.</paragraph>
<paragraph id="8BD12106FFCFD726FF20794D2653FD1A" blockId="57.[151,1437,151,1113]" pageId="57" pageNumber="58">
<emphasis id="B91AFD14FFCFD726FF20794D2113FD8A" bold="true" box="[199,533,582,609]" pageId="57" pageNumber="58">Live pigmentation of atokes</emphasis>
(after freshly collected specimens UF 4368,
<figureCitation id="13553D83FFCFD726FBF5794C2762FD8A" box="[1042,1124,582,609]" captionStart="FIGURE 25" captionStartId="56.[152,255,1727,1751]" captionTargetBox="[154,1433,417,1703]" captionTargetId="figure-123@56.[151,1436,417,1704]" captionTargetPageId="56" captionText="FIGURE 25. Chloeia fusca M’Intosh, 1885, non-type specimens (UF 4368) showing variation in reddish pigmentation and branchial contraction. A) Anterior region, dorsal view of specimen with visible dorsal oblique bands continued to the anterior notopodial surface, and bipinnate branchiae contracted. B) Same, right lateral view. C) Another specimen with oblique bands covered by fully extended bipinnate branchiae. D) Same, right lateral view, with less intense pigmentation along notopidal anterior surface. E) Another specimen with a few reddish chaetae, anterior region dorsal view, bipinnate branchiae covering oblique bands. F) Same, right lateral view showing pigmentation along a few anterior chaetigers on notopodial anterior surface. Scale bars: A, 1.1 mm; B, 1.4 mm; C, 1.8 mm; D, 1.3 mm; E, 0.9 mm; F, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621887" httpUri="https://zenodo.org/record/7621887/files/figure.png" pageId="57" pageNumber="58">Fig. 25</figureCitation>
, and
<bibRefCitation id="EFFF5CF7FFCFD726FB46794C2628FD8A" author="Belle, J." box="[1185,1326,583,609]" pageId="57" pageNumber="58" refId="ref103482" refString="Belle, J. (2022 b) Bristle worm (Chloeia fusca), Anilao, Philippines. Available from: https: // www. natureinstock. com / search / preview / bristle-worm-chloeia-fusca-anilao-philippines / 0 _ 10120610. html (accessed 31 July 2022)" type="url" year="2022">Belle 2022b</bibRefCitation>
). Dorsum pale to pinkish; middorsal bands reddish, continued throughout body, each band slightly expanded towards lateral margins. Brownish to reddish lateral bands approaching middorsal bands, running towards anterior notopodial surfaces. Caruncle pale. Dorsal cirri dark purple to blackish. Branchiae pale brownish to reddish. Notochaetae and neurochaetae with a distal yellowish band, rarely with red bases. Venter with midventral band pale (
<figureCitation id="13553D83FFCFD726FB1279DC264AFD1A" box="[1269,1356,727,753]" captionStart="FIGURE 4" captionStartId="10.[152,255,1827,1851]" captionTargetBox="[154,1433,635,1802]" captionTargetId="figure-246@10.[151,1436,633,1804]" captionTargetPageId="10" captionText="FIGURE 4. Chloeia fusca M’Intosh, 1885, non-type specimen (UF 4368). A. Ventral view. B. Ventral view after body wall removed. C. Same, after Methyl Green staining. D. Same, lower half of digestive tube removed. (AI, anterior intestine; MB, midventral band (nerve chord); MR, muscular ridge; Oe, oesophagus; PI, posterior intestine; R, rectum; St, stomach). All scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7621812" httpUri="https://zenodo.org/record/7621812/files/figure.png" pageId="57" pageNumber="58">Fig. 4A</figureCitation>
)
</paragraph>
<paragraph id="8BD12106FFCFD726FF2079F1215CFBB2" blockId="57.[151,1437,151,1113]" pageId="57" pageNumber="58">
<emphasis id="B91AFD14FFCFD726FF2079F12121FCFF" bold="true" box="[199,551,762,788]" pageId="57" pageNumber="58">Live pigmentation of epitokes</emphasis>
. Recently collected specimens (CAS 218217) have dark brown to blackish longitudinal middorsal bands (
<figureCitation id="13553D83FFCFD726FE2A78142131FCD2" box="[461,567,799,825]" captionStart="FIGURE 26" captionStartId="57.[152,255,1755,1779]" captionTargetBox="[178,1411,1140,1731]" captionTargetId="figure-439@57.[176,1411,1137,1732]" captionTargetPageId="57" captionText="FIGURE 26. Chloeia fusca M’Intosh, 1885, freshly collected epitokes (CAS 218217). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 10-11, dorsal view. D. Posterior region, slightly oblique dorsal view. Scale bars: A, 1.4 mm; B, 0.7 mm; C, 0.8 mm; D, 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621893" httpUri="https://zenodo.org/record/7621893/files/figure.png" pageId="57" pageNumber="58">Fig. 26A</figureCitation>
); in each segment, there are two thick orange-brownish thick bands, roughly parallel along anterior chaetigers, and in median chaetigers become divergent, connected laterally in darker wide bands, running along anterior parapodial surfaces, and another orange thinner band runs along the posterior parapodial surface (
<figureCitation id="13553D83FFCFD726FEC078802288FC4E" box="[295,398,907,933]" captionStart="FIGURE 26" captionStartId="57.[152,255,1755,1779]" captionTargetBox="[178,1411,1140,1731]" captionTargetId="figure-439@57.[176,1411,1137,1732]" captionTargetPageId="57" captionText="FIGURE 26. Chloeia fusca M’Intosh, 1885, freshly collected epitokes (CAS 218217). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 10-11, dorsal view. D. Posterior region, slightly oblique dorsal view. Scale bars: A, 1.4 mm; B, 0.7 mm; C, 0.8 mm; D, 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621893" httpUri="https://zenodo.org/record/7621893/files/figure.png" pageId="57" pageNumber="58">Fig. 26C</figureCitation>
). Prostomium grayish, anterior prostomial area pale. Eyes blackish, anterior eyes 4–6× larger than posterior ones, often in a dark area. Lateral antennae and palps purple, at least basally (
<figureCitation id="13553D83FFCFD726FB6178A427EDFC22" box="[1158,1259,943,969]" captionStart="FIGURE 26" captionStartId="57.[152,255,1755,1779]" captionTargetBox="[178,1411,1140,1731]" captionTargetId="figure-439@57.[176,1411,1137,1732]" captionTargetPageId="57" captionText="FIGURE 26. Chloeia fusca M’Intosh, 1885, freshly collected epitokes (CAS 218217). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 10-11, dorsal view. D. Posterior region, slightly oblique dorsal view. Scale bars: A, 1.4 mm; B, 0.7 mm; C, 0.8 mm; D, 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621893" httpUri="https://zenodo.org/record/7621893/files/figure.png" pageId="57" pageNumber="58">Fig. 26B</figureCitation>
), median antennae colorless to pale orange, with a purple basal ring below the base, slightly longer than caruncle when complete. Caruncle brownish with median ridge darker. Cirriform branchiae and dorsal cirri purple. Branchiae with orange stems, pinnules purple to paler with reddish tips. Chaetae with a wide yellow band. Venter with longitudinal median pale band. Anal cirri whitish (
<figureCitation id="13553D83FFCFD726FE027F342148FBB2" box="[485,590,1087,1113]" captionStart="FIGURE 26" captionStartId="57.[152,255,1755,1779]" captionTargetBox="[178,1411,1140,1731]" captionTargetId="figure-439@57.[176,1411,1137,1732]" captionTargetPageId="57" captionText="FIGURE 26. Chloeia fusca M’Intosh, 1885, freshly collected epitokes (CAS 218217). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 10-11, dorsal view. D. Posterior region, slightly oblique dorsal view. Scale bars: A, 1.4 mm; B, 0.7 mm; C, 0.8 mm; D, 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621893" httpUri="https://zenodo.org/record/7621893/files/figure.png" pageId="57" pageNumber="58">Fig. 26D</figureCitation>
).
</paragraph>
<caption id="DF11718EFFCFD726FF7F7DD02136F8D0" ID-DOI="http://doi.org/10.5281/zenodo.7621893" ID-Zenodo-Dep="7621893" httpUri="https://zenodo.org/record/7621893/files/figure.png" pageId="57" pageNumber="58" startId="57.[152,255,1755,1779]" targetBox="[178,1411,1140,1731]" targetPageId="57">
<paragraph id="8BD12106FFCFD726FF7F7DD02136F8D0" blockId="57.[151,1437,1755,1851]" pageId="57" pageNumber="58">
<emphasis id="B91AFD14FFCFD726FF7F7DD02227F918" bold="true" box="[152,289,1755,1779]" pageId="57" pageNumber="58">FIGURE 26</emphasis>
.
<taxonomicName id="4C6E5A85FFCFD726FED67DD0216AF918" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[305,620,1755,1779]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="57" pageNumber="58" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCFD726FED67DD022B9F918" box="[305,447,1755,1779]" italics="true" pageId="57" pageNumber="58">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFCFD726FE2E7DD0216AF918" author="M'Intosh, W. C." box="[457,620,1755,1779]" pageId="57" pageNumber="58" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
, freshly collected epitokes (CAS 218217). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 10-11, dorsal view. D. Posterior region, slightly oblique dorsal view. Scale bars: A, 1.4 mm; B, 0.7 mm; C, 0.8 mm; D, 1.2 mm.
</paragraph>
</caption>
<paragraph id="8BD12106FFCFD725FF207C62266BFDD6" blockId="57.[151,1437,1897,2032]" lastBlockId="58.[151,1437,151,2013]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<emphasis id="B91AFD14FFCFD726FF207C6222AFF868" bold="true" box="[199,425,1897,1923]" pageId="57" pageNumber="58">Variation of atokes</emphasis>
. Seventeen topotype specimens (RMNH 1245, ZMA VPol 156.5, ZMA VPol 156.7) complete, retaining pigmentation pattern. Body
<quantity id="4C968CE3FFCFD726FD6E7C86201FF843" box="[649,793,1933,1960]" metricMagnitude="-2" metricUnit="m" metricValue="1.675" metricValueMax="2.3" metricValueMin="1.05" pageId="57" pageNumber="58" unit="mm" value="16.75" valueMax="23.0" valueMin="10.5">10.5–23 mm</quantity>
long,
<quantity id="4C968CE3FFCFD726FC857C8620C7F843" box="[866,961,1933,1960]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="57" pageNumber="58" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
wide, 18–24 chaetigers. All with anterior prostomial area blackish; several specimens with darker lips, and a wide, diffuse brownish transverse ventral band, often displaced towards the anterior segmental half. Eyes blackish, separate; anterior eyes 3–4× larger than posterior ones, Median antenna pale, often long as caruncle, and even longer in some specimens, 2× longer than lateral antennae; lateral antennae and palps darker, often dark purple. Caruncle pale. Branchiae from chaetiger 5. Paired longitudinal bands dark purple, ill-defined along anterior chaetigers of smallest specimens (
<quantity id="4C968CE3FFCCD725FB847BD427C8FF12" box="[1123,1230,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="58" pageNumber="59" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
long,
<quantity id="4C968CE3FFCCD725FAF27BD5269BFF12" box="[1301,1437,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="58" pageNumber="59" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
wide), bands discontinuous with 1–2 wider areas per segment, becoming continuous and medially wider along posterior segments, band margins ill-defined. Median-sized specimens (
<quantity id="4C968CE3FFCCD725FC687A2C270EFEAA" box="[911,1032,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="58" pageNumber="59" unit="mm" value="15.0" valueMax="17.0" valueMin="13.0">13–17 mm</quantity>
long,
<quantity id="4C968CE3FFCCD725FBAA7A2D27AFFEAA" box="[1101,1193,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="58" pageNumber="59" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
wide) with additional bands visible, brownish, roughly parallel to middorsal bands, continued laterally along anterior notopodial surfaces; longitudinal bands thin to wide, not depending on segment contraction. Longest specimens (
<quantity id="4C968CE3FFCCD725FB557A64262FFE62" box="[1202,1321,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="58" pageNumber="59" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">18-22 mm</quantity>
long,
<quantity id="4C968CE3FFCCD725FA957A6523C5FE46" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="58" pageNumber="59" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
wide) with thinner brownish bands anteriorly, blackish posteriorly; lateral bands ill-defined, roughly parallel to middorsal bands. Lateral bands paler, parallel to middorsal ones, continued along anterior notopodial surfaces. Branchiae from chaetiger 5, with pale stems, lateral branches purple. Anal cirri fragile, easily detached, yellowish, ovoid to digitate, 2–4× longer than wide. Non topotype specimens from
<collectingCountry id="F3796196FFCCD725FC2C7AF4273EFDF2" box="[971,1080,511,537]" name="Indonesia" pageId="58" pageNumber="59">Indonesia</collectingCountry>
(ZMA VPol 156.2, 156.6), and from the
<collectingCountry id="F3796196FFCCD725FF197928227AFDD6" box="[254,380,547,573]" name="Philippines" pageId="58" pageNumber="59">Philippines</collectingCountry>
(ZMA VPol 156.3, 156.4) match the topotype specimens and are regarded as conspecific.
</paragraph>
<paragraph id="8BD12106FFCCD725FF20794D2143FCB6" blockId="58.[151,1437,151,2013]" pageId="58" pageNumber="59">
<emphasis id="B91AFD14FFCCD725FF20794D22B8FD8B" bold="true" box="[199,446,582,608]" pageId="58" pageNumber="59">Variation of epitokes</emphasis>
.
<materialsCitation id="3B062B5BFFCCD725FE2C794C2147FCB6" ID-GBIF-Occurrence="4029882484" collectionCode="ZMA" pageId="58" pageNumber="59" specimenCount="1" typeStatus="paratype">
Smaller epitokes (
<collectionCode id="ED7FB9C3FFCCD725FD74794C21D4FD8A" box="[659,722,583,609]" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="58" pageNumber="59">ZMA</collectionCode>
VPol 156.1) complete, with darker pigmentation than described for the largest epitoke (
<figureCitation id="13553D83FFCCD725FE727960211FFD6E" box="[405,537,619,645]" captionStart="FIGURE 21" captionStartId="49.[152,251,1277,1301]" captionTargetBox="[153,1433,185,1250]" captionTargetId="figure-18@49.[151,1436,181,1254]" captionTargetPageId="49" captionText="FIGURE 21. Chloeia fucata de Quatrefages, 1866, holotype (MNHN IA-TYPE 80). A. Right lateral view, after Methyl green staining (MGS). B. Anterior end, oblique right lateral view, after MGS. C. Anterior region, dorsal view (asterisks indicate first non-cirriform branchiae). D. Median chaetigers, dorsal view. E. Chaetiger 3, notochaetae; F. Chaetiger 13, notochaetal tips; G. Same, neurochaetal tips. G. Posterior end, dorsal view. Scale bars: A, 5 mm; B, C 0.7 mm; D, 1.1 mm; E, F, 100 μm; G, 40 μm; H, 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7621871" httpUri="https://zenodo.org/record/7621871/files/figure.png" pageId="58" pageNumber="59">Fig. 21C–F</figureCitation>
). Both with anterior prostomial area barely pigmented. Eyes blackish, coalescent, anterior eyes 6× larger than posterior ones. Median antenna pale to blackish basally, 2/3–4/5 as long as caruncle, 3 × longer than lateral antennae; lateral antennae bases separate from each other. Caruncle pale, reaching chaetiger 4. Mouth in chaetiger 2. Paired longitudinal bands dark purple to reddish, well defined along body; lateral bands pales, parallel, continued into internotopodial space. Branchiae from chaetiger 5, stem pale, lateral branches purple, parallel along body; median segments with branchiae with 6–7 lateral branches. Anal cirri yellowish (present in smallest
<typeStatus id="54D59FA4FFCCD725FF70784823FAFCB6" box="[151,252,835,861]" pageId="58" pageNumber="59" type="paratype">paratype</typeStatus>
), ovoid, 2× longer than wide
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFCCD724FF20786C20F1FCD2" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="discussion">
<paragraph id="8BD12106FFCCD725FF20786C21BCFBB2" blockId="58.[151,1437,151,2013]" pageId="58" pageNumber="59">
<emphasis id="B91AFD14FFCCD725FF20786C2232FC6A" bold="true" box="[199,308,871,897]" pageId="58" pageNumber="59">Remarks</emphasis>
.
<taxonomicName id="4C6E5A85FFCCD725FEA5786C2195FC6A" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[322,659,870,897]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCCD725FEA5786C22DBFC6A" box="[322,477,871,897]" italics="true" pageId="58" pageNumber="59">Chloeia fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFCCD725FE03786C2195FC6A" author="M'Intosh, W. C." box="[484,659,870,897]" pageId="58" pageNumber="59" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
was described with a single specimen collected in
<collectingCountry id="F3796196FFCCD725FB28786C263AFC6A" box="[1231,1340,871,897]" name="Indonesia" pageId="58" pageNumber="59">Indonesia</collectingCountry>
and it is characterized by having a brownish body and two dorsal longitudinal dark bands. By this pigmentation and having bipinnate branchiae from chaetiger 5, it belongs in the group fusca, together with
<taxonomicName id="4C6E5A85FFCCD725FBDE78A42654FC22" authority="Grube, 1868" authorityName="Grube" authorityYear="1868" box="[1081,1362,943,969]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="bistriata">
<emphasis id="B91AFD14FFCCD725FBDE78A427BFFC22" box="[1081,1209,943,969]" italics="true" pageId="58" pageNumber="59">C. bistriata</emphasis>
Grube, 1868
</taxonomicName>
, originally described from the Red Sea. As indicated above, the main differences between these species are the size of eyes and caruncular complexity. In
<taxonomicName id="4C6E5A85FFCCD725FDCC78FC218EFBFA" authorityName="M'Intosh" authorityYear="1885" box="[555,648,1015,1041]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCCD725FDCC78FC218EFBFA" box="[555,648,1015,1041]" italics="true" pageId="58" pageNumber="59">C. fusca</emphasis>
</taxonomicName>
the anterior eyes are 3–4× larger than posterior ones, and the median ridge of its caruncle has 18 vertical folds, whereas in
<taxonomicName id="4C6E5A85FFCCD725FD027F102062FBDE" authorityName="Grube" authorityYear="1868" box="[741,868,1051,1077]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="bistriata">
<emphasis id="B91AFD14FFCCD725FD027F102062FBDE" box="[741,868,1051,1077]" italics="true" pageId="58" pageNumber="59">C. bistriata</emphasis>
</taxonomicName>
anterior eyes are 6× larger than posterior ones, and its caruncular median ridge has 12 vertical folds.
</paragraph>
<paragraph id="8BD12106FFCCD725FF207F682256FAE6" blockId="58.[151,1437,151,2013]" pageId="58" pageNumber="59">
<bibRefCitation id="EFFF5CF7FFCCD725FF207F6822ABFB96" author="M'Intosh, W. C." box="[199,429,1122,1149]" pageId="58" pageNumber="59" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh (1885: 14)</bibRefCitation>
indicated the body of his
<typeStatus id="54D59FA4FFCCD725FD347F682033FB96" box="[723,821,1123,1149]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
was dusky brown (hence the specific name; Latin
<emphasis id="B91AFD14FFCCD725FA957F682218FB4B" italics="true" pageId="58" pageNumber="59">fuscus, -a, -um</emphasis>
: dark, dim, black, brown). The dorsum also had two longitudinal brownish lines with a paler median area. In comparison with the original description, most pigmentation patterns are retained in the specimen which has been almost 150 years in ethanol; the only difference is in the chaetal pigmentation because the greenish yellow banding is gone.
</paragraph>
<paragraph id="8BD12106FFCCD725FF207E1C224BFA0E" blockId="58.[151,1437,151,2013]" pageId="58" pageNumber="59">
<bibRefCitation id="EFFF5CF7FFCCD725FF207E1C22D7FADA" author="Treadwell, A. L." box="[199,465,1303,1329]" pageId="58" pageNumber="59" pagination="1145 - 1181" refId="ref113334" refString="Treadwell, A. L. (1906) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (1903), 3, 1145 - 1181." type="journal article" year="1906">Treadwell (1906: 1164)</bibRefCitation>
recorded this species with specimens collected in five stations in
<collectingRegion id="49AAEFE4FFCCD725FB507E1C260DFADA" box="[1207,1291,1303,1329]" country="United States of America" name="Hawaii" pageId="58" pageNumber="59">Hawaii</collectingRegion>
, in depths of
<quantity id="4C968CE3FFCCD725FF707E30220DFABE" box="[151,267,1339,1365]" metricMagnitude="1" metricUnit="m" metricValue="7.3" metricValueMax="10.8" metricValueMin="3.8" pageId="58" pageNumber="59" unit="m" value="73.0" valueMax="108.0" valueMin="38.0">38–108 m</quantity>
, but he identified them as
<taxonomicName id="4C6E5A85FFCCD725FDDF7E302194FABE" authorityName=": Treadwell 1906:" authorityYear="1164" box="[568,658,1339,1365]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="flava">
<emphasis id="B91AFD14FFCCD725FDDF7E302194FABE" box="[568,658,1339,1365]" italics="true" pageId="58" pageNumber="59">C. flava</emphasis>
</taxonomicName>
. He noted there were no middorsal dots but instead “two continuous brown lines extending along the dorsal surface.”
<bibRefCitation id="EFFF5CF7FFCCD725FD587E5420AAFA92" author="Hartman, O." box="[703,940,1375,1401]" pageId="58" pageNumber="59" pagination="163 - 252" refId="ref107718" refString="Hartman, O. (1966) Polychaetous annelids of the Hawaiian Islands. Occasional Papers of Bernice P. Bishop Museum, 23 (11), 163 - 252." type="journal article" year="1966">Hartman (1966: 179)</bibRefCitation>
indicated that the Hawaiian form has a dorsal “row or dark spots and two continuous brown lines, one on each side of the median line” and that “branchiae start “from fifth segment”. This pigmentation pattern does not match with typical
<emphasis id="B91AFD14FFCCD725FBD37EAC2793FA2A" box="[1076,1173,1447,1473]" italics="true" pageId="58" pageNumber="59">
<taxonomicName id="4C6E5A85FFCCD725FBD37EAC2797FA2A" authorityName=": Treadwell 1906:" authorityYear="1164" box="[1076,1169,1447,1473]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="flava">C. flava</taxonomicName>
,
</emphasis>
but resembles
<taxonomicName id="4C6E5A85FFCCD725FAA77EAC224EFA0E" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCCD725FAA77EAC269AFA2A" box="[1344,1436,1447,1473]" italics="true" pageId="58" pageNumber="59">C. fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFCCD725FF707EC0224EFA0E" author="M'Intosh, W. C." box="[151,328,1482,1509]" pageId="58" pageNumber="59" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD12106FFCCD725FF207EE42123F90A" blockId="58.[151,1437,151,2013]" pageId="58" pageNumber="59">
<bibRefCitation id="EFFF5CF7FFCCD725FF207EE422B6F9E2" author="Horst, R." box="[199,432,1519,1545]" pageId="58" pageNumber="59" refId="ref108337" refString="Horst, R. (1912) Polychaeta errantia of the Siboga Expedition. Part 1. Amphinomidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 a, 1 - 43, 10 pls." type="journal volume" year="1912">Horst (1912: 22–24)</bibRefCitation>
included several specimens, some from the
<typeStatus id="54D59FA4FFCCD725FC497EE420D8F9E2" box="[942,990,1519,1545]" pageId="58" pageNumber="59">type</typeStatus>
locality (Banda Islands), and provided additional details to the original description. He noted, however an interesting difference regarding the size of eyes among some of his material; he indicated “among the worms collected with the tow-net near the Paternoster islands (ZMA V.Pol 156.1) there is a gigantic specimen, measuring
<quantity id="4C968CE3FFCCD725FCD37D502085F99E" box="[820,899,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="58" pageNumber="59" unit="mm" value="37.0">37 mm</quantity>
in length; its eyes are extraordinarily developed and coalesced, whereas in the dorsal fascicles the long and slender bristles are prevailing” (
<bibRefCitation id="EFFF5CF7FFCCD725FB7E7D74264EF972" author="Horst, R." box="[1177,1352,1663,1689]" pageId="58" pageNumber="59" refId="ref108337" refString="Horst, R. (1912) Polychaeta errantia of the Siboga Expedition. Part 1. Amphinomidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 a, 1 - 43, 10 pls." type="journal volume" year="1912">Horst 1912: 24</bibRefCitation>
). There are actually two other, smaller specimens in the same lot, and all have coalescent eyes; they are herein regarded as epitokes and were described above.
</paragraph>
<paragraph id="8BD12106FFCCD725FF207DE02061F87E" blockId="58.[151,1437,151,2013]" pageId="58" pageNumber="59">
<bibRefCitation id="EFFF5CF7FFCCD725FF207DE02289F8EE" author="Fauvel, P." box="[199,399,1770,1797]" pageId="58" pageNumber="59" refId="ref106154" refString="Fauvel, P. (1953) The Fauna of India, including Pakistan, Ceylon, Burma and Malaya. Annelida Polychaeta. Indian Press, Allahbad, 507 pp." type="book" year="1953">Fauvel (1953: 97)</bibRefCitation>
indicated
<taxonomicName id="4C6E5A85FFCCD725FDE37DE0202EF8EE" authority="Potts, 1909" authorityName="Potts" authorityYear="1909" box="[516,808,1771,1797]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="longisetosa">
<emphasis id="B91AFD14FFCCD725FDE37DE021A2F8EE" box="[516,676,1771,1797]" italics="true" pageId="58" pageNumber="59">C. longisetosa</emphasis>
<bibRefCitation id="EFFF5CF7FFCCD725FD4D7DE0202EF8EE" author="Potts, F. A." box="[682,808,1771,1797]" pageId="58" pageNumber="59" pagination="355 - 371" refId="ref111949" refString="Potts, F. A. (1909) The Percy Sladen Trust Expediton to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, 20. Polychaeta of the Indian Ocean, Part 1. The Amphinomidae. The Transactions of the Linnean Society of London, Series 2 Zoology, 12 (4), 355 - 371, pls. 45 - 46. https: // doi. org / 10.1111 / j. 1096 - 3642.1909. tb 00147. x" type="journal article" year="1909">Potts, 1909</bibRefCitation>
</taxonomicName>
was the epitoke of
<taxonomicName id="4C6E5A85FFCCD725FBE57DE0275BF8EE" authorityName="M'Intosh" authorityYear="1885" box="[1026,1117,1771,1797]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCCD725FBE57DE0275BF8EE" box="[1026,1117,1771,1797]" italics="true" pageId="58" pageNumber="59">C. fusca</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFFF5CF7FFCCD725FB8C7DE02632F8EE" author="Horst, R." box="[1131,1332,1770,1797]" pageId="58" pageNumber="59" pagination="285 - 288" refId="ref108385" refString="Horst, R. (1917) A contribution towards our knowledge of the Polychaeta of South Africa. Zoologische Mededeelingen, 3, 285 - 288." type="journal article" year="1917">Horst (1917: 285)</bibRefCitation>
recorded it from Natal, and his specimens matched the thinner longitudinal bands found in the
<collectingCountry id="F3796196FFCCD725FB8A7C0427E3F8C2" box="[1133,1253,1807,1833]" name="Seychelles" pageId="58" pageNumber="59">Seychelles</collectingCountry>
by
<bibRefCitation id="EFFF5CF7FFCCD725FAF57C0423CBF8A6" author="Potts, F. A." pageId="58" pageNumber="59" pagination="355 - 371" refId="ref111949" refString="Potts, F. A. (1909) The Percy Sladen Trust Expediton to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, 20. Polychaeta of the Indian Ocean, Part 1. The Amphinomidae. The Transactions of the Linnean Society of London, Series 2 Zoology, 12 (4), 355 - 371, pls. 45 - 46. https: // doi. org / 10.1111 / j. 1096 - 3642.1909. tb 00147. x" type="journal article" year="1909">Potts (1909: 356)</bibRefCitation>
; however, Natal specimens had shorter median antenna, about half as long as caruncle, whereas in the topotype specimens, median antennae are as long as, or longer than caruncle (
<bibRefCitation id="EFFF5CF7FFCCD725FC727C5C2757F89A" author="Horst, R." box="[917,1105,1879,1905]" pageId="58" pageNumber="59" pagination="285 - 288" refId="ref108385" refString="Horst, R. (1917) A contribution towards our knowledge of the Polychaeta of South Africa. Zoologische Mededeelingen, 3, 285 - 288." type="journal article" year="1917">Horst 1917: 286</bibRefCitation>
). These records might belong in
<taxonomicName id="4C6E5A85FFCCD725FF537C7022DBF87E" authority="sensu Potts, 1909" authorityName="sensu Potts" authorityYear="1909" box="[180,477,1915,1941]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="58" pageNumber="59" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCCD725FF537C702253F87F" box="[180,341,1915,1941]" italics="true" pageId="58" pageNumber="59">C. fusca sensu</emphasis>
<bibRefCitation id="EFFF5CF7FFCCD725FEBB7C7022DBF87E" author="Potts, F. A." box="[348,477,1915,1941]" pageId="58" pageNumber="59" pagination="355 - 371" refId="ref111949" refString="Potts, F. A. (1909) The Percy Sladen Trust Expediton to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, 20. Polychaeta of the Indian Ocean, Part 1. The Amphinomidae. The Transactions of the Linnean Society of London, Series 2 Zoology, 12 (4), 355 - 371, pls. 45 - 46. https: // doi. org / 10.1111 / j. 1096 - 3642.1909. tb 00147. x" type="journal article" year="1909">Potts, 1909</bibRefCitation>
</taxonomicName>
, ande belong to a different species.
</paragraph>
<paragraph id="8BD12106FFCCD724FF207C942792FF3E" blockId="58.[151,1437,151,2013]" lastBlockId="59.[151,1437,151,897]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
There is little information regarding morphological modifications among amphinomids for reproduction; for example, in the closely related euphrosinids, an increase in the number of capillary chaetae, and in eye size were documented (
<bibRefCitation id="EFFF5CF7FFCDD724FEC97B9C22C2FF5A" author="Naville, A." box="[302,452,151,177]" pageId="59" pageNumber="60" pagination="171 - 208" refId="ref111273" refString="Naville, A. (1933) Quelques formes epitoques d'annelids polychetes nouveles ou peu connues pechees a la lumiere dans la baie de Banyuls. Annals des Sciences Naturelles, Zoologie, Series 10, 16, 171 - 208." type="journal article" year="1933">Naville 1933</bibRefCitation>
), and this would imply these specimens are epitokes. Nevertheless, these differences are regarded as sufficient for separating these coalescent-eyes specimens as a different species.
</paragraph>
<paragraph id="8BD12106FFCDD724FF207BD42133FD8A" blockId="59.[151,1437,151,897]" pageId="59" pageNumber="60">
The epitokes described above match
<taxonomicName id="4C6E5A85FFCDD724FD817BD42071FF12" authority="M'Intosh, 1885" authorityName="M'Intosh" authorityYear="1885" box="[614,887,222,249]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="59" pageNumber="60" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FFCDD724FD817BD421C7FF12" box="[614,705,223,249]" italics="true" pageId="59" pageNumber="60">C. fusca</emphasis>
<bibRefCitation id="EFFF5CF7FFCDD724FD2E7BD42071FF12" author="M'Intosh, W. C." box="[713,887,222,249]" pageId="59" pageNumber="60" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh, 1885</bibRefCitation>
</taxonomicName>
by having two longitudinal bands along dorsum, although there are some differences regarding the color of the longitudinal bands, the pigmentation of the anterior prostomial region, and the pigmentation of dorsal cirri along chaetigers 1–3. In the epitokes the longitudinal bands are reddish, the anterior prostomial region is pale to gray, and the first three dorsal cirri are pale, whereas in the nonepitoke specimens the bands are brownish, dark purple or blackish, the anterior prostomial region is black, and all dorsal cirri are dark purple or blackish. Chaetae are very similar in these two species, although in the epitokes there are more abundant long, thin furcates in notopodia, and neuropodia giving their chaetal fascicles a wooly outlook. This difference, however, is related to epitoky resulting in swimming specimens, as indicated by the fact that they were collected in the water column (
<bibRefCitation id="EFFF5CF7FFCDD724FDD37AF421BEFDF2" author="Horst, R." box="[564,696,511,537]" pageId="59" pageNumber="60" refId="ref108337" refString="Horst, R. (1912) Polychaeta errantia of the Siboga Expedition. Part 1. Amphinomidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 a, 1 - 43, 10 pls." type="journal volume" year="1912">Horst 1912</bibRefCitation>
). The main difference is the size of eyes; in epitokes eyes are coalescent, anterior eyes about 6× larger than posterior ones, whereas in atoke specimens they are separate, anterior eyes 3–4× larger than posterior ones.
</paragraph>
<paragraph id="8BD12106FFCDD724FF20796020F1FCD2" blockId="59.[151,1437,151,897]" pageId="59" pageNumber="60">
All images or videos made after living atoke specimens (
<bibRefCitation id="EFFF5CF7FFCDD724FCB9796020EBFD6E" author="Belle, J." box="[862,1005,619,645]" pageId="59" pageNumber="60" refId="ref103482" refString="Belle, J. (2022 b) Bristle worm (Chloeia fusca), Anilao, Philippines. Available from: https: // www. natureinstock. com / search / preview / bristle-worm-chloeia-fusca-anilao-philippines / 0 _ 10120610. html (accessed 31 July 2022)" type="url" year="2022">Belle 2022b</bibRefCitation>
,
<bibRefCitation id="EFFF5CF7FFCDD724FC1C7960279FFD6E" author="Charpin, F." box="[1019,1177,619,645]" pageId="59" pageNumber="60" refId="ref104295" refString="Charpin, F. (2022) Darklined fireworm. Available from: https: // reefguide. org / chloeiafusca. html (accessed 31 July 2022)" type="url" year="2022">Charpin 2022</bibRefCitation>
) corresponding to this species show reddish longitudinal bands over a pale background. The shift to brownish might be explained after the preservative used during the Challenger Expedition, which made the reddish bands turn brownish, and the pale dorsum became darkened, and this explains the name
<bibRefCitation id="EFFF5CF7FFCDD724FD0E79DC20A7FD1A" author="M'Intosh, W. C." box="[745,929,726,753]" pageId="59" pageNumber="60" refId="ref109895" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 12 (34), i - xxxvi + 1 - 554, pl. 1 - 55, 1 A - 39 A, &amp; Annelida stations map." type="book" year="1885">M’Intosh (1885)</bibRefCitation>
introduced for the species. On the other hand, after the clarification of the variations in pigmentation, the surprisingly wide distribution of this species, from the Indian to the Western Pacific should be reassessed with molecular indicators.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFCDD724FF20784922FFFC6A" pageId="59" pageNumber="60" type="distribution">
<paragraph id="8BD12106FFCDD724FF20784922FFFC6A" blockId="59.[151,1437,151,897]" pageId="59" pageNumber="60">
<emphasis id="B91AFD14FFCDD724FF207849225FFCB7" bold="true" box="[199,345,834,860]" pageId="59" pageNumber="60">Distribution</emphasis>
.
<collectingCountry id="F3796196FFCDD724FE8F784822DEFCB6" box="[360,472,835,861]" name="Indonesia" pageId="59" pageNumber="60">Indonesia</collectingCountry>
,
<collectingCountry id="F3796196FFCDD724FE03784821C7FCB6" box="[484,705,835,861]" name="Papua New Guinea" pageId="59" pageNumber="60">Papua New Guinea</collectingCountry>
, Line and Loyalty Islands,
<collectingCountry id="F3796196FFCDD724FC13784827B1FCB6" box="[1012,1207,835,861]" name="French Polynesia" pageId="59" pageNumber="60">French Polynesia</collectingCountry>
to Hawaii, from the intertidal to
<quantity id="4C968CE3FFCDD724FEC7786C2261FC6A" box="[288,359,871,897]" metricMagnitude="2" metricUnit="m" metricValue="2.04" pageId="59" pageNumber="60" unit="m" value="204.0">204 m</quantity>
water depth.
</paragraph>
</subSubSection>
</treatment>
</document>
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