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<document id="65F106CDC4F1EC7CDD96F14EFEF00127" ID-DOI="10.5852/ejt.2023.915.2379" ID-ISSN="2118-9773" ID-Zenodo-Dep="10410126" ID-ZooBank="78346F83-1177-4EAE-AFFD-D4DDCC1BFB83" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703073859257" checkinUser="plazi" docAuthor="Balkenohl, Michael" docDate="2023" docId="4A6487ECCF2AFFD0FDA5F905FC992DB5" docLanguage="en" docName="EJT.2023.915.1-57.pdf" docOrigin="European Journal of Taxonomy 915 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2379/10451" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sulciclivina andrewesi Balkenohl 2023, sp. nov." docType="treatment" docUuid="DF22D598-5910-4EA2-B4F1-6A5EF2506E7B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="48" masterDocId="B65DFF94CF07FFFFFFC5FFC1FF8F2A5D" masterDocTitle="Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="46" updateTime="1703074388710" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="88E989A272549C1BE8EE04D41E15A942">Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)</mods:title>
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<mods:date id="BCFB17B9B7CAC89EFA410B34383B4691">2023</mods:date>
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<treatment id="4A6487ECCF2AFFD0FDA5F905FC992DB5" ID-DOI="http://doi.org/10.5281/zenodo.10410158" ID-Zenodo-Dep="10410158" LSID="urn:lsid:zoobank.org:act:DF22D598-5910-4EA2-B4F1-6A5EF2506E7B" httpUri="http://treatment.plazi.org/id/4A6487ECCF2AFFD0FDA5F905FC992DB5" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46">
<subSubSection id="8AD76571CF2AFFD2FDA5F905FB4B2D58" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="C27236FACF2AFFD2FDA5F905FC5C2C82" blockId="45.[367,1220,1732,1834]" box="[608,979,1732,1759]" pageId="45" pageNumber="46">
<heading id="993A8196CF2AFFD2FDA5F905FC5C2C82" box="[608,979,1732,1759]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="46" reason="2">
<taxonomicName id="05CD4D79CF2AFFD2FDA5F905FCFE2C82" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[608,881,1732,1759]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="andrewesi" status="sp. nov.">
<emphasis id="F0B9EAE8CF2AFFD2FDA5F905FCFE2C82" bold="true" box="[608,881,1732,1759]" italics="true" pageId="45" pageNumber="46">Sulciclivina andrewesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB8A5793CF2AFFD2FCBCF904FC5C2C82" box="[889,979,1733,1759]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<paragraph id="C27236FACF2AFFD2FD07F8CEFCFE2D77" blockId="45.[367,1220,1732,1834]" box="[706,881,1807,1834]" pageId="45" pageNumber="46">
<figureCitation id="5AF62A7FCF2AFFD2FD07F8CEFC932D74" box="[706,796,1807,1833]" captionStart="Figs 2023" captionStartId="43.[189,242,1928,1954]" captionTargetBox="[209,1340,1340,1816]" captionTargetId="figure-373@43.[189,1398,1308,1893]" captionTargetPageId="43" captionText="Figs 2023. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 20. S. sagittaria sagittaria (Bates, 1892), ♂, paralectotype (NHMUK). 21. S. sagittaria singaporensis subsp. nov., ♀, holotype (NHMUK). 22. S. oculiangusta sp. nov., ♀, holotype (NHMUK). 23. S. andrewesi sp. nov., ♂, holotype (NHMB)." figureDoi="http://doi.org/10.5281/zenodo.10410140" httpUri="https://zenodo.org/record/10410140/files/figure.png" pageId="45" pageNumber="46">Figs 23</figureCitation>
, 5253
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF2AFFD2FF78F892FD1D2DB5" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="C27236FACF2AFFD2FF78F892FEB92D30" blockId="45.[189,310,1875,1901]" box="[189,310,1875,1901]" pageId="45" pageNumber="46">
<heading id="993A8196CF2AFFD2FF78F892FEB92D30" bold="true" box="[189,310,1875,1901]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="F0B9EAE8CF2AFFD2FF78F892FEB92D30" bold="true" box="[189,310,1875,1901]" pageId="45" pageNumber="46">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF2AFFD2FF78F845FD1D2DB5" blockId="45.[187,1400,1924,2024]" pageId="45" pageNumber="46">
The species is the distinctly largest-sized one of the
<emphasis id="F0B9EAE8CF2AFFD2FCEFF845FC2A2DC0" box="[810,933,1924,1949]" italics="true" pageId="45" pageNumber="46">sagittaria-</emphasis>
species group. Unique characters are in addition the elongated and falcate mandibles, the elongated and parallel-sided pronotum, and the elongate elytron with the lateral margins parallel.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF29FFD1FF78FEC8FD192B2B" pageId="46" pageNumber="47" type="etymology">
<paragraph id="C27236FACF29FFD1FF78FEC8FECB2B7E" blockId="46.[189,324,265,291]" box="[189,324,265,291]" pageId="46" pageNumber="47">
<heading id="993A8196CF29FFD1FF78FEC8FECB2B7E" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="F0B9EAE8CF29FFD1FF78FEC8FECB2B7E" bold="true" box="[189,324,265,291]" pageId="46" pageNumber="47">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF29FFD1FF78FEF9FD192B2B" blockId="46.[188,1400,312,374]" pageId="46" pageNumber="47">
The name is a patronym in honour of Herbert Edward Andrewes (18631950) who provided a solid basis for the investigation of Asian
<taxonomicName id="05CD4D79CF29FFD1FDDCFE9DFD1D2B2B" box="[537,658,348,374]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subFamily" subFamily="Scaritinae">Scaritinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF29FFD1FF78FE5EFE3B29AA" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="C27236FACF29FFD1FF78FE5EFEE32BE4" blockId="46.[189,364,415,441]" box="[189,364,415,441]" pageId="46" pageNumber="47">
<heading id="993A8196CF29FFD1FF78FE5EFEE32BE4" bold="true" box="[189,364,415,441]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="F0B9EAE8CF29FFD1FF78FE5EFEE32BE4" bold="true" box="[189,364,415,441]" pageId="46" pageNumber="47">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF29FFD1FF2FFE0EFED32BB4" blockId="46.[189,1399,463,598]" box="[234,348,463,489]" pageId="46" pageNumber="47">
<emphasis id="F0B9EAE8CF29FFD1FF2FFE0EFED32BB4" bold="true" box="[234,348,463,489]" pageId="46" pageNumber="47">
<typeStatus id="1D768858CF29FFD1FF2FFE0EFED32BB4" box="[234,348,463,489]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="C27236FACF29FFD1FF78FE32FB1D280B" blockId="46.[189,1399,463,598]" pageId="46" pageNumber="47">
<materialsCitation id="72A53CA7CF29FFD1FF78FE32FB1D280B" collectionCode="NHMUK" collectorName="H. E. Andrewes Coll." country="India" determinerName="H. E. Andrewes" location="INDIA" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="BADA766ACF29FFD1FF78FE32FE9F2850" box="[189,272,499,525]" name="India" pageId="46" pageNumber="47">INDIA</collectingCountry>
<specimenCount id="D4CBFD73CF29FFD1FEEDFE32FEB02850" box="[296,319,499,525]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
; with labels and data: white, black printed “8. IX. 17 1. Among Dabui Rarhi and Batri roots Pusa, Haq. Coll:” / “Agric. Res. Inst. Pusa” / handwritten in black ink and printed “
<taxonomicName id="05CD4D79CF29FFD1FB6DFDD6FF71280B" authority="Bates" authorityName="Bates H. E. Andrewes" authorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sagittaria">Clivina sagittaria Bates</taxonomicName>
(over)
<determinerName id="37AEDA3ECF29FFD1FE90FDFAFD8E2808" box="[341,513,571,597]" pageId="46" pageNumber="47">H.E.Andrewes</determinerName>
det.” / “
<collectorName id="6F38532CCF29FFD1FDA7FDFAFCDC2808" box="[610,851,571,597]" pageId="46" pageNumber="47">H.E.Andrewes Coll.</collectorName>
B.M.1945-97.”;
<collectionCode id="A4DCAE3FCF29FFD1FBD9FDFDFB00280B" box="[1052,1167,572,598]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="46" pageNumber="47" type="Museum">NHMUK</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="C27236FACF29FFD1FF78FDBFFB9D28E0" blockId="46.[187,1400,638,701]" pageId="46" pageNumber="47">
<emphasis id="F0B9EAE8CF29FFD1FF78FDBFFEA528C5" bold="true" box="[189,298,638,664]" pageId="46" pageNumber="47">Remark.</emphasis>
In the left hind leg, all tarsomeres are missing and the right elytron shows a pinhole and a crack. The mandibles are not as sharp as in the
<typeStatus id="1D768858CF29FFD1FD5DFD62FD7228E0" box="[664,765,675,701]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
due to individual wear.
</paragraph>
<paragraph id="C27236FACF29FFD1FF2FFD24FED328A2" blockId="46.[189,1398,741,840]" box="[234,348,741,767]" pageId="46" pageNumber="47">
<emphasis id="F0B9EAE8CF29FFD1FF2FFD24FED328A2" bold="true" box="[234,348,741,767]" pageId="46" pageNumber="47">
<typeStatus id="1D768858CF29FFD1FF2FFD24FED328A2" box="[234,348,741,767]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="C27236FACF29FFD1FF78FCC8FB202915" blockId="46.[189,1398,741,840]" pageId="46" pageNumber="47">
<materialsCitation id="72A53CA7CF29FFD1FF78FCC8FB242915" collectionCode="NHMUK" collectorName="H. E. Andrewes Coll." country="India" location="INDIA" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="BADA766ACF29FFD1FF78FCC8FE9F297E" box="[189,272,777,803]" name="India" pageId="46" pageNumber="47">INDIA</collectingCountry>
<specimenCount id="D4CBFD73CF29FFD1FEE2FCC8FEDB2979" box="[295,340,777,804]" count="1" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
; with labels and data: white, black printed and handwritten in black ink “2110 Pusa Behar At Light” / “Agric. Res. Inst. Pusa” / “
<collectorName id="6F38532CCF29FFD1FDBAFCECFCE0291A" box="[639,879,813,839]" pageId="46" pageNumber="47">H.E.Andrewes Coll.</collectorName>
B.M.1945-97.”;
<collectionCode id="A4DCAE3FCF29FFD1FBFDFCEFFB242915" box="[1080,1195,814,840]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="46" pageNumber="47" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C27236FACF29FFD1FF78FCB1FE3B29AA" blockId="46.[187,1400,880,1015]" pageId="46" pageNumber="47">
<emphasis id="F0B9EAE8CF29FFD1FF78FCB1FEB929D7" bold="true" box="[189,310,880,906]" pageId="46" pageNumber="47">Remarks.</emphasis>
The left elytron and the left hind leg are missing, the right elytron shows a pinhole. The ventral surface shows cracks and in the left intermediate leg, three tarsomeres are missing.
<bibRefCitation id="A65C4B0BCF29FFD1FB6DFC55FF7B298E" author="Andrewes H. E." pageId="46" pageNumber="47" refId="ref37824" refString="Andrewes H. E. 1929. The Fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. I., Carabinae. Taylor &amp; Francis, London." type="book" year="1929">Andrewes (1929: 359)</bibRefCitation>
already mentioned some of the outstanding characters of these
<specimenCount id="D4CBFD73CF29FFD1FC19FC78FB06298F" box="[988,1161,952,979]" count="2" pageId="46" pageNumber="47" type="generic">two specimens</specimenCount>
but did not describe them as a new taxon.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF29FFD0FF78FBDEFEB62D2E" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
<paragraph id="C27236FACF29FFD1FF78FBDEFEC12E64" blockId="46.[189,334,1055,1081]" box="[189,334,1055,1081]" pageId="46" pageNumber="47">
<heading id="993A8196CF29FFD1FF78FBDEFEC12E64" bold="true" box="[189,334,1055,1081]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="F0B9EAE8CF29FFD1FF78FBDEFEC12E64" bold="true" box="[189,334,1055,1081]" pageId="46" pageNumber="47">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF29FFD1FF78FB8EFDD82ED0" blockId="46.[187,1398,1103,1165]" pageId="46" pageNumber="47">
MEASUREMENTS.
<typeStatus id="1D768858CF29FFD1FE45FB8EFE632E34" box="[384,492,1103,1129]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
/
<typeStatus id="1D768858CF29FFD1FE3EFB91FDEB2E37" box="[507,612,1104,1130]" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
: body length 6.94/
<quantity id="05359B1FCF29FFD1FCF8FB8EFC2A2E37" box="[829,933,1103,1130]" metricMagnitude="-3" metricUnit="m" metricValue="6.57" pageId="46" pageNumber="47" unit="mm" value="6.57">6.57 mm</quantity>
, width 1.57 /
<quantity id="05359B1FCF29FFD1FBFCFB8EFB2C2E37" box="[1081,1187,1103,1130]" metricMagnitude="-3" metricUnit="m" metricValue="1.61" pageId="46" pageNumber="47" unit="mm" value="1.61">1.61 mm</quantity>
; L/W of pronotum 1.2 / 1.81; L/W of elytra 2.31 / 2.23.
</paragraph>
<paragraph id="C27236FACF29FFD1FF78FB77FE252F45" blockId="46.[189,1399,1206,1304]" pageId="46" pageNumber="47">COLOUR. Glossy. Piceous. Mouthparts, antenna, elytron, intermediate and hind leg, tarsomeres of front leg fuscous, palpomeres more pale fuscous. Anterior part of supraantennal plate at the margin slightly translucent-fuscous.</paragraph>
<paragraph id="C27236FACF29FFD0FF78FA80FCBA2B14" blockId="46.[187,1400,1345,2024]" lastBlockId="47.[189,1399,265,329]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
HEAD. A quarter narrower than pronotum. Clypeus with central part distinctly bilobed anteriorly; lateral lobe developed as acute tooth, nearly as projecting as central lobes, clypeal wing well developed, rounded, not as far projecting anteriorly as lateral tooth of clypeus, separated from lateral lobe by obtuse notch. Supraantennal plate laterally moderately convex, laterally somewhat less projecting than eye, indistinctly reflexed margined. Supraantennal plate overlapping eye anteriorly by about a fifth (dorsal view), with the supraorbital carina keeled posteriorly and extended up to mid-eye level, separated from wing by sharp notch. Clypeus transverse, separated from frons by sharp carina in form like an obtuse inverted V. Frons distinctly convex, with another but smaller carina like an inverted V which encloses elevated area, supraorbital carina short and tubercle-like, extending up to posterior gena-level. Clypeus rugose, area between V-like carinae with transverse rugae, supraantennal plates and frons rugose, posteriorly with distinct longitudinal rugae, clypeus and frons separated from supraantennal plate and supraorbital carina by deep and moderately wide furrow; with two supraorbital setigerous punctures situated at posterior eye and at end of gena-level. With some moderately sized punctures laterally posterior eye. Eye small, embedded, moderately convex in dorsal view, main part situated ventrally where it is more convex. Gena moderately convex, covering around a sixth of posterior eye in ventral view. Eye somewhat constricted due to the overlapping supraantennal plate and the genae. Antenna not reaching up to middle of pronotum, antennomeres five to ten moniliform (L/W around 1.1). Labrum straight anteriorly, with isodiametric-irregular reticulation, six setose. Mandibles distinctly falcate, bent up dorsally.
<collectingRegion id="0009F818CF29FFD1FB85F86AFB2D2D98" box="[1088,1186,1963,1989]" country="Myanmar" name="Chin" pageId="46" pageNumber="47">Mentum</collectingRegion>
with irregular fine wrinkles and indistinct isodiametric reticulation, shape of lobe wide-oval, flattened, with sub-marginal carina, with long median carina. Median tooth moderately wide, not as protruding anteriorly as lobe, hollowed out, obtuse angled and bicarinate anteriorly.
</paragraph>
<paragraph id="C27236FACF28FFD0FF78FEB2FEE628E6" blockId="47.[188,1400,371,699]" pageId="47" pageNumber="48">PRONOTUM. Disk slightly convex in lateral and semicircular in frontal view. Distinctly longer than wide, peltate. Reflexed lateral margin nearly smooth, straight laterally, nearly parallel, anterior angle rounded off, posterior angle missing; lateral channel moderately narrow, narrower before and behind posterior setigerous puncture. Median line moderately deep, sharp, subcrenulated, complete. Anterior transverse crenulated, wider than median line, joining median line, joining declivity at anterior margin. Surface glossy, completely covered with small punctures, with paralateral indistinct group of moderately sized punctures, with few fine transverse wrinkles laterally and basally. Base distinct, channel two times as wide as lateral channel, width about a quarter of width of pronotum. Proepisternum distinct, well visible in dorsal view.</paragraph>
<paragraph id="C27236FACF28FFD0FF78FD24FDA22E0F" blockId="47.[187,1400,741,1106]" pageId="47" pageNumber="48">ELYTRON. Shape elongate with margin nearly parallel. Disk flattened in anterior half in lateral view, semicircular in frontal view. More than two times longer than wide, with maximum at middle. Reflexed lateral margin smooth, setigerous punctures in lateral channel with regularly long setae, punctures widely interrupted at middle. Scutellar striole missing; setigerous tubercle at base of first stria, with tubercle at base of third interval, basal declivity with isodiametric reticulation. Humeral tooth situated at base of fourth interval. All striae distinctly deepened, punctuate-striate, one to three free at base, four reaching humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals distinctly convex, more convex laterally. Setigerous punctures missing on intervals. Surface of all intervals glossy, interval four to eight with indistinct reticulation at humerus, interval eight at apex sub-carinate and with indistinct reticulation.</paragraph>
<paragraph id="C27236FACF28FFD0FF78FBBCFC1E2ECB" blockId="47.[189,913,1148,1175]" box="[189,913,1148,1175]" pageId="47" pageNumber="48">
HIND WINGS. Alae fully developed (based on
<specimenCount id="D4CBFD73CF28FFD0FD17FBBCFC0A2ECB" box="[722,901,1148,1175]" count="2" pageId="47" pageNumber="48" type="generic">two specimens</specimenCount>
).
</paragraph>
<paragraph id="C27236FACF28FFD0FF78FB00FA852F7A" blockId="47.[187,1400,1216,1319]" pageId="47" pageNumber="48">LOWER SURFACE. Proepisternum covered in lateral half with dense and deep transverse wrinkles, with some moderately sized punctures, at the lateral margin with isodiametric reticulation. Sternite of abdomen smooth at middle but with micropunctures, covered laterally with moderately large punctures.</paragraph>
<paragraph id="C27236FACF28FFD0FF78FA91FCEE2FCD" blockId="47.[189,1399,1360,1424]" pageId="47" pageNumber="48">LEGS. Protibia covered with longitudinal reticulation, with three spines of moderate length, apical spine distinctly arcuate, movable spur robust, regularly arcuate.</paragraph>
<paragraph id="C27236FACF28FFD0FF78FA7BFD852F89" blockId="47.[189,522,1466,1492]" box="[189,522,1466,1492]" pageId="47" pageNumber="48">MALE GENITALIA. Unknown.</paragraph>
<paragraph id="C27236FACF28FFD0FF78FA3FFB212C3E" blockId="47.[189,1398,1534,1636]" pageId="47" pageNumber="48">FEMALE GONOCOXITE AND EPIPLEURITE (Fig. 52). Gonocoxite fir-cone shaped, flattened dorsally and ventrally, with three wide nematiform setae positioned in the apical half at the median side, with one preapical nematiform seta. Epipleurite semi-square, with long widened angulated rod.</paragraph>
<paragraph id="C27236FACF28FFD0FF78F94CFEBC2CFA" blockId="47.[189,307,1677,1703]" box="[189,307,1677,1703]" pageId="47" pageNumber="48">
<heading id="993A8196CF28FFD0FF78F94CFEBC2CFA" bold="true" box="[189,307,1677,1703]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="F0B9EAE8CF28FFD0FF78F94CFEBC2CFA" bold="true" box="[189,307,1677,1703]" pageId="47" pageNumber="48">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF28FFD0FF78F97FFADE2CA3" blockId="47.[189,1398,1726,1791]" pageId="47" pageNumber="48">
The carina on the frons of the head is arch-like rounded in the HT and obtuse-angled in the PT. The punctures on disk of the pronotum are different in intensity and density between the
<specimenCount id="D4CBFD73CF28FFD0FB5EF924FAC22CA3" box="[1179,1357,1764,1791]" count="2" pageId="47" pageNumber="48" type="generic">two specimens</specimenCount>
.
</paragraph>
<paragraph id="C27236FACF28FFD0FF78F8E9FE3F2D1F" blockId="47.[189,432,1832,1858]" box="[189,432,1832,1858]" pageId="47" pageNumber="48">
<heading id="993A8196CF28FFD0FF78F8E9FE3F2D1F" bold="true" box="[189,432,1832,1858]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="F0B9EAE8CF28FFD0FF78F8E9FE3F2D1F" bold="true" box="[189,432,1832,1858]" pageId="47" pageNumber="48">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF28FFD0FF78F898FEB62D2E" blockId="47.[189,313,1881,1907]" box="[189,313,1881,1907]" pageId="47" pageNumber="48">Unknown.</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF28FFD0FF78F85CFC992DB5" box="[189,790,1949,2025]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="C27236FACF28FFD0FF78F85CFED92DEA" blockId="47.[189,342,1949,1975]" box="[189,342,1949,1975]" pageId="47" pageNumber="48">
<heading id="993A8196CF28FFD0FF78F85CFED92DEA" bold="true" box="[189,342,1949,1975]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="F0B9EAE8CF28FFD0FF78F85CFED92DEA" bold="true" box="[189,342,1949,1975]" pageId="47" pageNumber="48">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF28FFD0FF78F80EFC992DB5" blockId="47.[189,790,1998,2025]" box="[189,790,1998,2025]" pageId="47" pageNumber="48">
Known from the
<typeStatus id="1D768858CF28FFD0FE43F80EFE362DB4" box="[390,441,1999,2025]" pageId="47" pageNumber="48">type</typeStatus>
locality Pusa,
<collectingRegion id="0009F818CF28FFD0FDA0F80FFD272DB5" box="[613,680,1998,2024]" country="India" name="Bihar" pageId="47" pageNumber="48">Bihar</collectingRegion>
, in
<collectingCountry id="BADA766ACF28FFD0FD17F80FFC9D2DB5" box="[722,786,1998,2024]" name="India" pageId="47" pageNumber="48">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>